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Swamp White Oak

Quercus bicolor Willd.

Comments

provided by eFloras
Putative hybrids between Quercus bicolor and Q . macrocarpa are common in areas of contact. The hybrids tend to have more deeply lobed leaves and varying degrees of development of awns as a fringe along the margin of the acorn cup. Such characteristics occur sporadically throughout many populations of Q . bicolor ; in some cases they may occur because of subtle introgression.

The Iroquois used Quercus bicolor in the treatment of cholera, broken bones, consumption, and as a witchcraft medicine (D. E. Moerman 1986).

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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Description

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Trees , deciduous, to 30 m. Bark dark gray, scaly or flat-ridged. Twigs light brown or tan, 2-3(-4) mm diam., glabrous. Buds light or dark brown, globose to ovoid, 2-3 mm, glabrous. Leaves: petiole (4-)10-25(-30) mm. Leaf blade obovate to narrowly elliptic or narrowly obovate, (79-)120-180(-215) × (40-)70-110(-160) mm, base narrowly cuneate to acute, margins regularly toothed, or entire with teeth in distal 1/2 only, or moderately to deeply lobed, or sometimes lobed proximally and toothed distally, secondary veins arched, divergent, (3-)5-7 on each side, apex broadly rounded or ovate; surfaces abaxially light green or whitish, with minute, flat, appressed-stellate hairs and erect, 1-4-rayed hairs, velvety to touch, adaxially dark green, glossy, glabrous. Acorns 1-3(-5) mm, on thin axillary peduncle (20-)40-70 mm; cup hemispheric or turbinate, 10-15 mm deep × 15-25 mm wide, enclosing 1/2-3/4 nut, scales closely appressed, finely grayish tomentose, those near rim of cup often with short, stout, irregularly recurved and sometimes branched, spinose awns emerging from tubercle; nut light brown, ovoid-ellipsoid or oblong, (12-)15-21(-25) × 9-18 mm, glabrous. Cotyledons distinct. 2 n = 24.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
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Flora of North America Editorial Committee
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Distribution

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Ont., Que.; Ala., Conn., Del., Ill., Ind., Iowa, Ky., Maine, Md., Mass., Mich., Minn., Mo., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., Tenn., Vt., Va., W.Va., Wis.
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of North America @ eFloras.org
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Flora of North America Editorial Committee
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Flowering/Fruiting

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Flowering in spring.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Habitat

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Low swamp forests, moist slopes, poorly drained uplands; 0-1000m.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of North America @ eFloras.org
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Synonym

provided by eFloras
Quercus bicolor var. angustifolia Dippel; Q. bicolor var. cuneiformis Dippel; Q. bicolor var. platanoides A. de Candolle; Q. platanoides (Lamarck) Sudworth
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 3 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
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Broad-scale Impacts of Fire

provided by Fire Effects Information System Plants
More info for the terms: fuel, prescribed burn, swamp

A prescribed burn on an Indiana savannah did not kill swamp white oak
saplings and larger trees [1]. However, trees did not sprout following
the burn. Average fuel loads were 560 g/sq m before the fire and from
400 to 650 g/sq m 1 year after the fire.

Fires during the dormant season are less damaging to oaks because of
lowered ambient temperatures and the tree's physiological state [32].
Crooked trees may be killed more easily than straight trees if the
crooked trees are leaning towards the flames. Overstocked stands may
suffer more damage from fire due to reduced vigor and size of
individuals [32]. Fire appears to affect acorn crops only in that,
dying trees tend to produce a massive crop. Acorns themselves are
easily destroyed by fire because of high moisture content [32].
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Common Names

provided by Fire Effects Information System Plants
swamp white oak
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Description

provided by Fire Effects Information System Plants
More info for the terms: fruit, tree

Swamp white oak is a native deciduous tree that reaches heights of 50 to
70 feet (15-20 m) and diameters of 2 to 3 feet (0.6-1 m) [31]. It has a
limby bole and an open, irregularly shaped crown. Its bark is flakey
and grey. Its leaves resemble those of chestnut (Castanea spp.); they
are shallowly lobed with serrate margins [22]. The fruit is an acorn
0.75 to 1.25 inches (2-3 cm) long. A mossy-like fringed cup covers from
one-third to one-half of the acorn [10]. Acorns are one seeded (rarely
two) and form singly or in clusters [25].
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Distribution

provided by Fire Effects Information System Plants
Swamp white oak occurs mainly in the midwestern states from Iowa,
southern Wisconsin, Michigan, and Ohio south to northern Kentucky.
Isolated populations occur in Minnesota, New England, Quebec, Ontario,
Tennessee, and North Carolina [18].
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Ecology

provided by Fire Effects Information System Plants
More info for the terms: fire regime, tree

