The Ericales are a large and diverse order of dicotyledons. Species in this order have considerable commercial importance including for tea, persimmon, blueberry, kiwifruit, Brazil nuts, argan, and azalea. The order includes trees, bushes, lianas, and herbaceous plants. Together with ordinary autophytic plants, the Ericales include chlorophyll-deficient mycoheterotrophic plants (e.g., Sarcodes sanguinea) and carnivorous plants (e.g., genus Sarracenia).
Many species have five petals, often grown together. Fusion of the petals as a trait was traditionally used to place the order in the subclass Sympetalae.[2]
Mycorrhizal associations are quite common among the order representatives, and three kinds of mycorrhiza are found exclusively among Ericales (namely, ericoid, arbutoid and monotropoid mycorrhiza). In addition, some families among the order are notable for their exceptional ability to accumulate aluminum.[3]
Ericales are a cosmopolitan order. Areas of distribution of families vary largely - while some are restricted to tropics, others exist mainly in Arctic or temperate regions. The entire order contains over 8,000 species, of which the Ericaceae account for 2,000-4,000 species (by various estimates).
The most commercially used plant in the order is tea (Camellia sinensis) from the family Theaceae. The order also includes some edible fruits, including kiwifruit (esp. Actinidia deliciosa), persimmon (genus Diospyros), blueberry, huckleberry, cranberry, Brazil nut, and Mamey sapote. The order also includes shea (Vitellaria paradoxa), which is the major dietary lipid source for millions of sub-Saharan Africans. Many Ericales species are cultivated for their showy flowers: well-known examples are azalea, rhododendron, camellia, heather, polyanthus, cyclamen, phlox, and busy Lizzie.
Primula rosea from the Primulaceae
These families are recognized in the APG III system[1] as members of the Ericales:
Likely phylogenetic relationships between the families of the Ericales:[5]
Ericales ericoidsCyrillaceae
Ericaceae
Clethraceae
sarracenioidsRoridulaceae
Actinidiaceae
Sarraceniaceae
styracoidsStyracaceae
Diapensiaceae
Symplocaceae
Theaceae
Pentaphylacaceae
primuloidsPrimulaceae
Ebenaceae
Sapotaceae
polemonioidsPolemoniaceae
Fouquieriaceae
Lecythidaceae
Mitrastemonaceae
balsaminoidsMarcgraviaceae
Tetrameristaceae
Balsaminaceae
These families are not recognized in the APG III system[1] but have been in common use in the recent past:
These make up an early diverging group of asterids.[6] Under the Cronquist system, the Ericales included a smaller group of plants, which were placed among the Dilleniidae:
The Ericales are a large and diverse order of dicotyledons. Species in this order have considerable commercial importance including for tea, persimmon, blueberry, kiwifruit, Brazil nuts, argan, and azalea. The order includes trees, bushes, lianas, and herbaceous plants. Together with ordinary autophytic plants, the Ericales include chlorophyll-deficient mycoheterotrophic plants (e.g., Sarcodes sanguinea) and carnivorous plants (e.g., genus Sarracenia).
Many species have five petals, often grown together. Fusion of the petals as a trait was traditionally used to place the order in the subclass Sympetalae.
Mycorrhizal associations are quite common among the order representatives, and three kinds of mycorrhiza are found exclusively among Ericales (namely, ericoid, arbutoid and monotropoid mycorrhiza). In addition, some families among the order are notable for their exceptional ability to accumulate aluminum.
Ericales are a cosmopolitan order. Areas of distribution of families vary largely - while some are restricted to tropics, others exist mainly in Arctic or temperate regions. The entire order contains over 8,000 species, of which the Ericaceae account for 2,000-4,000 species (by various estimates).