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Comprehensive Description

provided by Memoirs of the American Entomological Society
Laccophilus maculosus decipiens LeConte Laccophilus decipiens LeConte, 1852, p. 205. Type: Museum of Comparative Zoology, number 5974, male; type locality here restricted to 2.8 miles east of La Grange, Vizard Creek, Stanislaus County, California; Crotch, 1873, p. 400; Sharp, 1882, p. 10; Horn, 1894, p. 313; Leech, 1948b, p. 402; LaRivers, 1951, pp. 397-8; Leech & Chandler, 1956, p. 314; Gordon & Post, 1965, p. 12.
Laccophilus truncatus Mannerheim, 1853, p. 160.
Laccophilus calif ornicus Motschulsky, 1859, p. 172.
Laecophilus (sic!) maculosus Walker, 1866, p. 317.
Laccophilus fusculus Sharp, 1882a, p. 290.
DIAGNOSIS. — There are only three other irrorated Laccophilus that occur within the range of L. m. decipiens in western North America. All three, L. mexicanus, L. fasciatus terminalis, and L. proximus, are smaller than decipiens. Almost all decipiens exceed 5.2 mm which is a maximum length for any of those three. L. proximus which overlaps decipiens east of the Rocky Mountains, attains a length of only 4.6 mm. All males and most females of L. mexicanus have black on the venter instead of brown or reddish-brown as in decipiens. Some females of mexicanus from coastal California and Oregon have brown venters, however, and a careful comparison of size and elytral pattern is necessary to separate the two species. L. mexicanus has a more uniformly irrorate pattern. L. f. terminalis, on the other hand, is less evenly irrorate than decipiens and has a checkered appearance when viewed megascopically. Some depigmented specimens are not checkered, but they are much paler than decipiens.
L. in. decipiens differs from L. m. maculosus in its larger size (5.1 to 6.4 mm as compared to 4.7 to 5.8 mm), more evenly irrorate pattern, and the structure of the aedeagus. It is quite similar to L. m. shermani in size, color and pattern (figs. 276-279), but differs in the more anterior position of the female epipleural flange, a finer coxal file (about 45 lines to 35), and in the aedeagus. It is flattened at the tip in decipiens and twisted with a sinus and incomplete hook at the tip in shermani.
NOMENCLATURAL NOTES. — The synonymy has been confirmed except for truncatus Mannerheim and calijornicus Motschulsky. Dr. F. N. Young has examined the female type of fusculus Sharp and has sent me a camera lucida drawing and there is nothing to distinguish it from numerous specimens of decipiens. It is probably not shermani Leech since the epipleura are not expanded; and Nevada, given as the type locality, seems to be entirely out of the range of that race. Since the description is based on a single female specimen and there is no precise locality, it is probably impossible to be certain. Motschulsky's calijornicus was taken at San Francisco, California, and could conceivably prove to be either L. mexicanus atristernalis Crotch or L. fasciatus terminalis Sharp. Crotch (1873) had, however, synonymized truncatus and calijornicus when he described atristernalis and presumably knew their identity. Mannerheim's truncatus was described from Alaska and the Pribilof Islands which is within the range of biguttatus Kirby and it is remotely possible that it could be a synonym of that frequently misidentified species. I have examined LeConte's type and compared it to a specimen collected by H. B. Leech from Stanislaus County, California, and could find no difference other than that of normal individual variation.
VARIATION. — Although exhibiting more variation than L. m. maculosus, decipiens has relatively little considering its enormous geographical range and ecological and topographic amplitude. Specimens from near the Coast and in Nevada, however, do appear to be darker with more suffusion of color between the dots and a more pronounced delineation of the marginal clear areas than those from the Rocky Mountains and Great Plains. The lateral spots may also be
smaller. Specimens from Wyoming appear faded when compared to coastal ones.
The mean length of males is about 5.8 mm and that of females about 5.6 mm, with an overlap of males and females of about 50 percent. This contrasts with maculosus in which about 75 percent or more of the males and females overlap. The variation among localities is slight and an analysis of variance for 240 males from eleven scattered localities for total length shows no significance at the 95 percent level.
The PW/EL ratio also shows a strong uniformity within decipiens. Mean values vary from 0.598 to 0.626 for eleven localities. The ratio does show, however, that decipiens in addition to being slightly larger than m. maculosus, is also relatively wider. The latter has mean values ranging from 0.580 to 0.600. The ratio is about the same for L. m. shermani as decipiens, but the former may be slightly narrower since there are five samples with values below 0.595 (none in decipiens).
