Truncatelloidea Gray 1840

Figures 1-6. Daphniola eptalophos sp n. photographed in ethanol. Apical view 1, alive specimen carrying egg capsules with an embryo on last body whorl and inside umbilicus (photographed in water) 2 operculum 3 soft body, head with tentacles and penis in situ 4-5 penis 6. A background square represents 1 mm2 in Figs 1, 4, 5. Scale bar 1 mm and 0.5 mm in Figs 2 and 3 respectively. Black arrow points the penis in Figs 4-5 and the outgrowth of penis in Fig. 6.

Figure 5.Bithynia forcarti sp. n. a shell,frontal view b shell,lateral view.

Figures 1–15.Cyclostremiscus mohicanus sp. n. types 1 holotype MZSP 106551 (W 2.8 mm), apical view 2 same, umbilical view 3 same, apertural view 4 same, apertural-slightly apical view 5, same, apertural-slightly umbilical view 6 paratype MZSP 106552 #1, apical view (W 1.8 mm) 7 same, umbilical view 8 #2, apical view (W 1.7 mm) 9 #1, apertural-slightly apical view 10 same, apertural view 11 #3, SEM, apertural-slightly apical view (W 1.7 mm) 12 same, apertural view 13 same, apertural-slightly umbilical view 14 #4, SEM, apical view (W 1.5 mm) 15 same, detail of apical region, scale 0.1 mm.

Figure 2.Karucia sublacustrina sp. n. a–b shells (a holotype b paratype) c head with penis in situ (arrow showing outgrowth on the left side of penis) d penis. Abbreviations: e eye, p penis, s snout, t tentacle.

Figure 3.Shells, opercula and radula, Pyrgulopsis licina sp. n. A Holotype, USNM 850347 B, C Shells, USNM 1204732, USNM 1188732 D, E Opercula (outer, inner sides), USNM 850348 F Portion of radular ribbon, USNM 850348 G Central teeth, USNM 850348 H Lateral and inner marginal teeth, USNM 850348. Scale bars A–C 1.0 mm; D, E 100 µm; F, H 10 µm; G 2 µm.

Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis licina sp. n., USNM 850346 C, D Pyrgulopsis perforata sp. n., BellMNH 20891 E, F Pyrgulopsis sanchezi sp. n., USNM 883361 G, H Pyrgulopsis micrococcus, BellMNH 20663 I, J Pyrgulopsis turbatrix, USNM 860699 K, L Pyrgulopsis turbatrix, USNM 883373. Scale bars A–C 250 µm; D–L 500 µm. Pd penial duct Pf penial filament Pl penial lobe Tg terminal gland.

Figure 3.Hydrobioids collectedduring the survey in mainland and insular Greece. A Daphniola exigua (dorsal and ventral view) B Isimerope semele (dorsal and ventral view, Megali Vrysi) C Pseudoislamia balcanica (dorsal and ventral view, Ag. Sophia) D Pseudamnicola pieperi (Olympos) E Radomaniola cf. curta (spring of Louros river) F Radomaniola cf. curta (Ag. Sophia spring) G Trichonia trichonica H Pseudobithynia eubooensis. Scale bar 1 mm.

Figure 25.Drymaeus species. A Drymaeus (Mesembrinus) aurifluus (Pfeiffer, 1857), lectotype NHMUK 1975502 (H = 21.8) B Drymaeus (Mesembrinus) bugabensis (Martens, 1893), lectotype NHMUK 1901.6.22.958 (H = 26.3) C Drymaeus (Mesembrinus) dubius (Pfeiffer, 1853), lectotype NHMUK 1975519 (H = 27.1) D Drymaeus (Mesembrinus) (?) rufescens (J.E. Gray, 1825), lectotype of Bulimus erusbescens Pfeiffer, 1847 NHMUK 1975562 (H = 24.2) E Drymaeus (Mesembrinus) conicus da Costa, 1907, lectotype NHMUK 1907.11.21.32 (H = 17.5) F Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848), lectotype of Bulimus demotus Reeve, 1850 NHMUK 1975504 (H = 31.8) G Drymaeus (Mesembrinus) vexillum (Wood, 1828), lectotype of Bulimus keppelli Pfeiffer, 1853 NHMUK 1975538 (H = 33.2) H Drymaeus (Mesembrinus) attenuatus varicosus (Pfeiffer, 1853), lectotype NHMUK 1975466 (H = 35.5) I–K Drymaeus (Drymaeus) signifer (Pfeiffer, 1855), lectotype NHMUK 1975216 (H = 32.9). All enlarged.

Figure 3.Shells, opercula and radula, Pyrgulopsis marilynae n. sp. A Holotype, USNM 1135068 B Shell, USNM 1135067 C, D Opercula (outer, inner sides), USNM 1135067 E Portion of radular ribbon, USNM 1135067 F Central teeth, USNM 1135067 G Lateral and inner marginal teeth, USNM 1135067. Scale bars A, B 1.0 mm; C, D 100 µm; E 20 µm, F, G 10 µm.