All oaks can resprout from stems when top-killed by fire. The ability
to sprout decreases with an increase in age and tree size [33]. Many
seedlings develop an "S"-shaped crook in their stems, which protects
dormant buds from fire heat and enables seedlings to sprout [32]. With
repeated fire stems become calloused. This tissue is filled with
dormant buds that resprout.

FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find FIRE REGIMES".
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Management Considerations

provided by Fire Effects Information System Plants
More info for the terms: basal area, litter

Fire can reduce litter depth, allowing oak seedlings to become
established [32]. Fire can also reduce stocking rates of other species,
allowing oak species to increase in basal area. Fire can induce
vigorous sprouting from older root stock, which may be a preferred
reproductive method [32].
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Growth Form (according to Raunkiær Life-form classification)

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More info on this topic.

More info for the term: phanerophyte

Phanerophyte: Mesophanerophyte
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat characteristics

provided by Fire Effects Information System Plants
More info for the terms: forest, swamp

Swamp white oak occurs in river bottomlands, depressions, along
streamsides, swamp borders, and on moist peaty flats [10,20]. It is a
minor component in tamarack (Larix laricina) swamps of southwestern
Michigan [16]. Along the Ohio shores of Lake Erie, swamp white oak
grows in Toledo soil, a very poorly drained, silty clay. It also grows
on Nappanee soils, which are somewhat poorly drained silt loams [13].
Along the Kankakee River on the Illinois and Indiana border, swamp white
oak is a major overstory component of the floodplain forest. Here the
soils are highly permeable, frequently flooded sandy loams [21]. In
Quebec, swamp white oak occurs on sandy and loamy sand alluvium between
68 and 87 feet (22.6 and 28.9 m) in elevation [30].

Plant associates include pin oak (Quercus palustris), northern red oak
(Q. rubra), hickory (Carya spp.), beech (Fagus spp.), sweetgum
(Liquidambar styraciflua), yellow-poplar (Liriodendron tulipifera), red
maple (Acer rubrum), silver maple (A. saccharinum), green ash (Fraxinus
pennsylvanicum), tamarack, dogwood (Cornus spp.), sumac (Rhus spp.),
serviceberry (Amelanchier spp.), grape (Vitis spp.), holly (Ilex spp.),
and viburnum (Viburnum spp.) [3,5,9,12,30].
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Cover Types

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More info on this topic.

This species is known to occur in association with the following cover types (as classified by the Society of American Foresters):

26 Sugar maple - basswood
14 Northern pin oak
38 Tamarack
39 Black ash - American elm - red maple
52 White oak - black oak - northern red oak
53 White oak
59 Yellow poplar - white oak - northern red oak
60 Beech - sugar maple
62 Silver maple - American elm
64 Sassafras - persimmon
65 Pin oak - sweet gum
92 Sweetgum - willow oak
94 Sycamore - sweetgum - American elm
108 Red maple
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Ecosystem

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This species is known to occur in the following ecosystem types (as named by the U.S. Forest Service in their Forest and Range Ecosystem [FRES] Type classification):

FRES15 Oak - hickory
FRES17 Elm - ash - cottonwood
FRES18 Maple - beech - birch
FRES19 Aspen - birch
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Plant Associations

provided by Fire Effects Information System Plants
More info on this topic.

