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Onthophagus
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Onthophagus pennsylvanicus Harold 1871
collect
overview
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maps
names
Scientific Names
Preferred Names
Onthophagus pennsylvanicus Harold 1871
Species recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus pennsylvanicus
Species recognized by
wikipedia русскую Википедию
,
wikipedia VI
,
wikipedia ID
,
Mexico Species List
,
North Atlantic Species List
,
NCBI
,
wikipedia MIN
, and
United States Species List
Onthophagus pennsylvanicus
Harold 1871
Species recognized by
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
, and
GBIF classification
Onthophagus (Onthophagus) pennsylvanicus Harold 1871
Species recognized by
Catalogue of Life 26 March 2024
Onthophagus moeris
Recognized by
BHL data coverage
Onthophagus pennsylvanicus
Species recognized by
wikidata_hierarchy tar gz
,
wikipedia ES
,
wikipedia PT
,
wikipedia EN
,
Barcode of Life Data Systems
,
Global Biotic Interactions
,
Global Biotic Interactions
, and
iNaturalist data coverage
Onthophagus pennsylvanicus Harold 1871
Species recognized by
iNaturalist
,
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
,
GBIF data coverage
,
Dynamic Hierarchy Version 2.2.3
, and
EOL Dynamic Hierarchy Version 2.2.4
Alternative Names
Onthophagus moeris Sturm 1826
Not accepted according to
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus moeris
Sturm 1826
Synonym according to
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
Onthophagus falcipes Harold 1871
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Onthophagus moeris Sturm 1826
Not accepted according to
Dynamic Hierarchy Version 2.2.3
and
EOL Dynamic Hierarchy Version 2.2.4
Onthophagus moeris Sturm 1826
Synonym according to
Catalogue of Life 26 March 2024
Common Names
There are no common names associated with this taxon.
Curated hierarchies for Onthophagus pennsylvanicus Harold 1871
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Onthophagus
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D' Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D' Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D' Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D' Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger MacLeay 1888
Onthophagus cuniculus MacLeay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D' Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D' Orbigny 1902
Onthophagus flavimargo D' Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D' Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D' Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D' Orbigny 1908
Onthophagus juncticornis D' Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D' Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D' Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D' Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D' Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus undaticeps D' Orbigny 1902
Onthophagus viridivinosus Kohlmann & Solis 2001
2069 additional siblings truncated for brevity.
See the
resource file
for a full list.
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Onthophagini Streubel 1846
Onthophagus Latreille 1802
Onthophagus subgen. Onthophagus Latreille 1802
Onthophagus (Onthophagus) pennsylvanicus Harold 1871
(this page)
Onthophagus (Onthophagus) abacus Boucomont 1921
Onthophagus (Onthophagus) abas Balthasar 1946
Onthophagus (Onthophagus) abmisibilus Krikken & Huijbregts 2012
Onthophagus (Onthophagus) abreui Arrow 1931
Onthophagus (Onthophagus) abruptus Orbigny 1913
Onthophagus (Onthophagus) absyrtus Balthasar 1946
Onthophagus (Onthophagus) abyssinicus Gillet 1925
Onthophagus (Onthophagus) academus Balthasar 1946
Onthophagus (Onthophagus) aceroides Krikken & Huijbregts 2013
Onthophagus (Onthophagus) acerus Gillet 1930
Onthophagus (Onthophagus) aciculatulus Blatchley 1928
Onthophagus (Onthophagus) acrisius Balthasar 1946
Onthophagus (Onthophagus) acuminatus Harold 1880
Onthophagus (Onthophagus) adelaidae Hope 1846
Onthophagus (Onthophagus) adelphus Gillet 1930
Onthophagus (Onthophagus) admetus Balthasar 1946
Onthophagus (Onthophagus) adornatus Orbigny 1904
Onthophagus (Onthophagus) aegrotus Balthasar 1967