An attempt was made to count the lines in the metacoxal file of specimens from different localities. There was no apparent geographical difference. The highest count was 50, and was found in both Wyoming and Oregon specimens and the lowest count was 40 in individuals from Utah and California. The average number was about 44. Only about five to ten percent of the females have epipleural flanges. — L. m. decipiens (fig. 3) ranges from northern Baja California to Alaska along the Pacific Coast. It extends eastward to Alberta and central Colorado. It intergrades with shermani along the northern edges of New Mexico, Arizona and into eastern Riverside County, California. There is a single specimen from Hale County in the northern panhandle of Texas. It occurs from sea level to 9000 feet. The principal difference between the habitats of decipiens and maculosus is that the former is found in or near coniferous forest instead of temperate deciduous forest. Practically any non-marine aquatic environment other than large lakes and streams provides suitable habitat for the adults. CANADA. — ALBERTA. Calgary, 1 $, 19, vii.4.53; 1 <S , 12, iv.30.57; 1 $, 1 9, v.12.57; 1 S, xii.7.57, F. S. Carr (CARR): 10, x.16.27, O. Bryant (CAS). Champion, 1 2, v. 18.59, F. S. Carr (CARR). Cypress Hills, 1 2, vi.13.24; 1 2, v.13.28 (DAV). Edmonton, 1 2, v. 1.26, F. S. Carr (DAV). Ft. McCleod, 3 3,5 2, iii.18.56, F. S. Carr (CARR); 50+, ix.15.28, O. Bryant (CAS). High River, 1, ix.2.28, O. Bryant (CAS). Medicine Hat, 13,19, x.12.58; 4, viii.25.23, F. S. Carr (CAS); 4, ix.3.23; 4, viii.7.26, Shoemaker (USNM). Pincher Ck., 2 2, viii.6.28 (DAV). Porcupine Hills, 1 2, x.18.59, F. S. Carr (CARR). BRITISH COLUMBIA. Bon Accord, 21, Hubbard & Schwarz (USNM). Creston, 2, ix.4.54, G. Stace Smith (CNL). Fernie, 10, ix.16.28, O. Bryant (CAS). Golden, 2 3, 1 2, 8.00.03, P. Gregson (AMNH). Kamloops, 2, ii.1.29, O. Bryant (CAS). Marysville, 1, vi.6.56, E. Fodor (CNL). Osoyoos, 1, iii.29.41, H. B. Leech (CAS). Victoria Is., Royal Oak, 2, v.19.56, E. Argyle (CNL). Victoria, 1, Wickham (USNM). Vernon, 4, v.1.29, R. Hopping (ISU). Windermere, 8, ix.17.28, O. Bryant (CAS). Wynndel, 10, viii. 11.54, G. Stace Smith (CNL).
UNITED STATES OF AMERICA. — ARIZONA. Mohave County. Littlefield, 2 3, 1 2, ix.6.53, P. D. Ashlock (CAS); same date, 13,12 (DAV). CALIFORNIA. Alameda County. Albany, 3, iii.16.21 (BERK). Berkeley, 3 3,9 2, x.15.37, vi.7.37, J. J. Dubois (BERK); 5, x.4-25.19, H. Dietrich (BERK); 4 3, 2 2, xii.14.29, A. T. McClay (DAV); 2, iv.20.41, H. P. Chandler (BERK). Berkeley, Strawberry Pool, 3 3, iii.24.48, R. Johnston (BERK). Livermore, Arroyo Mucho, 2 3, x.7.56; 16 m. S., 12, x.7.56, J. Chemsak (BERK); 1 2, x.8.54, A. M. Barnes (BERK); 1 3, iii.24.64, W. W. Wade (CAS); 1 2, viii.7.29, A. T. McClay (DAV). Alpine County. Monitor Pass, 8300 feet, 1 2, viii.24.63, H. B. Leech (CAS). Amador County. Volcano Cn., 1 3, xi.3.57, R. E. Rice (DAV). Calaveras County. Altaville, 4.5 m. W., 1 3, viii.31.63, H. B. Leech (CAS). Mokelumne Hill, 1, vii. — . — , Blaisdell (CNL). Contra Costa County. Tilden Park, 2 2, x.27.48; 1 2, xi.9.48, R. Johnston, 1 3, xi.21.56, J. A. Chemsak (BERK). Vine Hill, 3, vi.20.10, Chamberlain (CNL); 1 3, vi.20.10, Chamberlain (AMNH). County, 1 3, vi.3.09, F. W. Nunenmacher (FM). Eldorado County. Camino, 3 m. S., 4 3, 6 2, vi.23.48, Carter, Myers, & Smith (BERK); 2 3,2 2, same date, H. P. Chandler (DAV). Chile Bar, 3 3,2 2, vii.5.48, H. P. Chandler (BERK). China Flat, 2 2, vii.15.48 (BERK). Echo Lake, 3, vii.13.32, E. O. Essig (BERK). Pollock Pines, 13,2 2, vi.29.38; 1 2, vi.24.38; 1 3, vii.3.36; 1 3, 5 2, vi.23.39; 13,12, vii.8.39; 1 3, viii.18.39; 1 3, vii.23.39; 1 2, ix.1.39; 1 3, 3 2, vii. 8.48, A. T. McClay (DAV). Rubicon R. (near Georgetown), 1 2, vii.28.63, H. B. Leech (CAS). Tahoe, 1, vii.3.50, R. Schuster (BERK). County, 1 2 , vi.8.06, F. W. Nunenmacher (FM) . Del Norte County. Smith R. at Hwy. 101, 1 3, ix.29.65, H. B. Leech (CAS). Fresno County. Fresno, 1, v.30.— , F. A. Schwarz (USNM). Wartham Ck., near Parkfield, 1 3, xi.30.63, H. B. Leech (CAS). Humboldt County. Ferndale, 1, ix.14.46, H. P. Chandler (BERK). Bull Ck. at Cuneo Ck., 1 2, x.2.65, H. B. Leech (CAS). Inyo County. Deep Springs, 1 3, vi.19.54, P. Raven (CAS). Keeler, 1, vii.6-14. — , Wickham (USNM). Lone Pine, Owens Valley, 1, iv.20.31, Blackwelder (USNM); 1 3, 2 2, v.20.37, J. J. Dubois (BERK). Olancha (?) 1, J. D.