This species is known to occur in association with the following plant community types (as classified by Küchler 1964):

More info for the term: forest

K098 Northern floodplain forest
K099 Maple - basswood forest
K100 Oak - hickory forest
K101 Elm - ash forest
K102 Beech - maple forest
K103 Mixed mesophytic forest
K106 Northern hardwoods
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cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Immediate Effect of Fire

provided by Fire Effects Information System Plants
More info for the terms: swamp, top-kill

Severe fires can top-kill swamp white oak [33]. Moderate fires may kill
seedlings and saplings, but older trees usually survive. Fire-damaged
surviving trees are susceptible to disease and insect attack.
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Importance to Livestock and Wildlife

provided by Fire Effects Information System Plants
More info for the term: cover

Swamp white oak acorns are an important food for wildlife such as
squirrels, white-tailed deer, beaver, black bear, and a variety of birds
[3,12,24]. It provides cover for birds and mammals [6].
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Life Form

provided by Fire Effects Information System Plants
More info for the term: tree

Tree
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Management considerations

provided by Fire Effects Information System Plants
Swamp white oak is a component of forested wetlands, many of which are
being converted to agricultural lands and subdivisions [7]. Oaks are
susceptible to many insect pests, fungi, cankers, and wilts. Refer to
Solomon and others [34] for information on how to recognize and control
these diseases [34]. Oak species can suffer from what is known as "oak
decline." This is when trees die or limbs die back due to environmental
stresses [35].
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Nutritional Value

provided by Fire Effects Information System Plants
More info for the term: swamp

Percent oven-dry weight nutrient values for swamp white oak leaves are
as follows [4]:

nitrogen 2.02
potassium 1.20
phosphorous 0.26
calcium 1.07
magnesium 0.31
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Occurrence in North America

provided by Fire Effects Information System Plants
CT DE IL IN IA KS KY ME MD MA
MI MN MO NH NJ NY NC OH PA RI
TN VT VA WV WI ON PQ
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Phenology

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More info on this topic.

Swamp white oak acorns ripen from August through December [25].
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Plant Response to Fire

provided by Fire Effects Information System Plants
Swamp white oak stems can resprout following fire. Sprouts can grow as
much as 3 to 6 feet (1-3 m) per year for the first 2 to 3 postfire years
[33].
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Post-fire Regeneration

provided by Fire Effects Information System Plants
More info for the term: seed

off-site colonizer; seed carried by animals or water; postfire yr 1&2
survivor species; on-site surviving root crown
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Regeneration Processes

provided by Fire Effects Information System Plants
More info for the terms: fruit, seed, swamp

Sexual: Swamp white oak reproduces by seed, which mature in 1 year
[31]. Good seed crops are produced every 4 to 7 years, but many acorns
are infested by insects [33]. Acorns must be collected shortly afer
falling to prevent early germination. Viability can be tested by
dumping acorns into water. Those that float are not viable. Acorns
cannot be stored for more than a few months. Cleaned seed averages 120
per pound (108/kg). One hundred pounds of fruit will average between 60
and 75 pounds (54-67.5/kg) of seed [25]. Seedlings grow slowly at less
than 6 inches (15 cm) per year [33].

Vegetative: Swamp white oak can sprout from its trunk [33].
license
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Successional Status

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More info on this topic.

More info for the terms: swamp, tree

Swamp white oak is intermediate in shade tolerance but not very drought
tolerant [20]. It is a dominant tree in wetlands on infertile to
fertile soils of oak ecosystems in southeastern Michigan [2]. Without
disturbance elm (Ulmus americana)-ash (Fraxinus spp.)-cottonwood
(Populus spp.) types will convert to oak-dominated types that include
swamp white oak [23]. White oak (Quercus alba) forests of southern Ohio
(of which swamp white oak is a component) will progress towards hickory
and beech forests if undisturbed [5].
license
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Taxonomy

provided by Fire Effects Information System Plants
More info for the term: swamp

The currently accepted scientific name of swamp white oak is Quercus
bicolor Willd. (Fagaceae) [10]. There are no recognized varieties or
forms. Swamp white oak hybridizes with the following [10,14]:

Q. alba (Q. X jackiana Schneider)
Q. stellata (Q. X substellata Trel.)
Q. lyrata (Q. X humidicola E.J. Palmer)
Q. macrocarpa (Q. X Hillii Trel.)