Onthophagus (Onthophagus) aemulus Gillet 1930
Onthophagus (Onthophagus) aeneoniger Orbigny 1913
Onthophagus (Onthophagus) aeneopiceus Orbigny 1902
Onthophagus (Onthophagus) aequepubens Orbigny 1905
Onthophagus (Onthophagus) aereidorsis Orbigny 1902
Onthophagus (Onthophagus) aeremicans Orbigny 1904
Onthophagus (Onthophagus) aereomaculatus Boucomont 1914
Onthophagus (Onthophagus) aerestriatus Orbigny 1913
Onthophagus (Onthophagus) aeruginosus Roth 1851
Onthophagus (Onthophagus) aesopus Lansberge 1882
Onthophagus (Onthophagus) affinis Gillet 1930
Onthophagus (Onthophagus) africanus Lansberge 1886
Onthophagus (Onthophagus) agaricophilus Arrow 1931
Onthophagus (Onthophagus) ahenicollis Orbigny 1902
Onthophagus (Onthophagus) ahenomicans Orbigny 1902
Onthophagus (Onthophagus) akamineae Moretto & Génier 2020
Onthophagus (Onthophagus) akhaus Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Onthophagus) alaindrumonti Krikken & Huijbregts 2012
Onthophagus (Onthophagus) albicomus Orbigny 1908
Onthophagus (Onthophagus) albicornis (Palisot de Beauvois 1805)
Onthophagus (Onthophagus) albipennis Péringuey 1908
Onthophagus (Onthophagus) albipodex Orbigny 1902
Onthophagus (Onthophagus) alfuricus Huijbregts & Krikken 2012
Onthophagus (Onthophagus) aliceae Josso 2019
Onthophagus (Onthophagus) alienus Frey 1958
Onthophagus (Onthophagus) allojavanus Huijbregts & Krikken 2011
Onthophagus (Onthophagus) alluaudi Orbigny 1902
Onthophagus (Onthophagus) alluvius Howden & Cartwright 1963
Onthophagus (Onthophagus) aloysiellus Zunino 1977
Onthophagus (Onthophagus) alquirta Matthews 1972
Onthophagus (Onthophagus) alternans Raffray 1877
Onthophagus (Onthophagus) altilamina Orbigny 1905
Onthophagus (Onthophagus) altivagans Howden & Génier 2004
Onthophagus (Onthophagus) ambang Krikken & Huijbregts 2011
Onthophagus (Onthophagus) amoenus Orbigny 1908
Onthophagus (Onthophagus) amphicoma Boucomont 1914
Onthophagus (Onthophagus) amphinasus Arrow 1931
Onthophagus (Onthophagus) amphioxus Arrow 1933
Onthophagus (Onthophagus) amplipennis Orbigny 1905
Onthophagus (Onthophagus) arnetti Howden & Cartwright 1963
Onthophagus (Onthophagus) batesi Howden & Cartwright 1963
Onthophagus (Onthophagus) brevifrons Horn 1881
Onthophagus (Onthophagus) browni Howden & Cartwright 1963
Onthophagus (Onthophagus) canelasensis Howden & Génier 2004
Onthophagus (Onthophagus) cartwrighti Howden 1973
Onthophagus (Onthophagus) cavernicollis Howden & Cartwright 1963
Onthophagus (Onthophagus) chevrolati Harold 1869
Onthophagus (Onthophagus) coahuilae Zunino & Halffter 1988
Onthophagus (Onthophagus) cochisus Brown 1927
Onthophagus (Onthophagus) concinnus Castelnau 1840
Onthophagus (Onthophagus) coproides Horn 1881
Onthophagus (Onthophagus) cuevensis Howden 1973
Onthophagus (Onthophagus) cyanellus Bates 1887
Onthophagus (Onthophagus) cynomysi Brown 1927
Onthophagus (Onthophagus) fuscus Boucomont 1932
Onthophagus (Onthophagus) hajimei Masumoto 1984
Onthophagus (Onthophagus) hayashii Masumoto 1991
Onthophagus (Onthophagus) hecate (Panzer 1794)
Onthophagus (Onthophagus) hidalgus Zunino & Halffter 1988
Onthophagus (Onthophagus) hoepfneri Harold 1869
Onthophagus (Onthophagus) incensus Say 1835
Onthophagus (Onthophagus) knausi Brown 1927
Onthophagus (Onthophagus) landolti Harold 1880
Onthophagus (Onthophagus) lecontei Harold 1871
Onthophagus (Onthophagus) masumotoi Ochi 1985
Onthophagus (Onthophagus) mcclevei Howden & Génier 2004
Onthophagus (Onthophagus) medorensis Brown 1929
Onthophagus (Onthophagus) mendicus Gillet 1924
Onthophagus (Onthophagus) mextexus Howden & Cartwright 1970
Onthophagus (Onthophagus) oklahomensis Brown 1927
Onthophagus (Onthophagus) orpheus (Panzer 1794)
Onthophagus (Onthophagus) polyphemi Hubbard 1894
Onthophagus (Onthophagus) pseudofuscus Zunino & Halffter 1988
Onthophagus (Onthophagus) sakainoi Masumoto 1991
Onthophagus (Onthophagus) schaefferi Howden & Cartwright 1963
Onthophagus (Onthophagus) setchan Masumoto 1984
Onthophagus (Onthophagus) subopacus Robinson 1940
Onthophagus (Onthophagus) subtropicus Howden & Cartwright 1963
Onthophagus (Onthophagus) taurus (Schreber 1759)
Onthophagus (Onthophagus) tuberculifrons Harold 1871
Onthophagus (Onthophagus) velutinus Horn 1875
Onthophagus (Onthophagus) yangi Masumoto, Tsai & Ochi 2006
1593 additional siblings truncated for brevity.
