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Behavioral diversity

provided by EOL authors

It is extremely difficult to generalize about any of the behaviors or nesting habits of passerines, because as a group they are so diverse. Perching birds exhibit a bewildering array of plumages and colors derived from diverse keratin structures as well as ingested pigments, such as carotenoids (Gray, 1996). Many passerines, such as some Old World Flycatchers (Muscicapidae) and African Widowbirds (Viduinae) have extremely long tail feathers or highly modified plumes (Birds of Paradise: Paradisaeidae) used in courtship displays. Several groups such as the Wattlebirds of New Zealand (Callaeidae) and Honeyeaters (Meliphagidae) have fleshy, bright blue, red or yellow wattles on the face and neck. Perching birds build their nests generally out of sticks or grass on the ground, in trees, and in the case of Dippers (Cinclidae) in the banks of fast-flowing rivers. Many passerines migrate from their nesting grounds in the Nearctic and Palearctic to more equatorial regions, or from southern temperate regions north to the tropics. Parental care by both sexes is common in passerines, although in some highly dimorphic and predominantly lekking groups, such as manakins (Prum, 1994) and birds of paradise (Diamond, 1986), females alone provide for young and build the nest. Cooperative breeding, in which young birds delay breeding and assist other individuals (often parents) in raising young and defending the territory, is common in several passerine groups, such as Australian fairy wrens (Maluridae) and New World Jays (Corvidae; Brown, 1987; Edwards and Naeem, 1993). Some of the most elaborate singers in the bird world are passerines (Kroodsma and Miller, 1996). Some passerine birds are poisonous to the touch and are avoided as prey by indigenous peoples (Dumbacher et al., 1992).

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Edwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/
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Introduction

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The Passeriformes is the largest and most diverse commonly recognized clade of birds. The Passeriformes (or ‘passerine’ birds) are synonymous with what are commonly known as "perching birds"; this group also contains within it a major radiation commonly known as songbirds (oscine Passerines or Passeri). Of the 10,000 or so extant species of birds, over half (~5,300) are perching birds.

Perching birds have a worldwide distribution, with representatives on all continents except Antarctica, and reaching their greatest diversity in the tropics. Body sizes of passerines vary from about 1.4 kg in northern populations of Ravens (Corvus corax) to just a few grams. Perching birds include some of the most colorful and mysterious of all birds, such as birds of paradise from New Guinea and the bright orange Cock of the Rock from tropical South America. Because of their high diversity, generally small body size and relative ease of observation, collection and field study, perching birds have historically attracted the attention of a wide range of descriptive and experimental biologists, including systematists, behavioral ecologists, and evolutionary biologists.

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Edwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/
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Monophyly and Sister Group

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Historically, it is generally agreed that the Passeriformes constitute a monophyletic group. Raikow (1982) established this monophyly in an explicitly phylogeneticcontext. He noted that Passeriformes possess a suite of distinguishing characteristics, including a unique sperm morphology, a distinctive morphology of the bony palate, a simple yet functionally diverse foot with three toes forward and one (the hallux) oriented backwards, and a distinctive fore- (wing) and hindlimb musculature. There are few if any species which pose problems for avian systematists as to whether they are or are not passerines. Most of the controversy lies in relationships within the clade.

The sister group of the Passeriformes is not so much hotly contested as it is poorly resolved by existing data sets. Traditionally, Passeriformes have been considered closely related to a large group known as the “higher non-Passerines”. These include a number of clades such as cuckoos (Cuculiformes), hornbills, kingfishers and related lineages (Coraciiformes), and woodpeckers and relatives (Piciformes). Many of these groups possess a zygodactyl foot, a condition in which two toes point forward and two point backward. The sister relationship of Passeriformes to woodpeckers, the hornbill group and allies is reflected in Joel Cracraft’s phylogenetic hypothesis for major groups of birds based on cladistic interpretation of morphological and molecular characters (Cracraft, 1988). However, in the other major classification bearing on the relationships of perching birds, that based on DNA-DNA hybridization, Passeriformes appear as the sister group to a large, diverse group containing pigeons and doves (Columbiformes), cranes and rails (Gruiformes) and storks (Ciconiiformes)! These latter three groups share few obvious morphological characteristics with Passeriformes. However, the DNA hybridization tree links Passeriformes with these groups at a very deep level in the tree, rendering this result tenuous. A recent study of nuclear DNA sequences by Hackett et al. (2008) finds Psittaciformes (parrots) to be the sister group of passerines, with Falconidae (falcons) also close. Clearly, more work on the sister-group relationship of Passeriformes is needed, since this relationship will be the basis of any study seeking to identify whether or not Passeriformes are a particularly diverse group (e.g., Nee et al. 1992).