Q. X introgressa is a hybrid cross formed with another hybrid parent
[28]. Q. meuhlenbergii is introgressed by Q. prinoides and Q. bicolor.
For more information on swamp white oak hybrids see Little [36].
license
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Value for rehabilitation of disturbed sites

provided by Fire Effects Information System Plants
Swamp white oak is planted on highway rights-of-way [15].
license
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bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Wood Products Value

provided by Fire Effects Information System Plants
More info for the terms: hardwood, swamp

Oak species account for one-third of the hardwood sawtimber volume in
the United States [34]. Swamp white oak is a heavy, hard wood that
machines well, but it can check and warp if not dried properly. It is
used for furniture, flooring, boxes, crates, barrels, kegs, ships and
boats [27].
license
cc-publicdomain
bibliographic citation
Snyder, S. A. 1992. Quercus bicolor. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Associated Forest Cover

provided by Silvics of North America
Swamp white oak is a consistent though mostly a minor component of hydromesophytic forest communities in which other species usually dominate. Tree species that commonly grow in association with swamp white oak are pin oak (Quercus palustris), sweetgum (Liquidambar styraciflua), red maple (Acer rubrum), silver maple (A. saccharinum), American elm (Ulmus americana), eastern cottonwood (Populus deltoides), sycamore (Platanus occidentalis), green ash (Fraxinus pennsylvanica), bur oak Quercus macrocarpa), shellbark and shagbark hickory (Carya laciniosa and C. ovata), blackgum (Nyssa sylvatica), black willow (Salix nigra), and American basswood (Tilia americana) (3,4,6).

Swamp white oak occurs in four forest cover types: Black Ash-American Elm-Red Maple (Society of American Foresters Type 39), Bur Oak (Type 42), Silver Maple-American Elm (Type 62), and Pin Oak-Sweetgum (Type 65). It is usually found singly in these types but occasionally may be abundant in small areas (6).

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Climate

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Within the range of swamp white oak, mean annual temperatures vary from 16° C (60° F) in Arkansas to 4° C (40° F) in southern Ontario. Extremes in temperature vary from 41° C (105° F) to -34° C (-30° F). Average annual precipitation is from 640 mm (25 in) in southeast Minnesota to 1270 mm (50 in) in northeast Arkansas. The frost-free period ranges from 210 days in the southern part of the growing area to 120 days in the northern part (4).

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Damaging Agents

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Windthrow may be a problem especially in recently thinned stands.

Disease and insects affecting swamp white oak are essentially the same as those found on white oak. Oak anthracnose can be damaging to individual trees but is generally not fatal. Swamp white oak is susceptible to the oak wilt fungus (Ceratocystis fagacearum) and in Illinois Phomopsis canker and Coniothyrium dieback were found on this oak. In addition, an Alternaria fungus was found on blighted petioles (4,7).

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Flowering and Fruiting

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Swamp white oak is monoecious; male and female flowers appear on the same tree in the spring at about the time leaves are one-third developed (May to June). The fruit, an acorn, matures in 1 year and is generally paired and home on slender stalks from 3 to 8 cm (1.25 to 3.25 in) long. The ovoid acorns, each 19 to 32 mm (0.75 to 1.25 in) long and 13 to 19 min (0.5 to 0.75 in) in diameter, fall during September and October.

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Genetics

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Two forms of swamp white oak have been described: a mesophytic form with leaves that are green and velvety on the lower surface and a more xerophytic form with leaves that are white-tomentulose beneath. The following six hybrids with swamp white oak are recognized: Quercus x jackiana Schneid. (Q. bicolor x alba); Q. x humidicola Palmer (Q. bicolor x lyrata); Q. x schuettei Trel. (Q. bicolor x macrocarpa) (1); Q. x introgressa P. M. Thomson (Q. bicolor x muehlenbergii x prinoides) (11); Q. x substellata Trel. (Q. bicolor x stellata); Q. x nessiana Palmer (Q. bicolor x virginiana). Swamp white oak also hybridizes with chestnut oak Quercus prinus) and English oak (Q. robur).

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Growth and Yield

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On the better drained lowland soils, the growth rate of swamp white oak is comparable to that of white oak. The root system is usually shallow, but the tree is relatively long lived-up to 300 years or more. Normally it is a mediumsized tree, 18 to 23 in (60 to 75 ft) in height and 61 to 91 cm (24 to 36 in) d.b.h., although trees up to 30 in (100 ft) tall and 213 cm (84 in) d.b.h. have been reported.

Swamp white oak normally grows in mixtures with other bottom-land species and is abundant only locally. Individual old growth trees may contain as much net volume as 3.4 m³ (600 fbm) but this is uncommon (4).