See the
resource file
for a full list.
Dynamic Hierarchy Version 2.2.3
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D'Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D'Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D'Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D'Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger MacLeay 1888
Onthophagus cuniculus MacLeay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D'Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D'Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D'Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D'Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D'Orbigny 1908
Onthophagus juncticornis D'Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D'Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D'Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus piceiceps D'Orbigny 1904
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D'Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus sylvipapuanus Krikken & Huijbregts 2012
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D'Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus viridivinosus Kohlmann & Solis 2001
2034 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Onthophagus
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D' Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D' Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D' Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D' Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger Macleay 1888
Onthophagus cuniculus Mac Leay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D' Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D' Orbigny 1902
Onthophagus flavimargo D' Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D' Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D' Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D' Orbigny 1908
Onthophagus juncticornis D' Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D' Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D' Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D' Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D' Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus undaticeps D' Orbigny 1902
Onthophagus viridivinosus Kohlmann & Solis 2001
2071 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy Version 2.2.4
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D'Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D'Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D'Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D'Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger MacLeay 1888
Onthophagus cuniculus MacLeay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D'Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D'Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D'Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D'Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D'Orbigny 1908
Onthophagus juncticornis D'Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D'Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D'Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus piceiceps D'Orbigny 1904
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D'Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus sylvipapuanus Krikken & Huijbregts 2012
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D'Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus viridivinosus Kohlmann & Solis 2001
2034 additional siblings truncated for brevity.
See the
resource file
for a full list.
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Onthophagus Latreille 1802
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus akhaus Masumoto, Ochi & Hanboonsong 2007
Onthophagus amicus (Gillet 1925)
Onthophagus anthracinus Harold 1873
Onthophagus antoinei Walter 1990
Onthophagus apunneea Masumoto, Ochi & Hanboonsong 2007
Onthophagus aztecus Zunino & Halffter 1988
Onthophagus babaulti D' Orbigny 1915
Onthophagus babirussa (Eschscholtz 1822)
Onthophagus bambra Matthews 1972
Onthophagus basilewskyi Frey 1961