Sherman, Jr. (USNM). Kern County. Red Canyon, 1 2, vi. 10.22 (AMNH). Lake County. Thurston Lake, 1, H. P. Chandler (BERK). Lassen County. Eagle Lake, 1, ix.19.46, H. P. Chandler (BERK) . Hallelujah-Jet., 1 2 , vii.4.49, J. W. McSwain (BERK); 13,12, vii.4.49, H. A. Hunt (DAV). Madeline, 1, vii.12.25 (USNM). Poison L., 1 3, vi.13.41, C. W. Anderson (BERK). Susanville, 1, viii.1.32, Blackwelder (USNM). Willow Ck., 13,12, ix.9.58, Allyn Smith (CAS) . Los Angeles County. Claremont, 6 3,72, Baker (CNG) . Lancaster, 3 m. N., 8, iv.3.53, R. K. Benjamin (USNM). Los Angeles, 5, vii.— .— , W. D. Richardson (USNM); 13, Coquillett (USNM); 1, ii.— .19, R. P. Dow (USNM). Palmdale, 1 2, vi.18.44, G. P. Mackenzie (LACM). Pasadena, 2 3,12, vi.27.16; 1 2, v.29.20 (AMNH). Pine Can., Hidden L., 13,12, ii.15.53; 1 3, iii.21.53, Menke & Truxal (DAV). Pomona, 4, J. D. Sherman, Jr. (USNM). San Franquito Can., 4 3, ii.29.53; 13,12, iii.30.53; 2 3, 1 2, iv.1.53; 1 3, iv.3.53, L. A. Stange (DAV). Soledad Can., 1 3, i.10.54, Menke & Sullivan (DAV). Tanbark Flat, 1 2, vii. 13.50 (BERK). Madera County. Chilkoot L., Jackass Mtns., 4 3, 2 2, vii.23-31.46, H. P. Chandler (NMSU). Chowcilla, 18 m. E., 1 3, ii.2.64, H. B. Leech (CAS). Coarsegold, 1 3,1 2, H. P. Chandler (BERK). O'Neal's, 1 3, iii. 10.46, H. S. Fitch (DAV). Windy Gap. 1, vii. 13.46, H. P. Chandler (CAS). Marin County. Dillon Beach, 1 2, vi.24.63, H. B. Leech (CAS). Inverness, 2 3, 1 2, vii.20.50 (BERK). Pt. Reyes Sta., 2 3, x.1.60, J. S. Buckett (DAV). Ross Valley, 1 2, ix.2.06, E. C. Van Dyke (AMNH). County, 2, J. D. Sherman, Jr. (USNM). Mariposa County. Miami Rngr. Sta., 1 2, vi.7.42, H. P. Chandler (FM). Yosemite, 13,12, vii.17.48, B. Adelson (BERK); 1 2, v.24.64, W. W. Wade (CAS). Mendocino County. 1, v.31.17, E. R. Leach (USNM). Albion, 1 m. S., 1 3, vii. 1.63, H. B. Leech (CAS). Modoc County. Cedarville, 5 m. E., 1 2, vi.5.60, J. Schuh (SCH). Juniper Flat, 1 3, vi.10.39, J. J. Dubois (BERK). Mono County. Bodie, 8500 feet, 1 2 , viii.13.62, H. B. Leech (CAS). Bridgeport, Travertine Hot Springs, 2 m. S.E., 6700 feet, 1 3,
1 2, viii. 11.62, H. B. Leech (CAS). Inyo Craters Lake, 8280 feet, 1 3, viii.17.62, H. B. Leech (CAS). June Lake, 13,5 2, ix.8-9.45, G. P. Mackenzie (LACM). Leavitt Meadows, 7200 feet, 1 3, viii. 10.62, H. B. Leech (CAS). Mammoth (Lakes?), 4 3,5 2, ix.15.45, G. P. Mackenzie (LACM). Twin Lakes, 1, vi.14.31, Blackwelder (USNM). Monterey County. Bryson,
2 2, iv.9.63, D. C. Rentz & K. A. Hale (CAS). Parkfield, 1 2, xi.20.63, H. B. Leech (CAS). Salina(s?), R., 3, vii. 17.25 (USNM). Napa County. Monticello, Putah Ck., 2, xii.17. — , J. C. Bradley (CNL); 1 3, vii.22.30, A. T. McClay (DAV); 2 3,3 2, iii. 30.31 (DAV). Nevada County. Hobart Mills, 1, H. D. Chandler (BERK). Sagehen Ck., 13,12, x. 15.55, J. L. Herring (BERK). Webber L., 13, viii.22.64, E. S. Ball (SCH). Orange County. Costa Mesa, 1 3, 2 2, iv.19.58, M. E. Irwin (DAV). Laguna Beach, 3, viii.25-26.— (CNL); 1, vii.— .21, C. T. Dodds (BERK). Laguna Can., 2 3, xii.29.58, M. E. Irwin (DAV). Placer County. Tahoe, 14, vii.—.— (USNM); 14, vii.6.09, J. D. Sherman, Jr. (USNM); 4 3, 6 2, vii.—.— (AMNH). Plumas County. Almanor Dam, 1 m. N.W., 2 3,2 2, vii.17.47, C. H. Spitzer (CAS). Buck's Lake, 13,12, vii. 14.49, E. I. Schlinger (DAV). Chester, 1 9, vii.29.61, H. B. Leech (CAS). Clio, 1 9, viii.28.61, H. B. Leech (CAS). Quincy, 4 m. W., 3 6,4 9, vii.13.49, W. H. Wade (BERK); 13,3 9, vii.6.49, L. W. Isaak (DAV). Riverside County. Camp Pendleton, 2, x.7.45, H. P. Chandler (BERK). Riverside, 3 6, 2 9, F. E. Winters (CAS); 6, viii. 16.37, Drake & Andre (USNM); 14, F. E. Winters (CNL). San Jacinto Mtns, 1 6,