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Edwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/
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Species diversity, origin and biogeography

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Species diversity:The tradition of recognizing perching birds (Passeriformes) as the most diverse and rapidly radiating clade has been questioned because there are few obvious “key innovations” that should cause systematists to recognize Passeriformes over any other arbitrarily larger or smaller monophyletic group within birds (Raikow, 1986). One point that has been missed in debates on this issue is that the branch leading to the songbirds (oscines), a group comprising 80% of extant perching birds, is the longest internal branch on the DNA hybridization tree produced by Sibley and Ahlquist (1990). This branch has also been one of the few to be well resolved in applications of mtDNA sequences to higher level questions in birds, presumably because it is long. Given the large number of clades that will require names under phylogenetic taxonomy, perhaps the length of branches leading to particular clades should be one criterion whereby systematists decide which of the many clades to name.

Origin and biogeography of passerines:The temporal and geographic origin of passerine birds is obscure. Traditionally the group was thought to have originated in the Tertiary, at about the same time as extant orders of mammals. Some recent workers favor a later, Eocene origin (Feduccia, 1995; Wilson, 1989), but the DNA -DNA hybridization data again favors an earlier origin (Sibley and Ahlquist, 1990). Recently some of the oldest oscine fossils have been uncovered in Queensland, Australia (Boles, 1995); this and other paleobiogeographical data suggest that passerines may have in fact originated in the Southern hemisphere (Olson, 1989).

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Edwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/
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Passerine

provided by wikipedia EN

A passerine (/ˈpæsərn/) is any bird of the order Passeriformes (/ˈpæsərɪfɔːrmz/; from Latin passer 'sparrow' and formis '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds or songbirds, passerines are distinguished from other orders of birds by the arrangement of their toes (three pointing forward and one back), which facilitates perching.

With more than 140 families and some 6,500 identified species,[1] Passeriformes is the largest order of birds and among the most diverse orders of terrestrial vertebrates, representing 60% of birds.[2][3] Passerines are divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni (suboscines), and Passeri (oscines).[4][5] The passerines contain several groups of brood parasites such as the viduas, cuckoo-finches, and the cowbirds. Most passerines are omnivorous, while the shrikes are carnivorous.

The terms "passerine" and "Passeriformes" are derived from the scientific name of the house sparrow, Passer domesticus, and ultimately from the Latin term passer, which refers to sparrows and similar small birds.

Description

The order is divided into three suborders, Tyranni (suboscines), Passeri (oscines), and the basal Acanthisitti.[6] Oscines have the best control of their syrinx muscles among birds, producing a wide range of songs and other vocalizations, though some of them, such as the crows, do not sound musical to human beings. Some, such as the lyrebird, are accomplished imitators.[7] The New Zealand wrens are tiny birds restricted to New Zealand, at least in modern times; they were long placed in Passeri.

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Pterylosis or the feather tracts in a typical passerine

Most passerines are smaller than typical members of other avian orders. The heaviest and altogether largest passerines are the thick-billed raven[8] and the larger races of common raven, each exceeding 1.5 kg (3.3 lb) and 70 cm (28 in). The superb lyrebird and some birds-of-paradise, due to very long tails or tail coverts, are longer overall. The smallest passerine is the short-tailed pygmy tyrant, at 6.5 cm (2.6 in) and 4.2 g (0.15 oz).[9]

Anatomy

The foot of a passerine has three toes directed forward and one toe directed backward, called anisodactyl arrangement, and the hind toe (hallux) joins the leg at approximately the same level as the front toes. This arrangement enables passerine birds to easily perch upright on branches. The toes have no webbing or joining, but in some cotingas, the second and third toes are united at their basal third.

The leg of passerine birds contains an additional special adaptation for perching. A tendon in the rear of the leg running from the underside of the toes to the muscle behind the tibiotarsus will automatically be pulled and tighten when the leg bends, causing the foot to curl and become stiff when the bird lands on a branch. This enables passerines to sleep while perching without falling off.[10][11]

Most passerine birds have 12 tail feathers but the superb lyrebird has 16,[12] and several spinetails in the family Furnariidae have 10, 8, or even 6, as is the case of Des Murs's wiretail. Species adapted to tree trunk climbing such as woodcreeper and treecreepers have stiff tail feathers that are used as props during climbing. Extremely long tails used as sexual ornaments are shown by species in different families. A well-known example is the long-tailed widowbird.

Eggs and nests

The chicks of passerines are altricial: blind, featherless, and helpless when hatched from their eggs. Hence, the chicks require extensive parental care. Most passerines lay colored eggs, in contrast with nonpasserines, most of whose eggs are white except in some ground-nesting groups such as Charadriiformes and nightjars, where camouflage is necessary, and in some parasitic cuckoos, which match the passerine host's egg. The vinous-throated parrotbill has two egg colors, white and blue, to deter the brood parasitic common cuckoo.

Clutches vary considerably in size: some larger passerines of Australia such as lyrebirds and scrub-robins lay only a single egg, most smaller passerines in warmer climates lay between two and five, while in the higher latitudes of the Northern Hemisphere, hole-nesting species like tits can lay up to a dozen and other species around five or six. The family Viduidae do not build their own nests, instead, they lay eggs in other birds' nests.