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Reaction to Competition

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The tree is classed as intermediate in tolerance to shade, and seedlings become established under moderate shade. Lowland forests in which swamp white oak grows are characterized by instability and successional uncertainty because of the variable effects of flooding, together with the presence of saturated soils. Swamp white oak may achieve dominance on the better drained lowland soils together with basswood, northern red oak (Quercus rubra), American beech (Fagus grandifolia), and sugar maple (Acer saccharum) (8). Once established, it is able to compete effectively with American elm, green ash, and black willow. Limited current evidence indicates clearcutting to be an adequate silvicultural system, particularly on the better sites (2,8).

In forest stands swamp white oak has a straight bole with ascending branches and a narrow crown. However, open-grown trees are generally poorly formed and often have persistent lower branches (4).

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Rooting Habit

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Seed Production and Dissemination

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Good crops of swamp white oak occur every 3 to 5 years, with light crops during intervening years. The minimum seed-bearing age is 20 years, optimum age is 75 to 200 years, and maximum age is usually 300 years. Because the seed of swamp white oak is not dormant, it germinates soon after falling. Seed collections should be made soon after ripening in order to delay early germination. These acorns are difficult to store without germination or loss of viability occurring. Sound acorns have a germinative capacity between 78 and 98 percent. Gravity, rodents, and water are the primary dispersing agents (4,10).

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Seedling Development

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Germination is hypogeal (10). After acorns germinate in the fall, roots continue to develop until growth is limited by low temperatures. Seedling establishment and early growth seem to be favored on the better drained lowland soils rather than on sites that are poorly drained or subjected to persistent flooding. In any case, adequate moisture and light are necessary for successful early development (4,8).

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Soils and Topography

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Throughout its range, swamp white oak is typically found on hydromorphic soils. These may be mineral soils that are imperfectly to poorly drained, as evidenced by high water tables and the presence of glei subsurface layers, or both; organic soils ranging from mucks (well decomposed) to peats (poorly decomposed) in which high water levels have favored organic accumulation; or alluvial soils underlain by a glei layer. These kinds of soils are associated with lands that are periodically inundated, such as broad stream valleys, low-lying fields, and the margins of lakes, ponds, or sloughs. Swamp white oak is not found where flooding is permanent (2,4,5,6,8). In general, the soils on which this oak most commonly is found are in the orders Entisols and Inceptisols.

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Special Uses

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The acorns are sweet, like others in the white oak group, and are eaten by squirrels and other rodents (9). In a study in Wisconsin, swamp white oak acorns were found to make up 27 percent of the diet of wild ducks. Several nongame bird species include these acorns in their diet (4).

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Vegetative Reproduction

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Like most oaks, swamp white oak produces seedling sprouts or stump sprouts when the top is cut or killed. The frequency of sprouting declines, however, with increasing d.b.h. (8):



D.b.h. classes Stumps likely to sprout cm in percent 15 to 27 6 to 10 75 27 to 39 11 to 15 30 39 to 52 16 to 20 10 52+ 20+ 5

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Brief Summary

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Fagaceae -- Beech family

Robert Rogers

Swamp white oak (Quercus bicolor) is a mediumsized tree of the north central and northeastern mixed forests. It is found in lowlands, along edges of streams, and in swamps subject to flooding. It is rapid growing and long lived, reaching 300 to 350 years. The hard strong wood is commercially valuable and is usually cut and sold as white oak. Many kinds of wildlife eat the acorns, particularly ducks.

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Distribution

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Swamp white oak, a lowland tree, grows from southwestern Maine west to New York, southern Quebec, and southern Ontario, to central Michigan, northern Wisconsin, and southeastern Minnesota; south to Iowa and Missouri; east to Kentucky, Tennessee, Virginia, and New Jersey. It is scattered in North Carolina and northeastern Kansas. This species is most common and reaches its largest size in western New York and northern Ohio (4).


-The native range of swamp white oak.


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Quercus bicolor

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Quercus bicolor, the swamp white oak, is a North American species of medium-sized trees in the beech family. It is a common element of America's north central and northeastern mixed forests. It can survive in a variety of habitats. It forms hybrids with bur oak where they occur together in the wild.

