Diagnostic Description
provided by Fishbase
Diagnosed from its congeners n Baltic and Black Sea basins by having following characters: last simple dorsal ray slightly thickened, with 0-30 small serrae posteriorly along median third; flexible segmented part of last simple dorsal ray about 50% of ray length; lower lip thick, with a pendulous median lobe; lateral line with 52-70 scales (usually 59-62); 11-12 scale rows between lateral line and dorsal fin origin; pelvic fin origin slightly to clearly behind dorsal fin origin; scales with free posterior part rounded; and scales without epithelial crests (Ref. 59043).
Life Cycle
provided by Fishbase
Eggs are released in holes dug by females with their anal fins. Probably spawning takes place in 2-3 batches during a spawning season (Ref. 59043).
Biology
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Inhabits small rivers with moderate to fast flowing water in piedmont and montane zones at 200-600 m above sea level, preferring stone, gravel, pebble or sand bottom and river stretches with rapids and deep holes. Undergoes upriver migration to shallow, smaller tributaries during the start of the spawning season. Does not migrate for long distances, usually moving along main course of stream during feeding season. Preys on benthic invertebrates. Breeds in shallow areas over pebble bottom with fast current and clear water with high oxygen concentration. Spawns mainly at night during sunset. Slow decline of population is expected to continue due to on-going economic development (Ref. 59043).