Origin and evolution

The evolutionary history of the passerine families and the relationships among them remained rather mysterious until the late 20th century. In many cases, passerine families were grouped together on the basis of morphological similarities that, it is now believed, are the result of convergent evolution, not a close genetic relationship. For example, the wrens of the Americas and Eurasia, those of Australia, and those of New Zealand, look superficially similar and behave in similar ways, yet belong to three far-flung branches of the passerine family tree; they are as unrelated as it is possible to be while remaining Passeriformes.[a]

Advances in molecular biology and improved paleobiogeographical data gradually are revealing a clearer picture of passerine origins and evolution that reconciles molecular affinities, the constraints of morphology and the specifics of the fossil record.[14][15] The first passerines are now thought to have evolved in Gondwana (in the Southern Hemisphere) in the late Paleocene or early Eocene, around 50 million years ago.[5][15]

The initial split was between the New Zealand wrens (Acanthisittidae) and all other passerines (Eupasserine), and the second split involved the Tyranni (suboscines) and the Passeri (oscines or songbirds).[3] A rupture of the Gondwanan continent caused the core split of the Eupasseres, which were divided into these groups, one in Western Gondwana (Tyranni) and the other in Eastern Gondwana (Passeri).[2] Passeri experienced a great radiation of forms out of the Australian continent. A major branch of the Passeri, parvorder Passerida, expanded deep into Eurasia and Africa, where a further explosive radiation of new lineages occurred.[15] This eventually led to three major Passerida lineages comprising about 4,000 species, which in addition to the Corvida and numerous minor lineages make up songbird diversity today. Extensive biogeographical mixing happens, with northern forms returning to the south, southern forms moving north, and so on.[3]

Fossil record

Earliest passerines

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Male superb lyrebird (Menura novaehollandiae): This very primitive songbird shows strong sexual dimorphism, with a peculiarly apomorphic display of plumage in males.

Perching bird osteology, especially of the limb bones, is rather diagnostic.[16][17][18] However, the early fossil record is poor because the first Passeriformes were apparently on the small side of the present size range, and their delicate bones did not preserve well. Queensland Museum specimens F20688 (carpometacarpus) and F24685 (tibiotarsus) from Murgon, Queensland, are fossil bone fragments initially assigned to Passeriformes.[16] However, the material is too fragmentary and their affinities have been questioned.[19] Several more recent fossils from the Oligocene of Europe, such as Wieslochia, Jamna, Resoviaornis and Crosnoornis,[20] are more complete and definitely represent early passeriforms, although their exact position in the evolutionary tree is not known.

From the Bathans Formation at the Manuherikia River in Otago, New Zealand, MNZ S42815 (a distal right tarsometatarsus of a tui-sized bird) and several bones of at least one species of saddleback-sized bird have recently been described. These date from the Early to Middle Miocene (Awamoan to Lillburnian, 19–16 mya).[21]

Early European passerines

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Wieslochia fossil

In Europe, perching birds are not too uncommon in the fossil record from the Oligocene onward, but most are too fragmentary for a more definite placement:

  • Wieslochia (Early Oligocene of Frauenweiler, Germany)
  • Resoviaornis (Early Oligocene of Wola Rafałowska, Poland)
  • Jamna (Early Oligocene of Jamna Dolna, Poland)
  • Winnicavis (Early Oligocene of Lower Silesian Voivodeship, Poland)
  • Crosnoornis (Early Oligocene of Poland)
  • Passeriformes gen. et sp. indet. (Early Oligocene of Luberon, France) – suboscine or basal[b]
  • Passeriformes gen. et spp. indet. (Late Oligocene of France) – several suboscine and oscine taxa[23][18]
  • Passeriformes gen. et spp. indet. (Middle Miocene of France and Germany) – basal?[c]
  • Passeriformes gen. et spp. indet. (Sajóvölgyi Middle Miocene of Mátraszőlős, Hungary) – at least 2 taxa, possibly 3; at least one probably Oscines.[d]
  • Passeriformes gen. et sp. indet. (Middle Miocene of Felsőtárkány, Hungary) – oscine?[e]
  • Passeriformes gen. et sp. indet. (Late Miocene of Polgárdi, Hungary) – Sylvioidea (Sylviidae? Cettiidae?)[26]

That suboscines expanded much beyond their region of origin is proven by several fossil from Germany such as a broadbill (Eurylaimidae) humerus fragment from the Early Miocene (roughly 20 mya) of Wintershof, Germany, the Late Oligocene carpometacarpus from France listed above, and Wieslochia, among others.[17][15] Extant Passeri super-families were quite distinct by that time and are known since about 12–13 mya when modern genera were present in the corvoidean and basal songbirds. The modern diversity of Passerida genera is known mostly from the Late Miocene onwards and into the Pliocene (about 10–2 mya). Pleistocene and early Holocene lagerstätten (chronospecies and paleosubspecies.

American fossils

In the Americas, the fossil record is more scant before the Pleistocene, from which several still-existing suboscine families are documented. Apart from the indeterminable MACN-SC-1411 (Pinturas Early/Middle Miocene of Santa Cruz Province, Argentina),[f] an extinct lineage of perching birds has been described from the Late Miocene of California, United States: the Palaeoscinidae with the single genus Palaeoscinis. "Palaeostruthus" eurius (Pliocene of Florida) probably belongs to an extant family, most likely passeroidean.

