Arenivaga erratica (Rehn & J. A. G. 1903)

Arenivaga erratica Rehn (Plate IX, figures 11 to 13.)
1903. Homoeogamia {Arenivaga) erratica Rehn, Proc. Acad. Xat. Sci. Phila., 1903, p. 187. [d^; Prescott, Arizona.]
The present insect is closely related to A. apacha; males may be separated by the distinctive dextro-ventral concealed genital plate in that species, while in erratica the interocular width is normally less than, very rarely equal to, that between the ocelli, and the pronotal marking is normally less decided and never pictured. Xo males of the present species before us have the tegmina much suffused with darker brown, a condition which occurs frequently in apacha. Females of the two species are separable by the decidedly heaier and shorter limbs in apacha, this most noticeable in the tibiae; in apacha the fringe of hairs about the cephalic margin of the body is also appreciably heavier.
This species, with A . rehni and apacha, averages decidedly smaller than A . hoUiana, but se-eral males of the present insect from central Texas are larger than the smallest males of boUiana before us. As in the other species of the group, this size xariation appears to be due rather to peculiar local environmental conditions, than to purely geographic influences.
In addition to the characters gien in the key, the following features are of diagnostic value in the male sex.
The interocular width sometimes varies decidedly; normally slightly less than that between the ocelli, we have a few specimens in which these dimensions are subequal, while rarely the interocular width is decidedly less, in one example before us equalling but one-quarter that between the ocelli. Concealed genital hook small and slender, curved gently inward; apex enlarged and blunt, bearing a sharp basal tooth on the inner side directed proximad, thus resembling the barb of a fish hook. Dextro-dorsal concealed genital plate, a large, smooth, ovate, chitinous lobe; dextro-ventral concealed genital plate a flattened, chitinous plate, which shows neither shagreenous surface (as in bolliana and rehni) nor sharp projection (as in apacha).
The adult female not having been previously recognized, we here describe that sex,
9 ; Round Mountain, Blanco County, Texas. (F. G. Schaupp.) [Academy of Natural Sciences of Philadelphia.]
Description of Female.— Wery unlike male, apterous, size much larger, form broad ovate. Dorsal surface rugulose, covered with very short, but stout, minute reddish hairs, these longest along the margins of the body, particularly cephalad. Pronotum moderately broad, cephalic margin evenly arcuate at an angle of less than 90°, caudal margin almost transverse, very weakly produced mesad, with sides showing a slight concavity. Supra-anal plate transverse, weakly trapeziform; caudal margin weakly convex laterad, with mesal fifth subtruncate, showing an indication of mesal emargination. Subgenital plate very large, roundly produced mesad.
Measurements {i)i millimeters)
-71 Length of Length of Width of Length of Width of
body pronotiim pronotum tegmen tegmen
Round Mountain, Texas(3) 15. 4-1 7. 5 5.1-5.3 7-7-8 20.3-21 7,4-7.8
Carrizo Springs, Texas. (9)5" 13. 5-18 4.2-5.7 6-8.7 16.6-22 6.2-8.8
Brownsville, Texas. ... (9) 14. 8-15. 3 4.6-4.9 6.4-6.8 16. 9-17. 6 6.3-6.4 Jemez Springs, New
Mexico (3) 1 3. 5-1 6. 5 4-4.7 5.4-6.6 17.8-20.8 7-7.8
Roebles Ranch, Arizona(3) 15-15. 8 4.4-4.6 6.4-6.7 18. 3-19. 2 6.8-7.1
Phoenix, Arizona (8) 13-15 38-4.7 5-3-6.2 I5-I7-7 5-7-^^-8
Cottonwood, California 13.8 4 5.8 15 6
•'"'• A single male furnishes the maximum measurements given, the others appro^imate much more closely the niinimuni. Q Length of Length of Width of Width of Width of
* body pronotum pronotum mesonotum nietunotum
Round Mountain, Texas 16 5.4 8.6 11 11. 6
Carrizo Springs, Texas. (6) 13.8-15.3 4.8-5.3 6.8-7.8 8.7-9.7 y. 4-10. 6
Sabinal, Texas (4) 13. 8-14. 8 5-5.2 6.7-7.9 '^■9~9-5 9-7-IO-3
Albuquerque, New
Mexico 1 1. 4 4.7 6.5 8 8.6
Adamana, Arizona ... . 12.8 5.2 8 <).7 10 Chiricahua Mountains,
Arizona 14.7 5-7 -"^5 9-9 i^-'j
Coloration. — cf. This sex shows less color variation than in other species of the genus. General coloration ochraceous-buff. A brown, shield-shaped marking is situated meso-caudad on the pronotum, this is normally finely and weakly marked with a darker shade (a conspicuous development of this darker pattern giing the allied a pacha a distinctive pictured appearance). In rare specimens of extreme recessive coloration, this blotch becomes very pale, with no darker markings whatever. In specimens of extreme intensive coloration it becomes dark brown, the markings again disappearing. In the other species of the genus, we find, in the maximum of intensive coloration, that the homologous pronotal blotch expands, covering all but the cephalic margin of the pronotum; this does not appear in any of the very large series of erratica now before us.