2 9, — . — .32, F. E. Winters (CAS). San Jacinto Mtns., Keen Camp, 1 9, R. F. Smith (FM). Sacramento County. Folsom, 1 9, iv.5.47, A. T. McClay (DAV). Franklin, 1, iii.10.21, C. M. Packard (USNM). Sacramento, v.7.41, A. T. McClay (DAV); 1 9, ii. — .20, H. F. Robinson (DAV). San Bernardino County. Afton Canyon, 4, iv. 18.31, Blackwelder (USNM). Barstow, 1 6 (NMSU); 2 6,29 (AMNH); 7, viii. 19.88 (ISU). Barton Flats, 2 9, x.4.58, M. E. Irwin (DAV). Colton, 1 (USNM). Cottonwood Sprgs., 7 6, 7 9, iv.5.48, MacSwain & Smith (BERK). Hesperia, 1 $, iv.7.39, G. P. MacKenzie (AMNH). Mtn. Pass, 1 6, iv. 11.42, O. Bryant (CAS). Sweetwater (Sprgs.?), 1, vi.2. — , Dyar & Candell (USNM). San Benito County. Gonzales, 16 m. E., 2 9, iii. 30.63, D. Reutz & K. Hale (CAS). San Diego County. Anza-Borrego St. Pk., 1, iv. — .53, A. & H. Dietrich (CNL). Banner, 6 m. E., 1 $, vi.26.63, J. D. Birchim (CAS). Borrego (Sprgs.?), 1 6, iv.12.50, W. F. Barr (FM). Campo, 1 9, iv.13.65, H. B. Leech (CAS). Descanso, 4 6, 1 9, ix.23.52, 1, viii.7.14, J. C. Bradley (CNL). Escondido, 10 m. N., 5, xii.17.54; 12 m. N., 1, xi. 20.53; 12 m. N., 1 9, iv.17.63, R. K. Benjamin (USNM). Glenncliff Cp. (near Buckman Sprgs.), 3 8, 5 9, ix.21.54, Menke & Stange (DAV). Laguna Mtns., near Buckman Sprgs., 2 6, 1 9, v.6.53, F. X. Williams (CAS). Mt. Palomar, 1 6, vi.21.59, M. E. Irwin (DAV); 2, x.21.45, H. P. Chandler (BERK). Oceanside, Camp Pendleton, 1 9, x.7.45, H. P. Chandler (FM). San Diego, 1, Mann (USNM); 1, G. H. Field (USNM); 2 (USNM); 2, iv.8.54, R. Straw (USNM). San Felipe Ck., 4 m. N., Scissors Crossing, 2 9, iv. 15.65, H. B. Leech (CAS). Sentenas Cn., 1 9, 1,26.56, Menke & Strange (DAV). Warners (Springs), 1 9, viii.19.25 (MCZ). County, 1 9 (FM). San Mateo County. San Francisco Bay Marsh, Redwood City, 1 9,x.l.50, PSB (CAS). Woodside, Pulgas Temple, 4 6, 10 9 , vi.10.51, PSB; 5 6,3 9, xi.4.51, PSB; 2 9, vi.23.52; 1 9, xi.25.54, PSB (CAS). Santa Barbara County. Los Prietos, 4 m. E., 1 6, vi.26.65, M. R. Gardner (DAV). Santa Barbara, 1, iv. — .— , F. E. Winters (CNL); 4 6, 3 9, F. E. Winters (CAS). Santa Clara County. Palo Alto, 2, ii.2.26, Blackwelder (USNM); 1 9, vi.6.92, V. L. Kellog (FM); 1, v.19.40, H. P. Chandler (BERK). Palo Alto, Stanford U., 6 6 , 1 9, ix.12.09, W. M. Mann (USNM). San Antonio Val., 2, iii.2.48, J. W. MacSwain (BERK). San Jose, 1, Hubbard & Schwarz (USNM). County, 1; 2 6, 2 9, C. F. Baker (USNM). Shasta County. Hat. L., 1 6, vii. 11.47, R. L. Usinger (BERK); 1 9, vii. 11.47, E. E. Seibert (CAS); Hat Ck., P.O., 1 9 , vi.27.55, A. J. Mueller (DAV) . Lake Britton, 1 6 , vi.29.47, C. H. Spitzer (CAS). Manzanita L., 5800 feet, 2 6, 3 9, x.1.44, B. Malkin (FM); Manzanita L., Lassen Nt. Pk., 1 9 , C. W. Anderson (BERK). Old Station, IS, 19, vi.26.47, E. E. Seibert (BERK). Viola, 4 m. W„ 2 3, vi.27.47, E. E. Seibert (BERK). Sierra County. Sattley, 4 6,3 9, vii. 11.62,