Systematics and taxonomy

   

Acanthisitti – New Zealand wrens (1 family containing 4 species)

     

Tyranni – suboscines (16 families containing 1,356 species)

   

Passeri – oscines (125 families containing 5,158 species)

      Phylogenetic relationship of the suborders within the Passeriformes. The numbers are from the list published by the International Ornithologists' Union in January 2020.[1][29]

The Passeriformes is currently divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni (suboscines) and Passeri (oscines). The Passeri has been traditionally subdivided into two major groups recognized now as Corvida and Passerida respectively containing the large superfamilies Corvoidea and Meliphagoidea, as well as minor lineages, and the superfamilies Sylvioidea, Muscicapoidea, and Passeroidea but this arrangement has been found to be oversimplified. Since the mid-2000s, literally, dozens of studies have investigated the phylogeny of the Passeriformes and found that many families from Australasia traditionally included in the Corvoidea actually represent more basal lineages within oscines. Likewise, the traditional three-superfamily arrangement within the Passeri has turned out to be far more complex and will require changes in classification.

Major "wastebin" families such as the Old World warblers and Old World babblers have turned out to be paraphyletic and are being rearranged. Several taxa turned out to represent highly distinct lineages, so new families had to be established, some of them – like the stitchbird of New Zealand and the Eurasian bearded reedlingmonotypic with only one living species.[30] In the Passeri alone, a number of minor lineages will eventually be recognized as distinct superfamilies. For example, the kinglets constitute a single genus with less than 10 species today but seem to have been among the first perching bird lineages to diverge as the group spread across Eurasia. No particularly close relatives of them have been found among comprehensive studies of the living Passeri, though they might be fairly close to some little-studied tropical Asian groups. Nuthatches, wrens, and their closest relatives are currently grouped in a distinct super-family Certhioidea.

Taxonomic list of Passeriformes families

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New Zealand rock wren (Xenicus gilviventris), one of the two surviving species of suborder Acanthisitti

This list is in taxonomic order, placing related families next to one another. The families listed are those recognised by the International Ornithologists' Union (IOC).[1] The order and the division into infraorders, parvorders and superfamilies follows the phylogenetic analysis published by Carl Oliveros and colleagues in 2019.[29][g] The relationships between the families in the suborder Tyranni (suboscines) were all well determined but some of the nodes in Passeri (oscines) were unclear owing to the rapid splitting of the lineages.[29]

Suborder Acanthisitti

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Javan banded pitta (Hydrornis guajanus), an Old World suboscine.
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Andean cock-of-the-rock (Rupicola peruvianus) a New World suboscine

Suborder Tyranni (suboscines)

Infraorder Eurylaimides: Old World suboscines

Infraorder Tyrannides: New World suboscines
Parvorder Furnariida

Parvorder Tyrannida

Suborder Passeri (oscines)

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Male stitchbird or hihi (Notiomystis cincta) showing convergence with honeyeaters
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Male regent bowerbird (Sericulus chrysocephalus, Ptilonorhynchidae)
  • Infraorder Corvides – previously known as the parvorder Corvida[32]
  • Superfamily Orioloidea[h]
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Tiny goldcrest (Regulus regulus) belongs to a minor but highly distinct lineage of Passeri
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Reed warblers, such as this Blyth's reed warbler (Acrocephalus dumetorum), are now in the Acrocephalidae
  • Infraorder Passerides – previously known as the parvorder Passerida[32]
 src=
Eurasian blue tit (Cyanistes caeruleus) and its relatives stand well apart from rest of the Sylvioidea sensu lato
  • Parvorder Sylviida[k] – previously known as the superfamily Sylviodea[32]
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Brown-headed nuthatch (Sitta pusilla), nuthatches can climb downwards head-first
  • Superfamily Locustelloidea
 src=
Hermit thrush (Catharus guttatus), like many Muscicapoidea a stout and cryptic bird with complex vocalizations.
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Like these male (right) and female Gouldian finches (Erythrura gouldiae), many Passeroidea are very colorful
  • Superfamily Aegithaloidea
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Lesser striped swallow (Cecropis abyssinica), showing some apomorphies of its ancient yet highly advanced lineage.
  • Parvorder Muscicapida – previously known as the superfamily Muscicapoidea[32]
  • Parvorder Passerida – previously known as the superfamily Passeroidea[32]

Phylogeny

Relationships between living Passeriformes families based on the phylogenetic analysis of Oliveros et al (2019).[29] Some terminals have been renamed to reflect families recognised by the IOC but not in that study.[1] The IOC families Alcippeidae and Teretistridae were not sampled in this study.

