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Lifespan, longevity, and ageing

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Maximum longevity: 17.6 years (captivity) Observations: Males are only fully mature at 8-9 years. Potential longevity has been estimated at 22 years (Ronald Nowak 1999), which is possible. Record longevity in captivity, however, is 17.6 years (Richard Weigl 2005).
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Life Cycle

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See Reproduction.

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Associations

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Eurasian lynx and wolves are the main predator of the chamois. (Gortazar, 2000). They are also hunted by humans. When alarmed, these animals flee to inaccesible locations. They can travel at speeds of up to 50 km per hour. They can jump 2 meters into the air, and a distance of up to 6 meters. (Nowak, 1999)

Known Predators:

  • Eurasian lynx (Lynx lynx)
  • gray wolves (Canis lupus)
  • humans (Homo sapiens)
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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Morphology

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Weight: 25 to 50 kg; Shoulder Height: 70 to 80 cm; Length 110 to135 cm. Chamois are a chestnut color but are lighter in the spring and summer. In the winter these animals grow long guard hairs over their dark brown under fur. Under parts are pale and the rump is white at the tail. A dark brown band runs from each side of the muzzle to the ears and eyes, and the rest of the head and throat is white. The horns of the male rise directly above the head then hook sharply back at the tips. The female also has horns, which although slimmer than the male's, can be longer. The female is smaller than the male. The hooves of the chamois are excellent for gripping slippery rock. (Nowak, 1999)

Range mass: 30 to 50 kg.

Range length: 110 to 135 cm.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: male larger; sexes shaped differently; ornamentation

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Life Expectancy

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The lifespan of the chamois ranges from 14-22 years. (Huffman, 1999)

Range lifespan
Status: wild:
14 to 22 years.

Average lifespan
Status: wild:
20.0 years.

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Habitat

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R. rupricapra lives in alpine and sub alpine meadows above the timberline. It winters in forested areas and steep slopes where snow does not accumulate. It is found in both relatively steep and flatter terrain. (Nowak, 1999)

Habitat Regions: temperate

Terrestrial Biomes: tundra ; mountains

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Distribution

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The range of Rupricapra rupicapra, or chamois, includes the Pyrenees, the mountains of south and central Europe, Turkey, and the Caucasus in Asia. It has been introduced on the South Island of New Zealand. (Huffman, 1999; Nowak, 1999)

Biogeographic Regions: palearctic (Native ); australian (Introduced )

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Trophic Strategy

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During the summer months the diet consists chiefly of herbs and flowers, but in winter the chamois eats lichens, mosses, and young pine shoots. It has been known to fast for two weeks and survive when the snow is so deep that food can not be found. (Nowak, 1999)

Plant Foods: leaves; flowers; bryophytes; lichens

Primary Diet: herbivore (Folivore )

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Dan Gunderson, University of Wisconsin-Stevens Point
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Associations

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This species provides food for two interesting predators, the eurasian lynx and the wolf. As a grazer, it also affects the plant community within its habitat.

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Benefits

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The meat of chamois is considered a prized food by some people. The skin is made into chamois (pronounces "shammy") leather for cleaning glass and polishing automobiles. The winter hair from the back is used to make the "gamsbart," the brush of Tyrelean hats. Chamois also bring increased tourism through hunting. (Nowak, 1999, Gortazar 2000)

Positive Impacts: food ; ecotourism

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Benefits

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Chamois compete with domestic sheep for grazing. (Gortazar, 2000)

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Conservation Status

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In the Caucasus Mountains, Tatra Mountains that run along the border of Poland and Slovakia, and in Massif de la Chartreuse in South Eastern France, excessive hunting, loss of habitat, competition with livestock, and harassment by people and dogs have greatly reduced the number of chamois. Otherwise, chamois now are generally increasing in number and have been introduced and reintroduced in various parts of Europe. Population in Europe is about 400,000. (Nowak 1999)

US Federal List: endangered

CITES: no special status

IUCN Red List of Threatened Species: least concern

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Dan Gunderson, University of Wisconsin-Stevens Point
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Behavior

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Perception Channels: tactile ; chemical

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Reproduction

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Males are generally solitary except during the breeding season. They join herds during the late summer. Older males are known to force younger males from the herd, and sometimes have killed them (Nowak, 1999). It is likely that breeding is polygynous.