R. L. Westcott (FM); 3 3, 5 9, vii. 11.62, Gill & Montgomery (DAV). Snag Lake, 2 3,3 2, vii. 5. 64, J. E. Slansky (DAV). Siskiyou County. Grass Lake, 5000 feet, 1 3, vii.4.52, M. Cozier, W. Gertsch, R. Schrammel (AMNH). Indian Tom L., 2 3, 2 2, ix.27.66, J. Schuh (SCH). Lower Klamath L., 4 2, vi.1.57; 1 3, vi.6.66, J. Schuh (SCH). Montague, 10 m. S.E., 1 3, viii.27.64; 10 m. N.E., 1 2, viii.27.64, J. Schuh (SCH). Mt. Hebron, 5 m. S., Butte Ck., 1 3,3 2, v.1.66, J. Schuh (SCH). Weed, 4 m. S., 1 2, viii.12.58, J. Howell (NMSU). Willow Ck., 1 2, vii.14.55, J. Schuh (MCHS); 1, ix.3.59, J. Schuh (SCH); Willow Ck., Jewell Sprg., 1 2, ix.22.62, J. Schuh (SCH). Stanislaus County. La Grange, 2.8 m. E., 2 3, 1 2, viii. 14.62, H. B. Leech (CAS). Sonoma County. Guerneville, 1, v.20.08, E. C. Van Dyke (USNM). Santa Rosa, 2 (CNL). Solano County. Rio Vista, 1, iii.26.21, C. M. Packard (USNM). Vacaville, 1 3, iv.15.43; 1 2, iv.8.49, A. T. McClay (DAV). Trinity County. County, 1, ix.16.— , E. R. Leach (USNM); 1 3, viii.22.29, E. R. Leech (CNG). Tulare County. Camp Potwisha, 4, vii. 16.31, E. C. Zimmerman (USNM). Kaweah, 4, Hopping (USNM). Porterville, 1 2, viii.19.62, E. E. Ball (SCH). Sequoia Ntl. Pk., 1 3, v.28.29; 1 2, v.19.30, A. T. McClay (DAV). Three Rivers, 1, vii. 12-14.07 (CNL). Tuolumne County. Leland Mdw., 13,12, viii.21.60, D. Q. Garagnaro (DAV). Near Knights Ferry, 1 3, ix.22.63, H. B. Leech (CAS). Priest, 4.2 m. S.E., 1 3, viii.19.62, H. B. Leech (CAS). Ventura County. Chuchupate Rgr. Sta., Frazier Mtn., 1 2 , v.2.59, C. W. O'Brien (BERK) . Ojai, 1 3 , vii.— .54, T. Barnes (LACM). Yolo County. Davis, 1, iv.— .49 (USNM); 1 3, ix.12.4, H. R. Cameron (ORES); 1 2, iii.28.49, E. I. Schlinger; 1 2, iii.5.50, M. E. Taylor 1 3, iv.18.59; 1 3, x.23.59, M. E. Irwin; 2 3, x.15.60; 1 2, ix.24.60, D. Q Caragnaro; 6 3, vi.26.61, V. L. Vesterby (DAV). Putah Cn., 1 2, iv.15.52 A. A. Grlgarick (DAV). COLORADO. Alamosa County. 1 8, vi.14.30; 1 8
1 2, vii.2.30, S. C. Bishop (NMSU); 1 8, iii.7.— , F. C. Bowditch (MCZ) Hooper, 9-10 m. E., San Luis Lakes, 2 8, 4 2, viii.13.65, H. B. Leech (CAS) Archuleta County. Piedra, 3 8, vii.19.52, B. Malkin (FM). Chaffee County Buena Vista, 1, vii. 6. 96, Wickham (USNM). Conejos County. Conejos R. at Hwy. 17, 1 8, 1 2, viii.12.65, H. B. Leech (CAS). N. Fork, Los Pinos R., at Hwy. 17, 1 8, viii.12.65, H. B. Leech (CAS). Denver County. Denver, 1, vi.25.— (CNL); 10, x.— .35, A. Thrupp (CAS); 4 8, 2 2, vii.19.— (MCZ). Garfield County. Glenwood Springs, 2 2, viii. — . — (MCZ). Gunnison County, 6, J. D. Sherman, Jr. (USNM). Gunnison, 1, ix. 19.33, E. A. Andrews (CAS). County, 7500 feet, 13,12, vi.26-30.85, F. C. Bowditch (MCZ). Saguache County. Mineral Hot Springs, 13,2 2, viii.13.65, H. B. Leech (CAS). Saguache, 24 3, 15 2, viii.18.52, B. Malkin & V.E.T. (FM); 10.5 m. N.W., 1 3 viii.12.65, H. B. Leech; 12.5 m. N.W., 1 3, viii.12.65, H. B. Leech (CAS) Weld County. Greeley, 1, Wickham (USNM). Adams County. Mesa, 2 m N. (Hwy. 95), 1 2, vii.20.6?, R. D. Anderson (RDA). Bear Lake County Paris (near), 1, vii.10.35 (CNG). IDAHO. Bingham County, 1, vii.24.— ; Hubbard & Schwarz (USNM). Blaine County. Clarendon Hot Springs, 5 3
2 2, vii.22.52, B. Malkin (FM). Bonneville County. Idaho Falls, 13,22 (CNG). Elmore County. Mountain Home, 1, vii. 17.34, Lanchester (CAS). Latah County. Moscow Mtn., 1, vii.20.38, H. M. Harris (ISU). Troy, 1, vii.16.38, H. M. Harris (ISU). Nez Perce County. Lenore, 1000 feet, 1 5, iv.1.51, W. F. Barr (FM). Oneida County. Malad City, 5 m. W., L. Malad R., 15,3 9, vii. 13.52, B. Malkin (FM). Owyhee County. Hot Springs, 1 9, vii.4.51, W. F. Barr (FM). Twin Falls County. Rogerson, 11 m. E., 2 9, vii.20.52, B. Malkin (FM). MONTANA. Dawson County. Glendive, 1, Wickham (USNM). Flathead County. Kalispell, 1, Wickham (USNM). Glacier County. Cutbank, 2 5,19, viii. 22.54, P.S.B. (CAS). Jefferson County. 