PasseriformesAcanthisitti

Acanthisittidae (New Zealand wrens)

      Tyranni Eurylaimides    

Philepittidae (asites)

   

Eurylaimidae (eurylaimid broadbills)

       

Calyptomenidae (African and green broadbills)

     

Sapayoidae (sapayoa)

   

Pittidae (pittas)

        Tyrannides Furnariida    

Melanopareiidae (crescent chests)

     

Conopophagidae (gnateaters)

   

Thamnophilidae (antbirds)

         

Grallariidae (antpittas)

     

Rhinocryptidae (tapaculos)

     

Formicariidae (antthrushes)

     

Scleruridae (leaftossers)

     

Dendrocolaptidae (woodcreepers)

   

Furnariidae (ovenbirds)

              Tyrannida

Pipridae (manakins)

     

Cotingidae (cotingas)

     

Tityridae (tityras, becards)

       

Oxyruncidae (sharpbill)

   

Onychorhynchidae (royal flycatchers & allies)

       

Pipritidae (piprites)

       

Platyrinchidae (spadebills)

     

Tachurididae (many-coloured rush tyrant)

     

Rhynchocyclidae (mionectine flycatchers)

   

Tyrannidae (tyrant flycatchers)

                        PasseriMenurida

Menuridae (lyrebirds)

   

Atrichornithidae (scrubbirds)

          Climacterida

Climacteridae (Australian treecreepers)

   

Ptilonorhynchidae (bowerbirds)

          Meliphagida

Maluridae (Australasian wrens)

     

Dasyornithidae (bristlebirds)

       

Pardalotidae (gerygones & allies)

   

Acanthizidae (Australasian warblers)

     

Meliphagidae (honeyeaters)

              Orthonychida

Pomatostomidae (Australasian babblers)

   

Orthonychidae (logrunners)

        Corvides

Cinclosomatidae (quail-thrushes, jewel-babblers)

     

Campephagidae (cuckooshrikes)

       

Mohouidae (whitehead & allies)

   

Neosittidae (sittellas)

        Orioloidea

Psophodidae (whipbirds & allies)

     

Eulacestomidae (ploughbill)

       

Falcunculidae (shriketits)

   

Oreoicidae (Australo-Papuan bellbirds)

         

Paramythiidae (painted berrypickers)

   

Vireonidae (vireos)

       

Pachycephalidae (whistlers)

   

Oriolidae (orioles, figbirds)

                Malaconotoidea

Machaerirhynchidae (boatbills)

     

Artamidae (woodswallows, butcherbirds)

     

Rhagologidae (mottled berryhunter)

       

Malaconotidae (bush-shrikes, puffbacks)

     

Pityriaseidae (bristlehead)

   

Aegithinidae (loras)

         

Platysteiridae (wattle-eyes, batsies)

   

Vangidae (vangas)

            Corvoidea

Rhipiduridae (fantails)

     

Dicruridae (drongos)

       

Monarchidae (monarchs)

     

Ifritidae (ifrit)

   

Paradisaeidae (birds-of-paradise)

         

Corcoracidae (Australian mudnesters)

     

Melampittidae (melampittas)

       

Laniidae (shrikes)

   

Platylophidae (crested jay)

     

Corvidae (crows, jays)

                        Passerides    

Cnemophilidae (satinbirds)

         

Melanocharitidae (berrypeckers)

           

Callaeidae (New Zealand warblers)

   

Notiomystidae (snitchbird)

           

Petroicidae (Australian robins)

     

Eupetidae (rail-babbler)

     

Chaetopidae (rock-jumpers)

   

Picathartidae (rockfowl)

            Sylviida    

Hyliotidae (hyliotas)

   

Stenostiridae (crested flycatchers)

         

Paridae (tits, chickadees)

   

Remizidae (penduline tits)

         

Panuridae (bearded reedling)

   

Alaudidae (larks)

       

Nicatoridae (nicators)

     

Macrosphenidae (crombecs, African warblers)

     

Cisticolidae (cisticolas)

      Locustelloidea    

"Graueriidae"

   

Acrocephalidae (acrocephalid warblers)

       

Locustellidae (grassbirds)

     

Donacobiidae (donacobius)

   

Bernieridae (Malagasy warblers)

               

Pnoepygidae (wren warblers)

   

Hirundinidae (martins, swallows)

      Sylvioidea

Pycnonotidae (bulbuls)

       

Paradoxornithidae (parrotbills, fulvettas)

   

Sylviidae (typical warblers, sylviid babblers)

       

Zosteropidae (white-eyes)

     

Timaliidae (babblers, tit-babblers, scimitar bablers)

     

Leiothrichidae (laughingthrushes)

   

Pellorneidae (fulvettas, ground babblers)

            Aegithaloidea

Phylloscopidae (leaf-warblers and allies)

     

Hyliidae (hylias)

     

Aegithalidae (long-tailed tits or bushtits)

     

Erythrocercidae (yellow flycatchers)

     

Scotocercidae (streaked scrub warbler)

   

Cettiidae (Cettia bush warblers and allies)

                                  MuscicapidaBombycilloidea

Dulidae (palmchat)

     

Bombycillidae (waxwings)

     

Ptiliogonatidae (silky-flycatchers)

     

Hylocitreidae (hylocitrea)

     

Hypocoliidae (hypocolius)