Description

Quercus bicolor grows rapidly and can reach 18 to 24 meters (60 to 80 feet) tall with the tallest known reaching 29 m (95 ft) and lives up to 285 years.[6] The bark resembles that of the white oak. The leaves are broad ovoid, 12–18 centimetres (4+34–7 inches) long and 7–11 cm (2+344+14 in) broad, always more or less glaucous on the underside, and are shallowly lobed with five to seven lobes on each side, intermediate between the chestnut oak and the white oak. In autumn, they turn brown, yellow-brown, or sometimes reddish, but generally, the color is not as reliable or as brilliant as the white oak can be. The fruit is a peduncled acorn, 1.5–2 cm (5834 in), rarely 2.5 cm (1 in), long and 1–2 cm (3834 in) broad, maturing about six months after pollination.[7]

Swamp white oak may live up to 300 years.

Distribution and habitat

Swamp white oak, a lowland tree, occurs across the eastern and central United States and eastern and central Canada, from Maine to South Carolina, west as far as Ontario, Minnesota, and Tennessee with a few isolated populations in Nebraska and Alabama. This species is most common and reaches its largest size in western New York and northern Ohio.[8][9]

The swamp white oak generally occurs singly in four different forest types: black ash–American elm–red maple, silver maple–American elm, bur oak, and pin oak–sweetgum. Occasionally the swamp white oak is abundant in small areas. It is found within a very wide range of mean annual temperatures from 16 to 4 °C (61 to 39 °F). Extremes in temperature vary from 41 to −34 °C (106 to −29 °F). Average annual precipitation is from 640 to 1,270 millimetres (25 to 50 in). The frost-free period ranges from 210 days in the southern part of the growing area to 120 days in the northern part. The swamp white oak typically grows on hydromorphic soils. It is not found where flooding is permanent, although it is usually found in broad stream valleys, low-lying fields, and the margins of lakes, ponds, or sloughs. It occupies roughly the same ecological niche as pin oak, which seldom lives longer than 100 years, but is not nearly as abundant.

Uses

It is one of the more important white oaks for lumber production. The wood is similar to that of Q. alba and is not differentiated from it in the lumber trade. In recent years, the swamp white oak has become a popular landscaping tree due to its relative ease of transplanting.

Being in the white oak group, wildlife such as deer, bears, turkeys, ducks, and geese as well as other animals are attracted to this tree when acorns are dropping in the fall.

Cultivars

A mix of Quercus robur fastigiata x Quercus bicolor, named 'Nadler' or the Kindred Spirit hybrid oak, exists.

References

  1. ^ Kenny, L.; Wenzell, K.; Jerome, D. (2017). "Quercus bicolor". IUCN Red List of Threatened Species. 2017: e.T194069A111189345. doi:10.2305/IUCN.UK.2017-2.RLTS.T194069A111189345.en. Retrieved 18 November 2021.
  2. ^ "NatureServe Explorer". Retrieved 27 May 2021.
  3. ^ First described in Muhlenberg, Heinrich Ernst (1801). With remarks by Carl Ludwig Willdenow. "Kurze Bemerkungen über die in der Gegend von Lancaster in Nordamerika wachsenden Arten der Gattungen Juglans, Fraxinus und Quercus" [Short remarks on the plants growing in the region of Lancaster in North America from the genera Juglans, Fraxinus, and Quercus]. Der Gesellschaft Naturforschender Freunde zu Berlin, Neue Schriften. 3: 396.
  4. ^ "Quercus bicolor". Tropicos. Missouri Botanical Garden.
  5. ^ "Quercus bicolor". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew – via The Plant List. Note that this website has been superseded by World Flora Online
  6. ^ "Eastern OLDLIST of maximum tree ages".
  7. ^ Nixon, Kevin C. (1997). "Quercus bicolor". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 3. New York and Oxford – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  8. ^ Rogers, Robert (1990). "Quercus bicolor". In Burns, Russell M.; Honkala, Barbara H. (eds.). Hardwoods. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). Vol. 2. Retrieved 14 December 2009 – via Southern Research Station.
  9. ^ "Quercus bicolor". County-level distribution map from the North American Plant Atlas (NAPA). Biota of North America Program (BONAP). 2014.

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Quercus bicolor: Brief Summary

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Quercus bicolor, the swamp white oak, is a North American species of medium-sized trees in the beech family. It is a common element of America's north central and northeastern mixed forests. It can survive in a variety of habitats. It forms hybrids with bur oak where they occur together in the wild.

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