Mating System: polygynous

After a gestation period of 170 days, kids are born in May and June in a shelter of grass and lichens. There is usually only one kid born to a female, but twins and triplets sometimes occur. Young weigh 2 o 3kg each and are weaned after 2 to 3 months. The precocial kids are able to follow their mother almost immediately after they are born and they rapidly improve their leaping ability within the first few days of life. If a mother is killed, other chamois take care of the young. Young males stay with the mother's group until they are 2 to 3 years old and then live nomadically until they are fully mature at 8 to 9 years, when they become attached to a definite area. Sexual maturity is reached at the age of 2.5 years in females and 3.5 to 4 years in males. (Nowak, 1999; Huffman, 1999)

Breeding season: Breeding occurs from October through December, with young born in May and June.

Range number of offspring: 1 to 2.

Range gestation period: 5.33 to 6.17 months.

Range weaning age: 2 to 3 months.

Range age at sexual or reproductive maturity (female): 2.5 to 4 years.

Range age at sexual or reproductive maturity (male): 2.5 to 4 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; fertilization (Internal ); viviparous

Average birth mass: 2400 g.

Average number of offspring: 1.

Young are precocial and able to follow their mother shortly after birth. The mother produces milk for the young, and nurses them for 2-3 months. Should the mother be killed, other chamois will care for the young. (Nowak, 1999)

Parental Investment: altricial ; female parental care ; post-independence association with parents

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Gunderson, D. 2003. "Rupicapra rupicapra" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rupicapra_rupicapra.html
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Biology

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The nimble chamois are well suited to the harsh terrain of the mountain ranges they inhabit. They are prodigious runners, capable of reaching speeds of 50 kilometres per hour on uneven ground, and daring leapers that can jump almost two metres in height and at least six metres in length (2). While in the summer months, chamois can feed on a relatively plentiful diet of herbs and flowers, during winter they turn to lichens, mosses and young pine shoots, and are able to survive for up to two weeks without food when snow blankets any sources of sustenance (2). Female chamois with their young generally live in herds of 15 to 30 individuals. Herds are believed to post 'sentinels' that warn the other members of the herd of any danger by stomping their feet and calling with a high-pitched whistle. Adult males live on their own for most of the year, but join the herds in late summer, in time for the autumn rut. During the rut, often in November, old, strong males drive the younger males away from the herd, and occasionally kill them (2). Following mating, and after a 170 day gestation period, females separate themselves from the herd to give birth in May and June. Females usually bear a single kid, giving birth in a shelter of grass and lichens. The kids are able to follow their mother immediately after they are born and are weaned after just two to three months. While female kids may remain with their mother's herds, young males leave the herd at the age of two or three. They then have a nomadic existence until they reach full maturity, at the age of eight or nine, when they become attached to a definite area (2).
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Conservation

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While the chamois is, as a species, still common, a number of the subspecies are threatened and require conservation action. The Chartreuse chamois has, since 1972, been the focus of conservation measures, beginning with efforts to educate local hunters. The hunters formed a Chamois Management Unit and together implemented measures including a shooting moratorium, lasting several years, and limiting livestock grazing on upland pastures. The efforts were a success, with the population multiplying by five between 1985 and 1997, but the population still faces threats, particularly hybridisation with the Alpine chamois (5). The Critically Endangered Tatra chamois is protected by law in Poland and Slovakia, and occurs solely within two national parks, each of which includes an area in which public access is strictly controlled to eliminate any disturbance during winter and during the birth season. Control of sheep grazing in Tatra National Park led to an increase in population numbers, but more recently numbers have again declined, possibly due to poor weather conditions and poaching (4). Although not so greatly threatened, the other five subspecies are subject to varying hunting laws and legal protection, occur in numerous protected areas, and some, such as the Balkan chamois and Carpathian chamois, have been successfully introduced to further areas (4).
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Description