   

Mohoidae (Hawaiian honeyeaters)

                  Muscicapoidea

Elachuridae (elechura)

     

Cinclidae (dippers)

       

Muscicapidae (Old World flycatchers, chats)

   

Turdidae (thrushes)

       

Buphagidae (oxpeckers)

     

Sturnidae (starlings, mynas)

   

Mimidae (mockingbirds, thrashers)

                Reguloidea

Regulidae (kinglets)

Certhioidea

Tichodromidae (wallcreeper)

     

Sittidae (nuthatches)

       

Certhiidae (treecreepers)

   

Salpornithidae

       

Polioptilidae (gnatcatchers, gnatwrens)

   

Troglodytidae (wrens)

                Passerida    

Promeropidae (sugarbirds)

   

Modulatricidae (dapple-throat & allies)

         

Nectariniidae (sunbirds)

   

Dicaeidae (flowerpeckers)

         

Chloropseidae (leafbirds)

   

Irenidae (fairy bluebirds)

       

Peucedramidae (olive warbler)

     

Urocynchramidae (Przevalski's finch)

        Phoceoidea  

Ploceidae (weavers)

     

Viduidae (indigobirds, whydals)

   

Estrildidae (estrildid finches)

           

Prunellidae (accentors)

     

Passeridae (sparrows)

     

Motacillidae (wagtails, pipits)

     

Fringillidae (finches, euphonias)

Emberizoidea

Rhodinocichlidae (thrush-tanager)

       

Calcariidae (longspurs, snow buntings)

   

Emberizidae (buntings)

         

Cardinalidae (cardinal grosbeaks)

     

Mitrospingidae (mitrosingus & allies)

   

Thraupidae (tanagers)

         

Passerellidae (American sparrows)

       

Parulidae (wood warblers)

     

Icteriidae (yellow-breasted chat)

   

Icteridae (New World blackbirds)

         

Calyptophilidae (chat-tanagers)

     

Zeledoniidae (wrenthrush)

       

Nesospingidae (Puerto Rican tanager)

   

Spindalidae (spindalises)

     

Phaenicophilidae (Hispaniolan tanagers and allies)

                                                                 

Notes

  1. ^ The name wren has been applied to other, unrelated birds in Australia and New Zealand. The 27 Australasian "wren" species in the family Maluridae are unrelated, as are the New Zealand wrens in the family Acanthisittidae; the antwrens in the family Thamnophilidae; and the wren-babblers of the families Timaliidae, Pellorneidae, and Pnoepygidae. For the monophyly of the "true wrens", Troglodytidae, see Barker 2004[13]
  2. ^ Specimen SMF Av 504. A flattened right hand of a passerine perhaps 10 cm long overall. If suboscine, perhaps closer to Cotingidae than to Eurylaimides.[22][18]
  3. ^ Specimens SMF Av 487–496; SMNS 86822, 86825-86826; MNHN SA 1259–1263: tibiotarsus remains of small, possibly basal Passeriformes.[17]
  4. ^ A partial coracoid of a probable Muscicapoidea, possibly Turdidae; distal tibiotarsus and tarsometatarsus of a smallish to mid-sized passerine that may be the same as the preceding; proximal ulna and tarsometatarsus of a Paridae-sized passerine.[24][25]
  5. ^ A humerus diaphysis piece of a swallow-sized passerine.[26]
  6. ^ Distal right humerus, possibly suboscine.[27][28]
  7. ^ Oliveros et al (2019) use the list of families published by Dickinson and Christidis in 2014.[29][31] Oliveros et al include 8 families that are not included on the IOC list. These are not shown here. By contrast, the IOC list includes 15 families that are not present in Dickinson and Christidis. In 13 of these cases, the position of the additional family in the taxonomic order can be determined from the species included by Oliveros and colleagues in their analysis. No species in the families Alcippeidae and Teretistridae were sampled by Oliveros et al so their position is uncertain.[1][29]
  8. ^ The order of the families within the superfamily Orioloidea is uncertain.[29]
  9. ^ The order of the families within the superfamily Malaconotoidea is uncertain.[29]
  10. ^ The order of the families within the superfamily Corvoidea is uncertain.[29]
  11. ^ The taxonomic sequence of the superfamilies Locustelloidea, Sylvioidea and Aegithaloidea is uncertain, although the order of the families within each of the superfamilies is well determined.[29]
  12. ^ The order of some of the families within the superfamily Emberizoidea is uncertain.[29]
  13. ^ The family Teretistridae (Cuban warblers) is tentatively placed here. The family was not included in the analysis published by Oliveros et al (2019).[29] Dickinson and Christidis (2014) considered the genus Teretistris Incertae sedis.[34] Barker et al (2013) found that Teretistridae is closely related to Zeledoniidae.[33]