9 . General coloration, above tawny to burnt umber, the cephalic margin of the pronotum usually vaguely paler, all but the darkest individuals faintly maculate on the pronotum, mesonotum and metanotum with a darker shade, and laterad a spot of this same color is found, with rare exceptions, in the series before us, on each abdominal segment both above and below. Minute hairy covering tawny. I'nderparts, particularly head and limbs, somewhat paler.
This species is know^n to range from Waco, south to Goliad and Brownsville, Texas. Westward the most northern localities are Ballinger, Texas; Durango, Colorado; Mescalero Apache Reservation and Jemez Hot Springs, New Mexico; Adamana and W'inslow^ Arizona, and St. (leorge, I'tah; while in California the species is known only from single records — Cottonwood and Rierside. The species undoubtedly has an extensive distribution in Mexico,
but is there unknown aside from our single record for the state of Sonora.
Specimens Examined: 252; 133 males. 39 females and 80 immature individuals.
Waco, Texas, V, 8 to VII, 16, (Belfrage), 4 a", [M. C. Z. and Hebard Cln.].
Bosque County. Tex.. (Belfrage). i c?. [M. C. Z.].
Ballinger, Tex.. (H. Pinkus; at light), i cT. [U. S. N. M.l.
Shovel Mountain, Burnet County, Tex., IX, 10, 1901, (F. G. Schaupp), 1 d",! 9, Austin. Tex., II. 19. 1903. (V. M. Wheeler). 2 9 .'"'■^ [A. N. S. P. and Hebard Cln.]; (W. M. Wheeler), 29,1 juv. o [Am. Mus. Nat. Hist.].
Round Mountain. Tex.. (F. G. Schaupp), 3 c?. i 9 ■''' l^^S?•]•
Georgetown, Tex., (E. Palmer; in cave), 5 o", 3 9 , 5 Juv. o", [M. C. Z., Hebard Cln. and A. N. S. P.].
Goliad, Tex., VII, 28, (R. A. Cushman), 2 d" . [U. S. N. M. and Hebard Cln.].
Brownsville. Tex., IV, 30, 1895(C H. T. Townsend), 4 d, [U. S. N. M.].
Kerrville. Tex., IV, ii, 1907, (F. C. Pratt; at light). 5 o^, [U. S. N. M. and Hebard Cln.].
San Antonio, Tex., IX, 18 to 27, (E. Palmer), 5 d" , 2 9 , [M. C. Z. and Hebard Cln.].
Cotulla, Tex., IV, 18, 1906, (F. C. Pratt), i juv. 9 ; V. 12. 1906. (Crawford and Pratt), 2 c?, [all U. S. N. M.].
Esperanza Ranch, Brownsville. Tex.. V, 22 and VI, 19. (C Schaeffer), 2 cT, [A. N. S. P. and Hebard Cln.].
Sabinal. Tex., V to VI, 19, 1910, (Pratt and Pierce), 3 o^, 4 ? • 2 juv. cf , 2 juv. 9 , [U. S. N. M. and Hebard Cln.].
Knippa, Tex., VII, 3. 1910, (F. C. Pratt), i juv. o^. [U. S. X. M.].
Carrizo Springs, Tex., (A. Wadgymar), 9 <f, h 9,4 Juv. c^ 8 juv. 9 , [Hebard Cln.. U. S. N. M. and A. N. S. P.].
Ringgold Barracks, [near Rio Grande], Tex.. (Schott), i c [MC. Z.].
Devils River. Tex.. V, 4, 1907, (F. C. Pratt), i juv. d, [U. S. X. M.].
Hackberry Creek, Boquillas Road. Brewster County, Tex.. IX, 2, 1912, (Rehn and Hebard; attracted to camp fire during two hours). 3 d, [Hebard Cln. and Chisos Mountains, VI, 10 to 12, 1908, (Mitchell and Cushman), i 9, i juv. 9, [U. S. X. M.]; VII, 16, 191 1, (H. A. Wenzel), 2 d', [A. N. S. P.].
