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Japanese Angelica Tree

Aralia elata (Miq.) Seem.

Description

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Shrubs or small trees, 2-5(-8) m tall, andromonoecious. Branches armed with sparse prickles. Leaves 2(or 3)-pinnately compound, with a pair of accessory leaflets at each division of rachis; petiole to ca. 50 cm, glabrous or pubescent, prickly; petiolules 3-5 mm; leaflets 5-11(-13) per pinna, broadly ovate to elliptic-ovate or narrowly ovate, 5-12(-19) × 2.5-8 cm, membranous, papery, or subleathery, abaxially glabrous or light yellow or gray pubescent, sometimes or more densely pubescent on veins, or sometimes glabrescent, adaxially sparsely strigose, secondary veins 6-10 pairs, raised abaxially, conspicuous adaxially, base cordate to subcordate or rounded, margin serrulate, apex acuminate. Inflorescence a terminal panicle of umbels, densely yellow-brown or gray pubescent, unarmed; primary axis 1-5 cm; secondary axes 20-35 cm; ultimate axes with a terminal umbel of bisexual flowers and 1 to several lateral umbels of male flowers; bracts persistent, lanceolate to subulate, 2-5 mm, sometimes ciliate; umbels 6-15-flowered; pedicels 1-10 mm, densely pubescent. Ovary 5-carpellate; styles 5, free or united to middle. Fruit globose, ca. 3-4 mm in diam.; styles persistent. Fl. Jul-Sep, fr. Sep-Dec. 2n = 24*.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 13: 480, 481, 484 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Distribution

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Anhui, Fujian, S Gansu, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, N Shandong, Shanxi, Sichuan, Yunnan, Zhejiang [Japan, Korea, E Russia].
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 13: 480, 481, 484 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
project
eFloras.org
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eFloras

Habitat

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Forests, forest margins, scrub fields, roadsides; near sea level to 2700 m.
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cc-by-nc-sa-3.0
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 13: 480, 481, 484 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
project
eFloras.org
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Cyclicity

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Flowering from July to September.
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Wen, Jun
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Wen, Jun
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Diagnostic Description

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Aralia elata differs from A. chinensis L. in the former’s less pubescent leaflets and inflorescences, glaucous (vs. green) abaxial leaflet surface, coronulate (vs. striated) cuticles on abaxial leaflet surface, fewer flowers (7-25 vs. 20-50) and shorter pedicels of terminal umbels [1.5-10 mm (usually 4-7 mm) vs. 10-20 mm (usually 12-15 mm)]. Aralia elata closely related to A. dasyphylloides. The major differences between these two species lie in the aggregation of flowers into heads (A. dasyphylloides) or umbels (A. elata), and densely tomentose (A. dasyphylloides) or somewhat pubescent to glabrous (A. elata) leaves.
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Distribution

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Aralia elata is the most widely distributed species within Aralia, occurring in Anhui, Chongqing, Fujian, Gansu, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan and Zhejiang provinces of China, Japan and Korea.
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Evolution

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The phylogeny of Aralia sect. Dimorphanthus has been inferred using the ITS sequences of nuclear ribosomal DNA (Wen, 2000). The ITS phylogeny suggests a close relationship among Aralia elata, A. dasphylloides, and A. scaberula. Aralia elata and A. dasphylloides have largely allopatric distributional areas with overlapping boundaries. A. dasphylloides is mostly distributed in southern China and also occurs (but is not common) in eastern and central China. A. elata has a more northern distribution and is most frequent in central and eastern China and Japan. It seems reasonable to hypothesize that A. elata perhaps gave rise to A. dasphylloides via geographical speciation (Wen, 2004). Based on the morphological variation, Wen (1993) provided a generic delimitation of Aralia, in which Arala elata belong to section Dimorphanthus.
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General Description

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Shrubs or treelets, 1.5-7 m tall. DBH up to 15 cm. Stem usually pricky, rarely inermis, sometime pickles in rows nearly encircling the stem; bark brownish gray or gray. Leaves bipinnate, 70-125 cm long, 50-90 cm wide, with 3-5 pinnae, each 5-15-foliolated, 20-45 cm long; stipules basal or nearly so, narrowly triangular, 1-1.5 cm long; petioles pubescent or becoming nearly glabrescent, umarmed, greenish; rachises subtended with a pair of accessory leaflets, rarely with a pair of 5-foliolated accessory pinnae; leaflets ovate to broadly ovate, sometimes narrowly ovate to lanceolate, chartaceous, 6-15 cm long, 3.5-8.5 cm wide, acute to acuminate at apex, rounded to obtuse, occasionally subcordate at base, serrulate to serrate, lateral veins 7-9 pairs, adaxial surfaces green, slightly rugose, sometimes scabrid, abaxial surfaces glaucous or brownish glaucous, pubescent to densely pubescent, sometimes pilose, pilose on veins, or glabrous, petiolules 0-6 mm long. Inflorescences terminal, 25-55 cm long, 35-80 cm wide, umbellately paniculate, branching in 3 to 4 orders, lacking a main axis or with a very short one, less than 12 cm long, consisting of 5-8 primary branches, each 30-55 cm long, yellowish green when young, turning into pink at fruiting, bracts and bracteoles persistent, secondary bracts lanceolate to triangular, ciliate, 4-10 mm long, bracteoles lanceolate, ciliate, 1.5-2 mm long; umbels 6-15-flowered, pedicels 1-6 mm long, pubescent. Floral buds yellowish green. Flowers greenish white. Sepals minute, triangular to rounded. Petals ovate, 1.8-2.2 mm long. Filaments 2-2.5 mm long; anthers yellowish white, oblong, 0.7-1.2 mm long. Ovary 5-locular; styles distinct. Floral disc projected. Fruit globose to subglobose, ca. 4 mm in diameter, purplish black.
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Genetics