References

  1. ^ a b c d e Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2020). "Family Index". IOC World Bird List Version 10.1. International Ornithologists' Union. Retrieved 26 April 2020.
  2. ^ a b Ericson, P.G.P. et al. (2003) Evolution, biogeography, and patterns of diversification in passerine birds. J. Avian Biol, 34:3–15.
  3. ^ a b c Selvatti, A.P. et al. (2015) "A Paleogene origin for crown passerines and the diversification of the Oscines in the New World". Molecular Phylogenetics and Evolution, 88:1–15.
  4. ^ Barker, F. Keith; Barrowclough, George F.; Groth, Jeff G. (2002). "A phylogenetic hypothesis for passerine birds: Taxonomic and biogeographic implications of an analysis of nuclear DNA sequence data". Proceedings of the Royal Society B: Biological Sciences. 269 (1488): 295–308. doi:10.1098/rspb.2001.1883. PMC 1690884. PMID 11839199.
  5. ^ a b Ericson, P.G.; Christidis, L.; Cooper, A.; Irestedt, M.; Jackson, J.; Johansson, U.S.; Norman, J.A. (7 February 2002). "A Gondwanan origin of passerine birds supported by DNA sequences of the endemic New Zealand wrens". Proceedings of the Royal Society B. 269 (1488): 235–241. doi:10.1098/rspb.2001.1877. PMC 1690883. PMID 11839192.
  6. ^ Chatterjee, Sankar (2015). The Rise of Birds: 225 Million Years of Evolution. JHU Press. pp. 206–207. ISBN 9781421415901.
  7. ^ Winkler, D. W., S. M. Billerman, and I.J. Lovette (2020). Lyrebirds (Menuridae), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.menuri1.01
  8. ^ Madge, S. (2020). Thick-billed Raven (Corvus crassirostris), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.thbrav1.01
  9. ^ Clock, B. (2020). Short-tailed Pygmy-Tyrant (Myiornis ecaudatus), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.stptyr1.01
  10. ^ Stefoff, Rebecca (2008), The Bird Class, Marshall Cavendish Benchmark
  11. ^ Brooke, Michael and Birkhead, Tim (1991) The Cambridge Encyclopedia of Ornithology, Cambridge University Press ISBN 0521362059.
  12. ^ Jones, D. (2008) "Flight of fancy". Australian Geographic, (89), 18–19.
  13. ^ Barker, F.K. (2004). "Monophyly and relationships of wrens (Aves: Troglodytidae): a congruence analysis of heterogeneous mitochondrial and nuclear DNA sequence data". Molecular Phylogenetics and Evolution. 31 (2): 486–504. doi:10.1016/j.ympev.2003.08.005. PMID 15062790.
  14. ^ Dyke, Gareth J.; Van Tuinen, Marcel (June 2004). "The evolutionary radiation of modern birds (Neornithes): Reconciling molecules, morphology and the fossil record". Zoological Journal of the Linnean Society. 141 (2): 153–177. doi:10.1111/j.1096-3642.2004.00118.x.
  15. ^ a b c d Claramunt, S.; Cracraft, J. (2015). "A new time tree reveals Earth history's imprint on the evolution of modern birds". Science Advances. 1 (11): e1501005. Bibcode:2015SciA....1E1005C. doi:10.1126/sciadv.1501005. PMC 4730849. PMID 26824065.
  16. ^ a b Boles, Walter E. (1997). "Fossil songbirds (Passeriformes) from the Early Eocene of Australia". Emu. 97 (1): 43–50. doi:10.1071/MU97004.
  17. ^ a b c Manegold, Albrecht; Mayr, Gerald & Mourer-Chauviré, Cécile (2004). "Miocene Songbirds and the Composition of the European Passeriform Avifauna". Auk. 121 (4): 1155–1160. doi:10.1642/0004-8038(2004)121[1155:MSATCO]2.0.CO;2.
  18. ^ a b c Mayr, Gerald & Manegold, Albrecht (2006). "A Small Suboscine-like Passeriform Bird from the Early Oligocene of France". Condor. 108 (3): 717–720. doi:10.1650/0010-5422(2006)108[717:ASSPBF]2.0.CO;2.
  19. ^ Mayr, G (2013). "The age of the crown group of passerine birds and its evolutionary significance–molecular calibrations versus the fossil record". Systematics and Biodiversity. 11 (1): 7–13. doi:10.1080/14772000.2013.765521. S2CID 85167051.
  20. ^ Bochenski, Z.M., Tomek, T., Bujoczek, M. (2021) A new passeriform (Aves: Passeriformes) from the early Oligocene of Poland sheds light on the beginnings of Suboscines. J Ornithol. https://doi.org/10.1007/s10336-021-01858-0
  21. ^ Worthy, Trevor H.; Tennyson, A.J.D.; Jones, C.; McNamara, J.A.; Douglas, B.J. (2007). "Miocene waterfowl and other birds from central Otago, New Zealand". Journal of Systematic Palaeontology. 5 (1): 1–39. doi:10.1017/S1477201906001957. hdl:2440/43360. S2CID 85230857.
  22. ^ Roux, T. (2002). "Deux fossiles d'oiseaux de l'Oligocène inférieur du Luberon" [Two bird fossils from the Lower Oligocene of Luberon] (PDF). Courrier Scientifique du Parc Naturel Régional du Luberon. 6: 38–57.