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The chamois is an agile and graceful animal (2), adapted to cold, highland terrain (3). Its winter coat consists of thick, woolly underfur and long guard hairs, suited to conserving heat in its chilly alpine habitat, and the somewhat elastic pads of the hooves provide reassuring grip on uneven and slippery ground (2). The winter coat is blackish-brown, with a white patch on the rump and white to yellow facial stripes (3), while in spring and summer, this is shed for a lighter tawny-brown coat of short, stiff, coarse hairs (2). Both male and female chamois bear slender, black horns. Measuring up to 203 millimetres, the closely set horns rise almost vertically from the forehead and then bend abruptly backwards to form hooks (2).
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Habitat

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During the spring and summer months, chamois inhabit alpine meadows, generally above altitudes of 1,800 metres and never more than a few hundred metres from the safe refuge of cliffs. In autumn and winter, chamois can be found at lower altitudes, often below 1,100 metres, where they stay on steep slopes where snow does not accumulate, and sometimes enter forests (2).
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Range

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Occurs in mountain ranges of south-central Europe and Asia Minor, including the Alps, Carpathians, Balkans, and Caucasus, where it occurs as seven subspecies. The chamois has also been introduced to New Zealand (2) (3).
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Status

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Classified as Lower Risk / Least Concern (LR/lc) on the IUCN Red List 2007 (1). The Tatra chamois (R. r. tatrica) and Chartreuse chamois (R. r. cartusiana) are classified as Critically Endangered (CR), the Caucasian chamois (R. r. caucasica) is classified as Vulnerable (VU), the Balkan chamois (R. r. balcanica), the Carpathian chamois (R. r. carpatica) and the Alpine chamois (R. r. rupicapra) are classified as Lower Risk / Least Concern (LR/lc), and the Turkish chamois (R. r. asiatica) is classified as Data Deficient (DD) on the IUCN Red List 2007 (1).
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Threats

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With the exception of the Alpine chamois (Rupicapra rupicapra rupicapra), many other subspecies of the chamois are rare and many populations are declining (4). Such declines are due to a combination of factors. The flesh of chamois is prized by some people, the skin is made into shammy leather for cleaning glass and polishing cars, and winter hair from the back is used to make 'gamsbart', the brush of Tyrolean hats (2). This has led, in some areas, to excessive hunting (2), and poaching remains a threat to many populations, particularly outside of protected areas (4). Habitat loss is impacting some subspecies, such as in Albania where suitable habitat is being lost to expanding human populations, as is competition with domestic livestock. Hybridisation with the Alpine chamois poses a threat to a number of the subspecies and sarcoptic mange (a skin disease) is also a problem in some regions (4). The two Critically Endangered subspecies, the Chartreuse chamois (R. r. cartusiana) and the Tatra chamois (R. r. tatrica) face many of the threats mentioned above, which are compounded by their small populations and limited distribution. The Chartreuse chamois is restricted to a 350 square kilometre area of the Chartreuse massif, at the western edge of French Alps, where an official census in 1986-87 estimated the population to consist of a minimum of 150 individuals. Only two populations of the Tatra chamois remain; one in the Tatra National Park, Poland and Slovakia, and another introduced population occurs in the Low Tatra National Park, Slovakia. Such small populations are vulnerable to natural disasters, disease, and a loss of genetic diversity (4).
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One Species at a Time Podcast

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Growing up in a village in the foothills of the French Alps, Francis Roucher used to hunt the chamois, a cross between a goat and an antelope. But on the day one of his shots went astray, Roucher was transformed from hunter to game manager, working to reverse the chamois’ decline.

Listen to the podcast on the EOL Learning + Education website where you can also meet the featured scientist and find intriguing extras.

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Chamois

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The chamois (Rupicapra rupicapra) or Alpine chamois is a species of goat-antelope native to mountains in Europe, from west to east, including the Alps, the Dinarides, the Tatra and the Carpathian Mountains, the Balkan Mountains, the RilaRhodope massif, Pindus, the northeastern mountains of Turkey, and the Caucasus.[2] The chamois has also been introduced to the South Island of New Zealand. Some subspecies of chamois are strictly protected in the EU under the European Habitats Directive.[3]

Names

Chamois herd engraved on reindeer antler from Gourdan grotto, Haute Garonne.

The English name comes from the French chamois. The latter is derived from Gaulish camox (attested in Latin, 5th century), itself perhaps borrowing from some Alpine language (Raetic, Ligurian). The Gaulish form also underlies German Gemse, Gams, Gämse, Italian camoscio, Ladin ciamorz.