1 5, vii.27.25 (USNM). Ravalli County. Corvallis, 2 9, v. 20.52, A. A. Hubert (USNM). Sanders County. Thompson Falls, 1 5, 1 9, vii.22-23.50, B. Malkin (FM) ; 1, vii.22-23.50, B. Malkin (CAS). NEVADA. Elko County. Carlin, 2, vii.26.12 (USNM); 1, vii.17.12 (CNL). Lamoille Cn., 8 5, 24 9, vi.14.58, F. D. Parker (DAV). Lincoln County. Hiko, Tonapah Jet., 1 5, vii.25.63, P.S.B. (CAS). Nye County. Ash Meadows, 15, 19, iii.30.66, J. Schuh (SCH). Beatty, 2m.N,H, iii.29.66; 3 m. N. Amargosa R.,3 5, iii.29.66, J. Schuh (SCH). Warm Springs, 6 5,5 9, vii.25.63, P.S.B. (CAS). Ormsby County. Carson City, 3, vii. 30. — , Wickham (USNM). Washoe County. IXL Ranch, 2 9, vii.— .51, N. Pasquale (ORES). Reno, 15,19 (AMNH); 2 9, i.2.57, F. Parker (DAV); 1 5, Wickham (ANSP); 1, Wickham (USNM). OREGON. Benton County. Buchanon, 4 5,5 9, ix. 10.49, V. Roth (ORES). Corvallis, 1 9, viii. 13. 35; 1 9, x.18.35, G. Ferguson (ORES); 3 9, xi.2.06 (ORES); 1 9, x.28.06, Wood (ORES); 2 5, 1 9, xi.1.08, Hays, Sprague & Reynolds (ORES); 3 9, vi.23.— ; vii.4.09, iv.5.36 (ORES); 1 9, iii.30.30, R. L. West (ORES); 2 5, 3 9, x.24.— , xi.3.33, J. Schuh (ORES); 1 5, v.26.35, I. Tarshis (ORES); 2 5,2 9, iii.7-16.36, N. D. Larsen (ORES); 1 9, iv.10.38, S. Jewett (ORES) . Corvallis, Oak Ck., 1 5,1 9 , v.— .39, G. G. Black (ORES) . Corvallis, Colorado L., 4 9, iv.7.30, L. Seghetti (ORES). 3, iv.22.09, G. F. Moznette (CNL); 3 (USNM); 5, iv.25.— , J. D. Sherman, Jr. (USNM); 44, J. D. Sherman, Jr. (USNM); 21, 1, iv.28.01, J. D. Sherman, Jr. (USNM); 8, vi.24.09, J. D. Sherman, Jr. (USNM); 17 5, 24 9, iv.27.— , 14 5, 11 9, vi.24.— ; 10 5, 1 9, vii. 18.—; 4 5,3 9, ix.20.— ; A. R. Woodcock (AMNH). Summit Prairie, 4 5,6 9, viii.9.39, Gray & Schuh (ORES). Yew Ck., near Mt. Alsea, 2 5, v.25.45, G. L. Bennett (ORES). Coos County. Coos Bay, 15,19, vii.9-10.51, B. Malkin (FM). Curry County. Pistol R., 6 5,6 9, ix.18.50; 1 5, vii.7.51, B. Malkin (FM); 1, vii.7.51 (CAS). Deschutes County. Bend, 4, vii.23.32, R. Blackwelder (USNM) . Bend, 20 m. S., 2 5 , 7 9 , iii.8.39, Gray & Schuh (ORES). Near Tumalo, Deschutes R., 3 5, 3 9, iii.22.39, Gray & Schuh (ORES). Douglas County. Gardiner, 8 m. E., Otter Slough, 2 9, vi.1.66, J. Schuh & J. Vertrees (SCH). Grant County. Kimberly, 15 m. S., 2 5, vii.1.60, J. D. Lattin (ORES). Silvies Valley, 2 5, iv.28.57, J. Schuh (SCH). Harney County. Blitzen Valley, 15,4 9, viii. 1.36, S. Jewett (ORES). Buena Vista, 1 5, v.19.56, J. D. Lattin (ORES). Burns, 25 m. E., 2 5, viii. 21.45, H. A. Scullen (ORES). Burns, 15 m. S., 1 5, vii. 13.53, Roth & Beer (ORES). Burns, 3 5,5 9, vi.21.51, B. Malkin (FM). Malheur, L., 1 5, vi.20-21.51, B. Malkin (FM). Malheur Wldlf. Ref., 1 5, viii.15.62
(MCHS). Jackson County. Trail, 13, 29., ix.9.50, Malkin & Thatcher (FM); 1, ix.9.50 (CAS). Medford, 3 8 , 3 9 , iv.1.39, F. Lawrence (DAV). Josephine County. Grants Pass, 1 8 , vii. 3.06, Eismann (ORES). Rogue River, 1, ix.8.02, Biederman (USNM). County, 1 9, vi.6.34, F. W. Nunenmacher (FM). Klamath County. Aspen L., 3 8, 3 9, v.5.57, J. Smith (SCH). Bly, 5 m. E., 4 8, v.5.62, R. Scott & J. Schuh. Bly Mt., 2 $, 4 9, ix.15.60, J. Schuh (SCH). Chiloquin, 12 m. E., Sprague R., 1 <?, 3 9, vii. 1-3.51, B. Malkin (FM). Chiloquin, 7 m. N., Williamson R., 1 8, 2 9, ix.23.66; 6 m. E., 