Figures 7-10. Daphniola eptalophos sp. n. shell images from SEM Shell habitus 7, 8 protoconch 9 teleoconch 10 Scale bar 0.5 mm in Figs 7, 9 and 0.05 mm in Figs 8,10.

Figure 6.Shell of Bithynia starmuehlneri sp. n. a frontal view b lateral view c juvenile shell with operculum.

Figures 16–27.Episcynia itanhura sp. n. types 16 holotype MZSP 106553, umbilical view (W 2.0 mm) 17 same, apical view 18 same, apertural view 19 paratype MZSP 106556#1, umbilical view (W 2.2 mm) 20 same, apertural view 21 #2, SEM, umbilical-slightly apertural view (W 2.5 mm) 22 #3, SEM, umbilical view (W 2.1 mm) 23 #4, SEM, apertural-slightly umbilical view (W 2.5 mm) 24 #5, SEM, apertural view (W 2.6 mm) 25 #4, SEM, apertural view 26 #6, SEM, apical view (W 1.0 mm) 27 #6, detail of protoconch (scale 50µm).

Figure 3.Comparative morphology of shell and penis in Bithynia radomani Glöer & Pešić, 2007 (a–b) Bithynia montenegrina (Wohlberedt, 1901) (c–d) and Bithynia hambergerae A. Reischütz, N. Reischütz & P.L. Reischütz, 2008 (e–f): a, c, e = shell, b, d, f = penis.

Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis licina sp. n., USNM 850346 C, D Pyrgulopsis perforata sp. n., BellMNH 20891 E, F Pyrgulopsis sanchezi sp. n., USNM 883361 G, H Pyrgulopsis micrococcus, BellMNH 20663 I, J Pyrgulopsis turbatrix, USNM 860699 K, L Pyrgulopsis turbatrix, USNM 883373. Scale bars A–C 250 µm; D–L 500 µm. Pd penial duct Pf penial filament Pl penial lobe Tg terminal gland.

Figure 9.Shells, opercula and radula, Pyrgulopsis turbatrix. A Paratype, USNM 860699. B–D Shells, SBMNH uncat., USNM 853508, USNM 853510 E, F Opercula (outer, inner sides), USNM 860699 G Portion of radular ribbon, USNM 860699 H Central teeth, USNM 860699 I Lateral and outer marginal teeth, USNM 860699. Scale bars A–D 1.0 mm; E, F 250 µm; G 20 µm; H, I 10 µm.

Figure 3.Hydrobioids collectedduring the survey in mainland and insular Greece. A Daphniola exigua (dorsal and ventral view) B Isimerope semele (dorsal and ventral view, Megali Vrysi) C Pseudoislamia balcanica (dorsal and ventral view, Ag. Sophia) D Pseudamnicola pieperi (Olympos) E Radomaniola cf. curta (spring of Louros river) F Radomaniola cf. curta (Ag. Sophia spring) G Trichonia trichonica H Pseudobithynia eubooensis. Scale bar 1 mm.

Figure L18.Labels. i Bulimus deshayesi Pfeiffer, 1845 ii Bulimus devians Dohrn, 1863 iii Bulinus discrepans Sowerby I, 1833 iv Bulimus dubius Pfeiffer, 1853 v Bulimus dombeyanus ‘Férussac’ Pfeiffer, 1846.

Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis marilynae n. sp., USNM 883175 C, D Pyrgulopsis similis n. sp., USNM 1135065 E, F Pyrgulopsis gilae, BellMNH 20898. Scale bars A–F 200 µm. Dg2 dorsal gland along left edge Dg3 dorsal gland along right distal edge Ipg inner (left) penial gland Opg outer (right) penial gland Tg terminal gland Vg ventral gland.

Figures 11-13.Female genitalia of Daphniola species.Daphniola eptalophos sp. n. female genitalia drawn from the only one female individual found among dissected specimens 11 Daphniola louisifemale genitalia re-drawn from Falniowski and Szarowska (2000) 12 Daphniola exiguafemale genitalia re-drawn from Radoman (1983) 13 Scale bar 0.5 mm.

Figure 7.Bithynia mazandaranensissp. n. a, b shell c operculum d detail of the shell surface.

Figure 3.Comparative morphology of shell and penis in Bithynia radomani Glöer & Pešić, 2007 (a–b) Bithynia montenegrina (Wohlberedt, 1901) (c–d) and Bithynia hambergerae A. Reischütz, N. Reischütz & P.L. Reischütz, 2008 (e–f): a, c, e = shell, b, d, f = penis.

Figure 2.Map showing the distribution of mtDNA clades A–F with color codes matching those in Fig. 1.

Figure 1.Map showing the administrative units of Greece where “hydrobioid” localities were sampled during the fieldwork of this study.

Figure 50.Drymaeus species. A–C Drymaeus (Drymaeus) acervatus (Pfeiffer, 1857), lectotype NHMUK 1975461 (H = 41.5) D–F Drymaeus (Drymaeus) dunkeri (Pfeiffer in Philippi 1846), lectotype NHMUK 1975512 (H = 34.8) G–I Drymaeus (Drymaeus) lattrei (Pfeiffer in Philippi 1846), lectotype NHMUK 1975555 (H = 42.6). All enlarged.