The usual pronunciation for the animal is UK: /ˈʃæmwɑː/ or US: /ʃæmˈwɑː/, approximating the French pronunciation [ʃamwa]. However, when referring to chamois leather, and in New Zealand often for the animal itself, it is /ˈʃæmi/, and sometimes spelt shammy or chamy. The plural of chamois is spelled the same as the singular, and it may be pronounced with the final "s" sounded. However, as with many other quarry species, the plural for the animal is often pronounced the same as the singular.

The Dutch name for the chamois is gems, and the male is called a gemsbok. In Afrikaans, the name gemsbok came to refer to a species of Subsaharan antelope of the genus Oryx, and this meaning of gemsbok has been adopted into English.

Taxonomy

The species R. rupicapra is categorized into seven subspecies:

Description

Rupicapra rupicapra tatrica in the Tatra Mountains

The chamois is a very small bovid. A fully grown chamois reaches a height of 70–80 cm (28–31 in) and measures 107–137 cm (42–54 in) (the tail is not generally visible except when mating).[6] Males, which weigh 30–60 kg (66–132 lb), are slightly larger than females, which weigh 25–45 kg (55–99 lb).[6] Both males and females have short, straightish horns which are hooked backwards near the tip, the horn of the male being thicker. In summer, the fur has a rich brown colour which turns to a light grey in winter. Distinct characteristics are white contrasting marks on the sides of the head with pronounced black stripes below the eyes, a white rump and a black stripe along the back.

Biology and behaviour

Footprint at Rila National Park, Bulgaria, 2014
Chamois in the Gran Paradiso National Park in Italy

Female chamois and their young live in herds of up to 15 to 30 individuals; adult males tend to live solitarily for most of the year. During the rut (late November/early December in Europe, May in New Zealand), males engage in fierce battles for the attention of unmated females. An impregnated female undergoes a gestation period of 170 days, after which a single kid is usually born in May or early June. On rare occasions, twins may be born.[6] If a mother is killed, other females in the herd may try to raise the young.[7] The kid is weaned at six months of age and is fully grown by one year of age. However, the kids do not reach sexual maturity until they are three to four years old, although some females may mate at as early two years old.[6] At sexual maturity, young males are forced out of their mother's herds by dominant males (who sometimes kill them), and then wander somewhat nomadically until they can establish themselves as mature breeding specimens at eight to nine years of age.[7]

Chamois eat various types of vegetation, including highland grasses and herbs during the summer and conifers, barks and needles from trees in winter. Primarily diurnal in activity, they often rest around mid-day and may actively forage during moonlit nights.[6]

Chamois can reach an age of 22 years in captivity, although the average recorded age in the wild ranges from 15 to 17 years. Common causes of mortality can include avalanches, epidemics and predation. At present, humans are the main predator of chamois. In the past, the principal predators were Eurasian lynxes, Persian leopards and gray wolves; with some predation possibly by brown bears and golden eagles.[6] Chamois usually use speed and stealthy evasion to escape predators and can run at 50 kilometers per hour (31 mph) and can jump 2 m (6.6 ft) vertically into the air or over a distance of 6 m (20 ft).[7]

Distribution and habitat

Rupicapra rupicapra carpatica in the Retezat Mountains

The chamois is native to the Pyrenees, the mountains of south and central Europe, Turkey, and the Caucasus. It lives in precipitous, rugged, rocky terrain at moderately high elevations of up to at least 3,600 m (11,800 ft). In Europe, the chamois spends the summer months in alpine meadows above the tree line, but moves to elevations of around 800 m (2,600 ft) to spend the winter in pine-dominated forests.