  23. ^ Hugueney, Marguerite; Berthet, Didier; Bodergat, Anne-Marie; Escuillié, François; Mourer-Chauviré, Cécile & Wattinne, Aurélia (2003). "La limite Oligocène-Miocène en Limagne: changements fauniques chez les mammifères, oiseaux et ostracodes des différents niveaux de Billy-Créchy (Allier, France)" [The Oligocene-Miocene boundary in Limagne: faunal changes in the mammals, birds and ostracods from the different levels of Billy-Créchy (Allier, France)]. Geobios. 36 (6): 719–731. doi:10.1016/j.geobios.2003.01.002.
  24. ^ Gál, Erika; Hír, János; Kessler, Eugén & Kókay, József (1998–99). "Középsõ-miocén õsmaradványok, a Mátraszõlõs, Rákóczi-kápolna alatti útbevágásból. I. A Mátraszõlõs 1. lelõhely" [Middle Miocene fossils from the sections at the Rákóczi chapel at Mátraszőlős. Locality Mátraszõlõs I.]. Folia Historico Naturalia Musei Matraensis. 23: 33–78.
  25. ^ Gál, Erika; Hír, János; Kessler, Eugén; Kókay, József & Márton, Venczel (2000). "Középsõ-miocén õsmaradványok a Mátraszõlõs, Rákóczi-kápolna alatti útbevágásból II. A Mátraszõlõs 2. lelõhely" [Middle Miocene fossils from the section of the road at the Rákóczi Chapel, Mátraszõlõs. II. Locality Mátraszõlõs 2]. Folia Historico Naturalia Musei Matraensis. 24: 39–75.
  26. ^ a b Hír, János; Kókay, József; Venczel, Márton; Gál, Erika & GKessler, Eugén (2001). "Elõzetes beszámoló a felsõtárkányi "Güdör-kert" n. õslénytani lelõhelykomplex újravizsgálatáról" [A preliminary report on the revised investigation of the paleontological locality-complex "Güdör-kert" at Felsõtárkány, Northern Hungary] (PDF). Folia Historico Naturalia Musei Matraensis. 25: 41–64.
  27. ^ Noriega, Jorge I. & Chiappe, Luis M. (1991). "El más antiguo Passeriformes de America del Sur. Presentation at VIII Journadas Argentinas de Paleontologia de Vertebrados" [The most ancient passerine from South America]. Ameghiniana. 28 (3–4): 410.
  28. ^ Noriega, Jorge I. & Chiappe, Luis M. (1993). "An Early Miocene Passeriform from Argentina" (PDF). Auk. 110 (4): 936–938. doi:10.2307/4088653. JSTOR 4088653.
  29. ^ a b c d e f g h i j k l Oliveros, C.H.; et al. (2019). "Earth history and the passerine superradiation". Proceedings of the National Academy of Sciences of the United States of America. 116 (16): 7916–7925. doi:10.1073/pnas.1813206116. PMC 6475423. PMID 30936315.
  30. ^ The former does not even have recognized subspecies, while the latter is one of the most singular birds alive today. Good photos of a bearded reedling are for example here Archived 16 October 2007 at the Wayback Machine and here Archived 31 July 2008 at the Wayback Machine.
  31. ^ Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World. Vol. 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. ISBN 978-0-9568611-2-2.
  32. ^ a b c d e Cracraft, Joel (2014). "Avian higher-level relationships and classification: passeriformes". In Dickinson, E.C.; Christidis, L. (eds.). The Howard & Moore Complete Checklist of the Birds of the World. Vol. 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. pp. xvii-xxxiii [xxiv]. ISBN 978-0-9568611-2-2.
  33. ^ a b Barker, F.K.; Burns, K.J.; Klicka, J.; Lanyon, S.M.; Lovette, I.J. (2013). "Going to extremes: contrasting rates of diversification in a recent radiation of New World passerine birds". Systematic Biology. 62 (2): 298–320. doi:10.1093/sysbio/sys094. PMID 23229025.
  34. ^ Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World. Vol. 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. p. 358. ISBN 978-0-9568611-2-2.
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Passerine: Brief Summary

provided by wikipedia EN

A passerine (/ˈpæsəraɪn/) is any bird of the order Passeriformes (/ˈpæsərɪfɔːrmiːz/; from Latin passer 'sparrow' and formis '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds or songbirds, passerines are distinguished from other orders of birds by the arrangement of their toes (three pointing forward and one back), which facilitates perching.

With more than 140 families and some 6,500 identified species, Passeriformes is the largest order of birds and among the most diverse orders of terrestrial vertebrates, representing 60% of birds. Passerines are divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni (suboscines), and Passeri (oscines). The passerines contain several groups of brood parasites such as the viduas, cuckoo-finches, and the cowbirds. Most passerines are omnivorous, while the shrikes are carnivorous.

The terms "passerine" and "Passeriformes" are derived from the scientific name of the house sparrow, Passer domesticus, and ultimately from the Latin term passer, which refers to sparrows and similar small birds.

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