El Paso, Tex., IV, 4, 1899, (H. Solton), i juv. cf ; VIII, (G. W Dunn), i c?, [all Hebard Cln.].
Durango, Colorado, (E. J. Oslar), I c?, [U. S. X. M.].
Mescalero Apache Reservation, New Mexico, X, 2. 1896. (T. D. A. Cockcrell), I juv. c^. [M. C. Z.].
362 Incorrectly recorded by Rehn as bolliana. Proc. Acad. Nat. Sci. Phila., 1903,
p. 187, (1903)-
363 Incorrectly recorded by Rehn as bolliana. Trans. Am. Ent. Soc. xxvn, p. 331, (1902). Proc. Acad. Nat. Sci. Phila., 1903, p. 187, (1903).
Highrolls, Sacramento Mountains, N. M., 'I, 23, 1902, (H. L. Viereck), i juv.
d^3«^ [A. X. s. p.].
Jemez Hot Springs, X. M., 'II, 29 to IX, 2, 1913 and 1915, (J. Woodgate), 3 cf, IHebard Cln.].
Albuquerque, X. M.. V, 1905, (V. M. Wheeler), i 9 , [M. C. Z.]; (H. Wickham). I juv. 9, [U. S. X. M.].
Las Cruces. X. M.. IX. i,S95. (T. D. A. Cockerell; at light^ss), 4 ^, [M. c. Z. and Hebard Cln.].
Deming, X. M., 'II. is to 20. i()07, (Rehn and Hebard; at light), 1 1 cf, [Hebard Cln.].
Sheridan Canyon, Hachita C.rande Mountains, X. M., (H. A. Pilsbry), i d" , [A. X. Adamana, Arizona. 'II. 22, 1907. (Miss Stella McSaws), i 9 , [Hebard Cln.].
Winslow, Ariz.. (H. Wickham). i juv. cf , [Hebard Cln.].
Fort Grant, Graham County, Ariz., 1882, 3 d", [U. S. X. M. and Hebard Cln.].
San Simon. Ariz.. '. 7, (H. G. Hubbard), i cf , [U. S. X. M.].
Paradise. Ariz.. VIII, 10, 1915, (O. C. Poling), 2 a^, [Hebard Cln.].
Chiricahua Mountains, Ariz., i 9, i juv. 9, [A. X. S. P.].
Roosevelt, Ariz., 2 cf, [Cornell Univ. and Hebard Cln.].
Redington, Ariz.. I cT, [U. S. X. M.].
Lowell Ranger Station, Pima County, Ariz., 'II, 6 and 20, 1916, (Lutz and Rehn), 5 a^, [A. M. X. H. and A. X. S. P.].
Sabino Basin, Santa Catalina Mountains, Arizona. 'III, 151021, 1916, (Lutz). 3 cf , I 9 , [A. M. X. H. and A. X. S. P.].
Sabino Canon, Santa Catalina Mountains. Ariz., 11, 22 to III, 15, 1916. (J. F. Tucker), 6 9 . 44 juv.; 'I, 5 to XI. 2, 1915 and 1916, (J. F.Tucker), 4 o [all Hebard Cln.].
Tucson, Ariz., (H. Wickham). i 9, [Hebard Cln.]; VH, 21 to 2^,, 1916. (Lutz and Rehn), i d" , [A. M. X. H.].
San Xaier. near Tucson, Ariz.. TI. 24, 1916, (Lutz and Rehn), 2 cf . [A. M. X. H. and A. X. S. P.].
Snyder's Hill, Pima Count-, Ariz., c. 2500 feet, X, 11, 1910, (Rehn and Hel)ard; attracted to camp fire), i c . [Hebard Cln.].
Roebles Ranch. Pima County. Ariz., VHI, 13. 191 6, (Lutz and Rehn), 2 o'', [A. M. X. H. and A. X. S. P.]; X, 5, 1910, (Rehn and Hebard; attracted to camp fire), 3 cf . [Hebard Cln. and A. X. S. P.].
Palo Alto Ranch. Pima County. Ariz., VII, 29 and 30, 1916. (Lutz and Rehn), I cf, [A. M. X. H.].
Coyote Mcnmtains, Ariz.. X'lII. 4 to 7, 191 6. (Lutz and Rehn). i 9, [.. M. '^ Incorrectly recorded and described by Rehn as immature subdiaphana. Proc. Acad. Xat. Sci. Phila., 1903, pp. 190 to 191, (1903).
