Emarginula is a genus of small keyhole limpets, marine gastropod molluscs in the family Fissurellidae.[1]
Description
The oval shell is obliquely conical with a backward-pointing recurved apex. There is a deep incision in the anterior margin. A distinct anal fasciole extends upwards from this incision. The surface is latticed, sculptured with radial ribs. There is no septum inside.[2]
Radula
The radula has a dental formula of [many].1.4.1.4.1.[many],[3] with a wide, rectangular, acuspate rachidian tooth.[4] The various genera have a few dozen rows of teeth.[3]
Species
According to the World Register of Marine Species (WoRMS), species within the genus Emarginula include:[1]
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Emarginula adamsiana Sowerby in Adams & Sowerby, 1863
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Emarginula adriatica Costa O.G., 1829
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Emarginula agulhasensis Thiele, 1925
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Emarginula alba Watson, 1897
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Emarginula amyda Shikama, 1962
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Emarginula angusta McLean, 1970
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Emarginula annielangleitae Poppe & Tagaro, 2020
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Emarginula apolonia Pacaud, 2015
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Emarginula balicasagensis Poppe & Tagaro, 2020
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Emarginula bicancellata Montrouzier, 1860
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Emarginula bonfittoi Smriglio & Mariottini, 2001
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Emarginula boucheti Poppe & Tagaro, 2020
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Emarginula camilla Melvill & Standen, 1903
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Emarginula candida Adams, 1852
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Emarginula choristes Dall, 1925
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Emarginula christiaensi Piani, 1985
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Emarginula circumalbum Poppe & Tagaro, 2020
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Emarginula clypeus Adams, 1852
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Emarginula colipanoae Poppe & Tagaro, 2020
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Emarginula compta Habe, 1953
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Emarginula concinna Adams, 1852
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Emarginula connectens Thiele, 1915
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Emarginula connelli Kilburn, 1978
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Emarginula convexa Hedley, 1907
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Emarginula costulata Deshayes, 1863
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Emarginula crassa Sowerby, 1812
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Emarginula crassicostata Sowerby in Adams & Sowerby, 1863
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Emarginula curvamen Iredale, 1925
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Emarginula cuvieri Audouin, 1827 (species inquirenda)
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Emarginula decorata Deshayes, 1863
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Emarginula delonguevilleae Poppe & Tagaro, 2020
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Emarginula devota Thiele, 1915
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Emarginula dictya McLean, 1970
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Emarginula dilecta Adams, 1852
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Emarginula divae (van Aartsen & Carrozza, 1995)
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Emarginula dubia Schepman, 1908
- † Emarginula espibosensis Lozouet, 1999
- † Emarginula fenestrata Deshayes, 1861 †
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Emarginula fenestrella Deshayes, 1863
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Emarginula fissura (Linnaeus, 1758) - slit limpet
- † Emarginula fossorensis Lozouet, 1999
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Emarginula foveolata Schepman, 1908
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Emarginula foveolata fugitai Habe, 1953
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Emarginula fragilis Yokoyama, 1920
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Emarginula fulginea Adams, 1851
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Emarginula gabensis (Gabriel, 1961)
- † Emarginula galeriformis Marwick, 1928
- † Emarginula gigantea Poppe, 2008
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Emarginula gustavi Poppe & Tagaro, 2020
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Emarginula hataii Habe, 1953
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Emarginula hawaiiensis Dall, 1895
- † Emarginula haweraensis Powell, 1931
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Emarginula hosoyai Habe, 1953
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Emarginula huzardii (Payraudeau, 1826)
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Emarginula icosisculpta Simone & Cunha, 2014
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Emarginula imaizumi Dall, 1926
- † Emarginula imbricata Millet, 1865
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Emarginula imella Dall, 1926
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Emarginula intervecta Locard, 1898: synonym of Emarginula fissura (Linnaeus, 1758)
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Emarginula janae Cossignani, 2020
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Emarginula japonica Sowerby in Adams & Sowerby, 1863
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Emarginula jungcheni K.-Y. Lai, 2019
- † Emarginula kaiparica Laws, 1939
-
Emarginula kashimaensis Shikama, 1962
- † Emarginula komitica Laws, 1939
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Emarginula koon Kilburn, 1978
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Emarginula lata J.R.C. Quoy & J.P. Gaimard, 1834
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Emarginula liuzzii Poppe & Tagaro, 2020
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Emarginula longifissa Sowerby, 1866
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Emarginula lorenzoi Giusti & Micali, 2019
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Emarginula macclurgi Kilburn, 1978