1 8, 1 9, ix.21.65, J. Schuh (SCH). Crescent, 4<5,59, ix.10.50, B. Malkin (FM); 1 9, v.12.57; 1 9, J. Schuh (SCH); 2 m. N.E., 2 9, v.13.66; 11 m. N.E., 1 9, v.13.66, J. Schuh; 13 m. S.E. Bly, 1 9, iv.26.62, Vertrees & Schuh (SCH). Ft. Klamath, 5 <5, 3 9, viii.10.50, B. Malkin (FM). Klamath Falls, 10 m. N.W., 1 $, vi.16.52 (ORES); 1 8, iv.22.57; 1 8, v.2.57; 1 $, iv.23.53; 1 9, v.31.48, J. Schuh (ORES); Modoc Pt., 1 9, vi.1.55, Schuh (ORES); Poe Valley, 1 9, v.27.55, Schuh (ORES); 5 8, 2 9, ix.3.50, B. Malkin (FM). Klamath (?), 1, G. M. Greene (USNM). Mare's Egg Spring, 3 8, ix.7.59, J. Schuh (SCH). Midlland, 3 m. S., 20 8, 13 9, ix.3.50, B. Malkin (FM). Poe Valley, 4 $, v.13.66, J. Schuh (SCH). Lake County. Abert L., 1 8,3 9, vi.7.58, Vertrees & Schuh (SCH). Chandler St. Pk., 4 <$, vi.29-30.51, B. Malkin (FM). Near Valley Fall, Chewaucan R., 1 $, 19, vi.7.55, J. Schuh (ORES); 1 $, 11 9, iii.16.39, Schuh & Gray (ORES). Lakeview, 1 9, vi.2728.51, B. Malkin (FM); 1 $, vi.— .58, J. Schuh (SCH). Silver L., 10 m. N., 3 9, v.12.57, J. Schuh (SCH). Summer L., 1 9, v.18.57, J. D. Lattin (ORES). Lane County. Eugene, 2 9, vi. 26.41; 2 9, viii.2.41; 1 $, 1 9, iv.5.42; 2 9, iv.11.42; 2 $, vii. 13-14.51, B. Malkin (FM). Linn County. Albany, 1, vi.15-28.— , Wickham (USNM). Tangent, 2(5,39, v.4.38, Edman (ORES). Malheur County. Drewsey, 4 m. E., 1 $, 1 9, ix. 10.64, J. Schuh & W. Peters (SCH). Multnomah County. Gaston, 4, J. D. Sherman, Jr. (USNM). Portland, 1, J. D. Sherman, Jr. (USNM). Wasco County. Simnasho Hot Springs, 14 m. S., 1 <?, v.22.59, J. D. Lattin (ORES). Washington County. Forest Grove, 1 8, x. — .06, J. C. Birdwell (ORES). Yamhill County. McMinnville, 1 8, vii.— .36, K. Fender (FM); 1 8, v.— .38, K. M. & B. M. Fender (LACM). TEXAS. Hale County. Plainview, 1 8, vii. 14.54, F. C. Harmston (RDA). UTAH. Boxelder County. Rabbit Sp. Reservoir, 18, 19, vii.2.59, R. D. Anderson (RDA). Showville, 1 8, 1 9, viii.1.59, R. D. Anderson (RDA). Cache County. Benson, Logan Meadows, 13, viii.7.52, Knowlton & Wood (USNM). Wasatch Range, Beaver Mtn., 7700 feet, 1 8, vii.6.52, B. Malkin (FM). Juab County. Callao, 1 8, 1 9, xii.27.57, R. D. Anderson (RDA). Sevier R., at Hwy. 91, 1 8, 1 9, vi. 11.59, G. Musser (RDA). Kane County. Zion Ntl. Pk., 2 m. E., 2 9, vi.21.59, R. D. Anderson (RDA). Piute County. Greenwich, 5 m. S., 1 9, viii.3.64, H. B. Leech (CAS). Rich County. Bear L., 18, vii.4.59, R. D. Anderson (RDA). Salt Lake County. Saltair, 1, vi. 15.41, H. P. Chandler (CAS). Salt Lake, 1, iv.7.— ; vi.13.— , Hubbard & Schwarz (USNM). Salt Lake City, 1 9, vii. 14.52, B. Malkin & S. Mulaik (FM). Salt Lake City, Jordan R., 3 8, 1 9, iii.19.58; 2 8, 19, iv.20.57, R. D. Anderson (RDA). Summit County. Kimball Jet., 3 m. W., 1 9, v.12.57, R. D. Anderson (RDA). Pineview, 1 m. E, 1 5, vii.2.59, R. D. Anderson (RDA). Tooele County. Mills, 3 m. S, 2 3,3?, v.2.58, R. D. Anderson (RDA). Unita County. County, 2 2 , G. E. Wallace (CNG). Utah County. Goshen, 1 m. W., 2 J, iv.27.58, R. D. Anderson (RDA). Provo, 6, x.10.36, H. P. Chandler (CAS). Utah L., 13, vi.10.— , Hubbard & Schwarz (USNM);