"^ One specimen incorrectly recorded by Scudder and Cockerel! as boUiana. Proc. Davenport Acad. Sci., ix, p. 19, (1901).
Kits Peak Rincon, Baboquivari Mountains, Ariz., VIII, 7 to 9, 191 6. (Lutz and Rehn). 2 cf , [A. M. N. H. and A. N. S. P.].
Santa Cruz Village, Comobabi Mountains, Ariz., VIII, 10 to 12, 191 6, (Lutz and Rehn). i cf . [A. M. N. H.].
Prescott, Ariz., VI, 10, 1902, (E. J. Oslar), 2 cT, type, paratype, [A. N. S. P.].
Florence, Ariz.. VI, 8, 1903, (C. R. Biedennan), i juv. d' ; VII, 17 to 22. 1903, (C. R. Biederman), 11 &, [A. N. S. P.].
Phoenix, Ariz., VI, 17, 1892, (A. B. Cordley), i d'; VIII, 1890. i juv. cT ; IX, 14 to X, 9, 1903 and 1904, (R. E. Kunze), 7 o", [Hebard Cln. and A. N. S. P.].
Fort Mojave, Ariz., Ill, 25, 191 1, i d", [Hebard Cln.].
Yuma, Ariz., VII, 27, 1907, (Rehn and Hebard; at light), i o". [Hebard Cln.].
St. George, Utah, 1875, (E. Palmer), 49,1 juv. o i juv. 9. [M. C. Z. and Hebard Cln.].
Cottonwood, California, 2274 feet, IX, 9, 1907, (Hebard; at light on train), I cf, [Hebard Cln.].
Riverside, Cal., (H. Wickham), i 9, [Hebard Cln.].
Sonora, Mexico, i cf , [Hebard Cln.].

Arenivaga erratica, commonly known as the erratic sand cockroach, is a species of cockroach in the family Corydiidae. It is found in Central America and North America.

Male. Measurements. Holotype stand-in TL = 17.3 mm, GW = 8.8 mm, PW = 5.73 mm, PL = 3.76 mm, TL/GW = 1.97, PL/PW = 0.66. EW = 0.25 mm; OW = 0.40 mm. Among paratypes range of TL 16.3–23.5 mm; range of GW 7.4–10.0 mm; range of PW 5.10–6.62 mm; range of PL 3.76–4.70 mm. Head. Two ocelli large, ovoid and protruding (0.40 × 0.30 mm); vertex medium brown, with small ridges between apices of eyes extending on to ocellar tubercles; interocellar space slightly concave, medium brown with light brown medial line; two oval indentations laterally at base of interocellar space. Frons waxy white, flat; bound on either side by ridges extending from inner apex of ocelli outwards to lateral edges of clypeus; scattered long setae on frons and ridges. Anterior portion of frons waxy white, bulbous; clypeal suture demarcates waxy white anteclypeus. See Fig. 48d. Pronotum. Pronotum translucent waxy beige; variable length orange-brown setae along anterior margin; dorsal surface of pronotum covered with short orange-brown setae that are denser and longer anteriorly and laterally; pronotal pattern medium orange-brown “panther face”, but runs through every shade to dark brown in other specimens; with no aura and little discernible detail, though considerable detail seen in some specimens. See Fig. 48c. Body. Wing brace present. Legs and body light orange-brown; subgenital plate asymmetrical with posterior edge emarginated, rounded apices. See Fig. 48b. Forewings. Wings extended beyond abdominal apex (up to ~35% of total wing length); color highly variable from light beige, to light brown, to light orange-brown, to medium brown; usually blotchy; surface semi-transparent and matte, or with faint sheen on many specimens. See Fig. 48a. Genitalia. Right dorsal phallomere composed of bulbous lightly sclerotized hook-shaped lobe, articulated with right ventral phallomere on lateral side; central field lightly sclerotized; medial margin sclerotized, shagreened, often with one or two small dorsally projecting teeth; medial margin extends beyond rest of phallomere at each end. Small central sclerite punctate, with sclerotized shagreened ridge, toothed along edge, along anterolateral side. Right ventral phallomere extends from articulation into bulbous love that narrows anteriorly; after narrow gap, rounded, punctate arm extending to depth of rest of phallomere. Genital hook with pointed head and moderate hook; arm moderate with smooth curve. See Fig. 49.

This is a widespread species widely in New Mexico, the eastern half of Arizona, the far western part of Texas located south of New Mexico, south into central Mexico. Isolated records from Utah, Nevada, and a record from Colorado not included in the distribution map. See Fig. 50.