In New Zealand

Alpine chamois arrived in New Zealand in 1907 as a gift from the Austrian Emperor, Franz Joseph I in exchange for specimens of living ferns, rare birds and lizards. Mr Albert E.L. Bertling, formerly head keeper of the Zoological Society's Gardens, Regents Park, London, accepted an invitation from the New Zealand Government to deliver a consignment of chamois (two bucks and six does) to the colony. They arrived in Wellington, New Zealand, on 23 January 1907, on board the SS Turakina. From Wellington the chamois were transhipped to the Manaroa and conveyed to Lyttelton, then by rail to Fairlie in South Canterbury and a four-day horse trek to Mount Cook. The first surviving releases were made in the Aoraki / Mount Cook region and these animals gradually spread over much of the South Island.[8][9]

In New Zealand, chamois hunting is unrestricted and even encouraged by the Department of Conservation to limit the animal's impact on New Zealand's native alpine flora.[9][10]

New Zealand chamois tend to weigh about 20% less than European individuals of the same age, suggesting that food supplies may be limited.[11]

Chamois on the Piz Beverin mountain, Switzerland

Hunting and wildlife management

As their meat is considered tasty,[12] chamois are popular game animals. Chamois have two traits that are exploited by hunters: the first is that they are most active in the morning and evening when they feed; the second is that they tend to look for danger originating from below, which means that a hunter stalking chamois from above is less likely to be observed and more likely to be successful.[13]

The tuft of hair from the back of the neck, the gamsbart (chamois "beard"), is traditionally worn as a decoration on hats throughout the alpine countries.

Chamois leather

Chamois leather

Chamois leather, traditionally made from the hide of the chamois, is very smooth and absorbent and is favoured in cleaning, buffing, and polishing because it produces no scratching. Modern chamois leather may be made from chamois hides, but hides of deer or domestic goats or sheep are commonly used.

Chamois fabric

A fabric known as chamois is made variously from cotton flannel, PVA, Viscose, and other materials with similar qualities. It is napped to produce a plush surface similar to moleskin or chamois leather.

See also

References

  1. ^ Anderwald, P.; Ambarli, H.; Avramov, S.; Ciach, M.; Corlatti, L.; Farkas, A.; Papaioannou, H.; Peters, W.; Sarasa, M.; Šprem, N.; Weinberg, P. & Willisch, C. (2021) [amended version of 2020 assessment]. "Rupicapra rupicapra". IUCN Red List of Threatened Species. 2021: e.T39255A195863093. doi:10.2305/IUCN.UK.2021-1.RLTS.T39255A195863093.en. Retrieved 17 February 2022.
  2. ^ "Il Camoscio d'Abruzzo". www.camosciodabruzzo.it.
  3. ^ "EUR-Lex - 31992L0043 - EN". eur-lex.europa.eu.
  4. ^ Current status of the Balkan chamois (Rupicapra rupicapra balcanica) in Greece: Implications for conservation at Royal Belgian Institute of Natural Sciences Archived April 10, 2008, at the Wayback Machine
  5. ^ a b "Mineral Supply and Fertility of Chamois" (PDF).
  6. ^ a b c d e f Macdonald, D.W.; Barrett, P. (1993). Mammals of Europe. New Jersey: Princeton University Press. ISBN 0-691-09160-9.
  7. ^ a b c Dan Gunderson. "ADW: Rupicapra rupicapra: INFORMATION". Animal Diversity Web.
  8. ^ "A Note on the Chamois in New Zealand at New Zealand Ecological Society" (PDF).
  9. ^ a b "Recreational hunting in Nelson/Marlborough - Chamois at the Department of Conservation" (PDF).
  10. ^ "Heritage Preservation (p. 40 and 45) at the Department of Conservation" (PDF). Archived from the original (PDF) on 26 September 2012. Retrieved 12 February 2008.
  11. ^ "Trophy Chamois Buck Hunting New Zealand Free Range Safari Park Record Horns". Archived from the original on 4 February 2017. Retrieved 1 September 2012.
  12. ^ Tschohl, Christl. "Chamois goulosh: wild on wild". Montafan.at. Montafon Tourismus GmbH. Archived from the original on 13 February 2019. Retrieved 12 February 2019.
  13. ^ "Beginners Chamois Hunting Guide".

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Chamois: Brief Summary

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The chamois (Rupicapra rupicapra) or Alpine chamois is a species of goat-antelope native to mountains in Europe, from west to east, including the Alps, the Dinarides, the Tatra and the Carpathian Mountains, the Balkan Mountains, the RilaRhodope massif, Pindus, the northeastern mountains of Turkey, and the Caucasus. The chamois has also been introduced to the South Island of New Zealand. Some subspecies of chamois are strictly protected in the EU under the European Habitats Directive.

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