1 $ y vi.19.— (ANSP). Utah L., American Fork, 1 2, iv.23.58, R. D. Anderson (RDA). Washington County. Hurricane, 1 2, vi.22.59, R. D. Anderson (RDA). Near Central, Santa Clara Ck., 23,25, v.10.58, R. D. Anderson (RDA). New Harmony, 1 5, 3 2, x.11.64, R. D. Anderson (RDA). St. George, 1, vii.— .— , Wickham (USNM); 12, A. M. Woodbury (USNM); 2 5, 5 2, iv.— .25 (USNM); 1 m. W., 3 5, 3 2, xii.29.59, G. Musser (RDA). Virgin R., 1 5 , 2 2, G. Weldt (AMNH). Weber County. Ogden, 8, vii. 3.—, Hubbard & Schwarz (USNM). WASHINGTON. Adams County. Ritzville, 4, viii. 29.24, D. Dunavan (USNM); 2, viii.10.21, M. C. Lane (CAS). Benton County. Pasco, 4 $, iii.26.60, J. Shepard (ORES). Chelan County. Leavenworth, 8, v. 29.42, J. G. Needham (CNL). Douglas County. Stratford, 2, ix.3. 20, M. C. Lane (USNM). Grant County. Vantage, E. Side Columbia R.,
2 2, vii. — .53, Crabtree (FM). Jefferson County. Pt. Townsend, 15,12 (AMNH). King County. Issoquady, 1 5, ii.5.41, H. J. Jensen (BERK). Seattle, 1, iv.3.72 (CNL). Skykomish R., 1, vi.1.92 (USNM). Seattle, 1 2, iv. 19.45, H. J. Jensen (BERK); 1 2, v.12.— (ANSP). Kitsap County. Winslow, 1, ix.10.— , Wickham (USNM). Kittitas County. Ellensburg, 1 2, ix.26.54, B. Malkin (FM). Vantage, 8 m. N., Whiskey Dick Can., 1 2, viii.4.54, B. Malkin (FM). Okonagan County. Okonagan, 4, iv.25.29, M. D. Leonard (CNL). Pierce County. Buckley, 1, v.28.42, J. G. Needham (CNL). Ft. Lewis, 24, vii. 7.51, R. Schuster (BERK). Snohomish County. Chase Lake, 7 5,7 2, viii.23.54; 2 5, 1 2, ix.25.52; 15,12, v.4.52; 15,12, x.17.54, B. Malkin (FM). Sultan, 15,12, vi.1.53, B. Malkin & D. Boddy (FM). Walla Walla County. College Place, 1 5, v.2.45, W. Blehm (MCHS). Walla Walla, 1 5, ix.2.26, M. W. Stone (ORES). Whitman County. Pullman, 3, viii.17.01, C. Y. Piper (USNM); 1, vi.20.40, C. Yocum (ISU). Yakima County. Toppenish, 1, iii.20.25; 2, vii.20-27.24, D. Dunavan (USNM). Washington County. Terry, 50, Morrison (USNM). East Washington, 2, C. V. Piper (USNM). WYOMING. Albany County. Rock Creek, 1, viii.24.41, H. P. Chandler (CAS). Converse County. Douglas, 4.5 m. W., 1 5,1 2, vii.27.64, H. B. Leech (CAS). Crook County. Devils Tower N. M., 4 m. S., 5 5 , 6 2, viii.4.52, B. Malkin (FM). Goshen County. Ft. Laramie, Laramie R., 1 5, 4 2, vii.29.64, H. B. Leech (CAS). Ft. Laramie, Natl. His. Site, 1 5, 1 2, viii.19.65, H. B. Leech (CAS). Ft. Laramie, 5.3 m. E., 3 2, viii.18.65, H. B. Leech; Jay Jay Em, 2.5 m. N., 3 2, viii.20.65, H. B. Leech (CAS). Lingle, 3.8 m. E., 15, viii.19.65, H. B. Leech (CAS). Johnson County. Crazy Woman Ck., S. Fork, 1 5 ,2 2 , vii.26.64, H. B. Leech (CAS) . Laramie County. Cheyenne, 12.5 m. N.E., Lodgepole Ck., 3 5,2 2, viii.18.65, H. B. Leech (CAS). Natrona County. Natrona, 2.5 m. S.E., 1 2, viii.20.65, H. B. Leech (CAS). Midwest, 8.5 m. N.W., Duquot Ck., 15, 2 2, vii.27.64, H. B.
Leech (CAS). Niobrara County. Manville, 3 m. W, 2 i, viii.20.65, H. B. Leech (CAS). Platte County. Glendo, 2 m. S., Horseshoe Ck., 1 $, 2 5, vii.29.64, H. B. Leech (CAS). Sweetwater County. Farson, 1 $, viii.22.65, H. B. Leech (CAS). Green River, 1 <$, 1933, Wickham (USNM). Rock Springs, 10 m. W., 1 $, viii.23.65, H. B. Leech (CAS).
MEXICO. — BAJA CALIFORNIA. La Suerte, 2 S, vi.4.63, R. K. Benjamin (CAS) ; on southern slope of Sierra San Pedro Martir, Hamilton Ranch, 8 m. upstream near Colonia Guerro, 1 2, iv.23.63, H. B. Leech (CAS). San Felipe, 1, J. R. Sherman, Jr. (USNM). Tijuana, 45 km. S., 1 5,1 9, viii.21.53, L. A. Stange (DAV).
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bibliographic citation
Zimmerman, J.R. 1970. A Taxonomic Revision of the aquatic beetle genus Laccophilus (Dytiscidae) of North America. Memoirs of the American Entomological Society vol. 26. Philadelphia, USA