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There are some reports for the chromosomal data of Aralia elata. All counts are 2n = 24 (Sokolovskaya, 1966; Sun et al., 1988; Wu, 1995; Yi et al., 2004). The base chromosome number of A. elata has been suggested to be x = 12 (Wen, 2002).
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Habitat

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Growing in hillside, roadside, streamsides, disturbed areas, forest edges, along ravines, thickets and secondary forest; from sea level to 2100 m.
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Morphology

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Aralia elata is characterized by its lack of a main inflorescence axis or with a very short one (less than 12 mm long), secondary inflorescence braches consisting of many umbels (more than 7, rarely a few), leaflet margins serrulate to serrate, petioles usually unarmed, and bracts small (secondary bracts 4-10 mm long) and narrowly triangular.
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Aralia elata

provided by wikipedia EN

Aralia elata, the Japanese angelica tree,[1] Chinese angelica-tree,[2] or Korean angelica-tree,[3] is a woody plant belonging to the family Araliaceae. It is known as tara-no-ki (タラノキ; 楤木) in Japanese, and dureup-namu (두릅나무) in Korean.

Description

It is an upright deciduous small tree or shrub growing up to 10 m (33 ft) in height,[4] native to eastern Russia, China, Korea, and Japan.

The bark is rough and gray with prickles. The leaves are alternate, large, 60–120 cm long, and double pinnate. The flowers are produced in large umbels in late summer, each flower small and white. The fruit is a small black drupe.

Aralia elata is closely related to the American species Aralia spinosa, with which it is easily confused.

Cultivation

Aralia elata is cultivated, often in a variegated form, for its exotic appearance. It prefers deep loamy soils in partial shade, but will grow in poorer soils and in full sun. The cultivars 'Variegata'[5] and 'Aureovariegata'[6] have gained the Royal Horticultural Society's Award of Garden Merit.

Gastronomy

Japan

Tara-no-me on a plate

In Japan, the shoots are called tara-no-me and are eaten in the spring. They are picked from the end of the branches and are commonly fried in a tempura batter. They can also be blanched or stir fried, among other methods.

Korea

Dureup on sokuri (bamboo basket)

In Korean, the young shoot is called dureup (두릅), and the plant is called dureupnamu (두릅나무, "dureup tree"). Young shoots are harvested during a month, from early April to early May, when they are soft and fragrant. In Korean cuisine, the shoots are commonly eaten blanched as namul, pickled as jangajji, pan-fried as jeon, or deep-fried as bugak.

Invasive species

Aralia elata in the vicinity of Philadelphia, PA.

The tree was introduced into the United States in 1830. Birds like to eat the fruits, and are spreading its seeds, allowing the tree to expand its range as an invasive species in the Northeastern United States.[7]

References

  1. ^ "Angelica elata". www.rhs.org. Royal Horticultural Society. Retrieved 5 May 2020.
  2. ^ "Aralia elata". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA).
  3. ^ Korea National Arboretum (2015). English Names for Korean Native Plants (PDF). Pocheon: National Arboretum. p. 357. ISBN 978-89-97450-98-5. Archived from the original (PDF) on 25 May 2017. Retrieved 6 December 2016 – via Korea Forest Service.
  4. ^ "RHS Plant Selector - Aralia elata". Retrieved 29 May 2013.
  5. ^ "Aralia elata 'Variegata'". RHS. Retrieved 12 April 2020.
  6. ^ "Aralia elata 'Aureovariegata'". RHS. Retrieved 12 April 2020.
  7. ^ U.S. Dept. of Agriculture: Map of distribution in U.S. and Canada
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Aralia elata: Brief Summary

provided by wikipedia EN

Aralia elata, the Japanese angelica tree, Chinese angelica-tree, or Korean angelica-tree, is a woody plant belonging to the family Araliaceae. It is known as tara-no-ki (タラノキ; 楤木) in Japanese, and dureup-namu (두릅나무) in Korean.

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