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Emarginula mactanensis Poppe & Tagaro, 2020
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Emarginula maculata Adams in Adams & Sowerby, 1863
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Emarginula margarita Poppe & Tagaro, 2020
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Emarginula micans Adams, 1852
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Emarginula multistriata Jeffreys, 1882
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Emarginula natalensis Barnard, 1963
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Emarginula nigromaculata (Thiele, 1930)
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Emarginula nordica Pérez Farfante, 1947 - northern emarginula
- † Emarginula oblonga Sandberger, 1859
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Emarginula obovata Adams, 1852
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Emarginula octaviana Coen, 1939
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Emarginula oppressa Barnard, 1963
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Emarginula paivana (Crosse, 1867)
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Emarginula patula Cotton, 1930
- † Emarginula paucicostata Laws, 1936
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Emarginula paucipunctata Schepman, 1908
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Emarginula philippinensis Poppe & Tagaro, 2020
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Emarginula phrixodes Dall, 1927 - ruffled emarginula
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Emarginula pittensis Marwick, 1928
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Emarginula poppeorum Romani & Crocetta, 2017
- † Emarginula praesicula Lozouet, 1999
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Emarginula puncticulata Adams, 1852
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Emarginula punctulum Piani, 1980
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Emarginula pustula Thiele in Küster, 1913
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Emarginula quadrata Poppe & Tagaro, 2020
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Emarginula regia Habe, 1953
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Emarginula retecosa Adams, 1851
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Emarginula rosea Bell, 1824
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Emarginula scabriuscula Adams, 1852
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Emarginula sicula Gray, 1825 - dagger emarginula
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Emarginula sinica Lu, 1986
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Emarginula solidula Costa, 1829
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Emarginula souverbiana Pilsbry, 1890
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Emarginula spinosa Deshayes, 1863
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Emarginula striatula Quoy and Gaimard, 1834 - dagger emarginula [5]
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Emarginula subtilitexta Verco, 1908
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Emarginula superba Hedley, 1906
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Emarginula suspira Simone & Cunha, 2014
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Emarginula tenera Locard, 1892
- † Emarginula teneraeformis Lozouet, 1999
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Emarginula tenuicostata Sowerby, 1863
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Emarginula teramachii Habe, 1953
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Emarginula textilis Gould, 1859 in 1859-61
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Emarginula thomasi Crosse, 1864
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Emarginula thorektes Kilburn, 1978
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Emarginula tosaensis Habe, 1953
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Emarginula tuberculosa Libassi, 1859 - tuberculate emarginula
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Emarginula undulata Melvill & Standen, 1903
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Emarginula velascoensis Shasky, 1961
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Emarginula velascoi Rehder, 1980
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Emarginula viridibrunneis Poppe & Tagaro, 2020
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Emarginula viridicana Herbert & Kilburn, 1985
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Emarginula viridis Poppe & Tagaro, 2020
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Emarginula xishaensis Lu, 1986
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Emarginula yangeorum Poppe & Tagaro, 2020
The following species are mentioned in other databases : The Indo-Pacific Molluscan Database also mentions the following species with names in current use :[6]
- Subgenus Emarginula Lamarck, 1801
- Subgenus Emarginula (Subzeidora) Iredale, 1924
- Species brought into synonymy
- [1]
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Emarginula altilis Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
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Emarginula amitina Iredale, 1925: synonym of Emarginula dilecta A. Adams, 1852
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Emarginula angustata Thiele, 1915: synonym of Tugali decussata A. Adams, 1852
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Emarginula arabica A. Adams, 1852: synonym of Octomarginula arabica (A. Adams, 1852)
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Emarginula arconatii Issel, 1869: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
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Emarginula australis Quoy & Gaimard, 1834: synonym of Montfortia subemarginata (Blainville, 1819)
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Emarginula bella Gabb, 1865: synonym of Scelidotoma bella (Gabb, 1865)
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Emarginula biangulata Sowerby, 1901: synonym of Emarginella biangulata (G. B. Sowerby III, 1901)
- † Emarginula blainvillii Defrance, 1825: synonym of † Rimula blainvillii (Defrance, 1825)
-
Emarginula brevirimata Deshayes, 1862: synonym of Montfortula brevirimata (Deshayes, 1862)
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Emarginula clathrata A. Adams & Reeve, 1850: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
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Emarginula clathrata Pease, 1863: synonym of Emarginula dilecta A. Adams, 1852
