Rubus nagasawanus Koidz.

Shrubs scrambling. Branches robust, grayish soft hairy and tomentose, with reddish glandular hairs 2–3 mm and few prickles; prickles ca. 2 mm, curved, base dilated; petiole 1–2 cm, tomentose, with minute prickles and glandular hairs; stipules free, 8–10 mm, margin lacerate to pinnatipartite; lobes linear, with soft hairs and intermixed sparse, glandular hairs; blade suborbicular or broadly ovate, 4–6 × 4.5–7 cm, 5-veined, veins prominent abaxially, slightly impressed adaxially, abaxially gray or grayish yellow tomentose, adaxially dark and slightly hirsute especially along veins, base broadly to deeply cordate, margin 5-lobed, unevenly sharply serrate, apex acute or obtuse. Inflorescences terminal, cymose panicles, much branched, 8–15(–20) cm, many flowered; rachis and pedicels densely tomentose; bracts 5–8 mm, margin lacerate to pinnatipartite; lobes linear to linear-lanceolate, puberulous, with few glandular hairs. Pedicel 1–2 cm. Flowers ca. 1 cm in diam. Calyx yellowish tomentose, with sparse, intermixed glandular hairs; tube cupular, ca. 3 mm, unarmed; sepals triangular-ovate, 6–7 × 3–4 mm, apex acute to acuminate, usually undivided. Petals white, spatulate-orbicular to obovate, 3–4 × 2–3 mm, pubescent, base shortly clawed. Stamens many; filaments 3–4 mm, glabrous. Pistils 4–5 mm, glabrous. Aggregate fruit subglobose, 7–9 mm in diam., glabrous, enclosed in calyx; pyrenes 2–3 mm, slightly foveolate. Fl. summer, fr. autumn. 2n = 28*.

E Taiwan [Indonesia (Java, Sumatra), Philippines].

Thickets; 700--2500 m.

Rubus alceifolius Poiret var. emigratus Koidzumi (1913), not Focke (1904); R. formosensis Matsumura (1901), not Kuntze (1879); R. polyanthus H. L. Li; R. tephrodes Hance var. setosissimus Koidzumi (1939), not Handel-Mazzetti (1933).