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Emarginula conica Lamarck, 1801: synonym of Emarginula fissura (Linnaeus, 1758)
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Emarginula conica Schumacher, 1817 non Lamarck, 1801: synonym of Emarginula rosea Bell, 1824
-
Emarginula dentigera Heilprin, 1889: synonym of Hemimarginula dentigera (Heilprin, 1889)
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Emarginula elata Locard, 1898: synonym of Emarginula christiaensi Piani, 1985
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Emarginula elongata O. G. Costa, 1829: synonym of Emarginula octaviana Coen, 1939
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Emarginula eximia A. Adams, 1852: synonym of Emarginella eximia (A. Adams, 1852)
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Emarginula flindersi Cotton, 1930: synonym of Emarginula convexa Hedley, 1907
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Emarginula fuliginea Kuroda, 1941: synonym of Variemarginula variegata (A. Adams, 1852)
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Emarginula guernei Dautzenberg & Fischer, 1896: synonym of Emarginula tuberculosa Libassi, 1859
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Emarginula hedleyi Thiele, 1915: synonym of Emarginula candida A. Adams, 1852
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Emarginula incisula(sic): synonym of Emarginella incisura (A. Adams, 1852)
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Emarginula incisura A. Adams, 1852: synonym of Emarginella incisura (A. Adams, 1852)
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Emarginula kimberti Cotton, 1930: synonym of Laeviemarginula kimberti (Cotton, 1930)
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Emarginula modesta H. Adams, 1872: synonym of Hemimarginula modesta (H. Adams, 1872)
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Emarginula muelleri Forbes & Hanley, 1849: synonym of Emarginula fissura (Linnaeus, 1758)
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Emarginula ostheimerae Abbott, 1958: synonym of Octomarginula ostheimerae (Abbott, 1958)
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Emarginula panhi Quoy & Gaimard, 1834: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
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Emarginula papillosa Risso 1826: synonym of Emarginula huzardii Payraudeau, 1826
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Emarginula parmophoidea Quoy & Gaimard, 1834: synonym of Tugali elegans Gray, 1843
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Emarginula peasei Thiele, 1915: synonym of Emarginula dilecta A. Adams, 1852
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Emarginula phrygium Herbert & Kilburn, 1985: synonym of Agariste phrygium (Herbert & Kilburn, 1986) (original combination)
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Emarginula picta Dunker, 1860: synonym of Montfortula picta (Dunker, 1860)
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Emarginula pileata Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
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Emarginula pileum Heilprin, 1889: synonym of Hemimarginula pileum (Heilprin, 1889)
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Emarginula planulata A. Adams, 1852: synonym of Emarginella planulata (A. Adams, 1852)
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Emarginula polygonalis A. Adams, 1852: synonym of Hemitoma polygonalis (A. Adams, 1852)
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Emarginula pulchreclathrata Tomlin, 1932: synonym of Emarginula agulhasensis Thiele, 1925
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Emarginula pumila A. Adams, 1852: synonym of Hemimarginula pumila (A. Adams, 1852)
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Emarginula punctata A. Adams, 1852: synonym of Variemarginula punctata (A. Adams, 1852)
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Emarginula reticulata Sowerby, 1813: synonym of Emarginula fissura (Linnaeus, 1758)
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Emarginula rugosa Quoy & Gaimard, 1834: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
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Emarginula scutellata Deshayes, 1863: synonym of Octomarginula scutellata (Deshayes, 1863)
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Emarginula sibogae Schepman, 1908: synonym of Emarginella sibogae (Schepman, 1908)
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Emarginula simpla Christiaens, 1987: synonym of Hemimarginula simpla (Christiaens, 1987)
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Emarginula stellata A. Adams, 1852: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
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Emarginula subclathrata Pilsbry, 1890: synonym of Emarginula dilecta A. Adams, 1852
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Emarginula subemarginata Blainville, 1819: synonym of Montfortia subemarginata (Blainville, 1819)
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Emarginula subrugosa Thiele, 1916: synonym of Hemimarginula subrugosa (Thiele, 1916)
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Emarginula sulcifera A. Adams, 1852: synonym of Montfortulana sulcifera (A. Adams, 1852)
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Emarginula tricarinata Pilsbry, 1890: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
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Emarginula tricostata G.B. Sowerby I, 1823: synonym of Hemitoma octoradiata (Gmelin, 1791)
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Emarginula vadososinuata Yokoyama, 1922: synonym of Tugalina vadososinuata (Yokoyama, 1922)
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Emarginula vadum Barnard, 1963: synonym of Emarginula undulata Melvill & Standen, 1903
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Emarginula variegata A. Adams, 1852: synonym of Variemarginula variegata (A. Adams, 1852)
References
- "Emarginula". Integrated Taxonomic Information System.
-
Powell A W B, New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
- Piani P. (1985). Revisione del genere Emarginula Lamarck, 1801 in Mediterraneo. Lavori, Società Italiana di Malacologia 21: 193-238
- Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213
- Rolán E., 2005. Malacological Fauna From The Cape Verde Archipelago. Part 1, Polyplacophora and Gastropoda.
- Spencer, H.; Marshall. B. (2009). All Mollusca except Opisthobranchia. In: Gordon, D. (Ed.) (2009). New Zealand Inventory of Biodiversity. Volume One: Kingdom Animalia. 584 pp
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