Onthophagus bassariscus Zunino & Halffter 1988
Onthophagus batesi Howden & Cartwright 1963
Onthophagus bayeri Balthasar 1942
Onthophagus bennigseni Paulian 1937
Onthophagus bidentatus Drapiez 1819
Onthophagus bilingula Balthasar 1965
Onthophagus bituberifrons Josso 2018
Onthophagus campestris Masumoto, Kiuchi & Ho 2018
Onthophagus canescens Zunino & Halffter 1988
Onthophagus chilapensis Gasca-Álvarez, Zunino & Deloya 2018
Onthophagus curvicornis Latreille 1812
Onthophagus curvifrons D' Orbigny 1906
Onthophagus curvispina Reitter 1892
Onthophagus cyaneiceps D' Orbigny 1913
Onthophagus cyanellus Bates 1887
Onthophagus cyaneoniger D' Orbigny 1902
Onthophagus cyanochlorus D' Orbigny 1902
Onthophagus dorsofasciatus Fairmaire 1893
Onthophagus flaviclava D' Orbigny 1902
Onthophagus formosanus Gillet
Onthophagus fukuyamai Ochi, Kon & Ueda 2018
Onthophagus gagates Hope 1831
Onthophagus genuinus Kohlmann & Solis 2001
Onthophagus gibbicollis Lansberge 1885
Onthophagus ginyunensis Všetečka 1942
Onthophagus glasunowi Koshantschikov 1894
Onthophagus graniceps D' Orbigny 1908
Onthophagus granosus D' Orbigny 1913
Onthophagus granulatus Boheman 1858
Onthophagus granulifer Harold 1886
Onthophagus granulifrons Frey 1961
Onthophagus granulum D' Orbigny 1904
Onthophagus granum Lansberge 1885
Onthophagus graphicus Wallengren 1881
Onthophagus grataehelenae Kohlmann & Solis 2001
Onthophagus hingstoni Arrow 1931
Onthophagus hiroyukii Ochi 2007
Onthophagus inediapterus Kohlmann & Solis 2001
Onthophagus iumienus Masumoto, Ochi & Hanboonsong 2007
Onthophagus japonicus Harold 1875
Onthophagus kachinicus Kabakov 2006
Onthophagus kawaharai Ochi & Kon 2007
Onthophagus khonkaenus Masumoto, Ochi & Hanboonsong 2008
Onthophagus kivuensis Walter 2018
Onthophagus kuraruanus Matsumura 1938
Onthophagus lenzii Harold 1875
Onthophagus limonensis Kohlmann & Solis 2001
Onthophagus longipilis D' Orbigny 1905
Onthophagus luctuosus Boucomont 1914
Onthophagus ludio Boucomont 1914
Onthophagus lugens Fahraeus 1857
Onthophagus lugubris Fahraeus 1857
Onthophagus lunulifer Boucomont 1914
Onthophagus marginipennis Josso 2018
Onthophagus millingeni D' Orbigny 1898
Onthophagus monforti Josso & Prévost 2006
Onthophagus mycetorum Zunino & Halffter 1988
Onthophagus nemorivagus Kohlmann & Solis 2001
Onthophagus nubilus Kohlmann & Solis 2001
Onthophagus obenbergeri Balthasar 1965
Onthophagus oberthueri d'Orbigny 1898
Onthophagus obliquus (Olivier 1789)
Onthophagus papulatorius Kabakov 2008
Onthophagus parachandrai Ochi, Kon & Tsubaki 2009
Onthophagus pexatus Harold 1869
Onthophagus philippinensis Boucomont 1919
Onthophagus phoenicocerus Lea 1923
Onthophagus picatus D' Orbigny 1902
Onthophagus piceiceps D' Orbigny 1904
Onthophagus picipennis Hope 1841
Onthophagus pictus Reitter 1892
Onthophagus pseudoarai Ochi & Kon 2018
Onthophagus rojkoffianus Josso 2018
Onthophagus schoolmeestersi Ochi & Kon 2007
Onthophagus seniculus (Fabricius 1781)
Onthophagus shimba Cambefort 1980
Onthophagus shinichii Ochi & Kon 2018
Onthophagus simplex Raffray 1877
Onthophagus simpliciceps D' Orbigny 1905
Onthophagus simplicifrons Reitter 1892
Onthophagus simulator D' Orbigny 1905
Onthophagus sinuosicollis D' Orbigny 1905
Onthophagus sinuosus D' Orbigny 1913
Onthophagus sisyphoides Krikken 1977
Onthophagus skelleyi Sanchez-Huerta, Zunino & Halffter 2018
Onthophagus tigrinus D' Orbigny 1908
Onthophagus tungkamangensis Masumoto, Ochi & Hanboonsong 2007
Onthophagus ulula Balthasar 1966
Onthophagus zimmermanni Balthasar 1959
Onthophagus zumpti Frey 1954
2113 additional siblings truncated for brevity.
See the
resource file
for a full list.
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Onthophagini Burmeister 1846
Onthophagus Latreille 1802
Onthophagus subgen. Onthophagus Latreille 1802
Onthophagus pennsylvanicus Harold 1871
(this page)
Onthophagus aciculatulus Blatchley 1928
Onthophagus alluvius Howden & Cartwright 1963
Onthophagus antivagans Howden & Génier 2004
Onthophagus arnetti Howden & Cartwright 1963
Onthophagus batesi Howden & Cartwright 1963
Onthophagus brevifrons Horn 1881
Onthophagus browni Howden & Cartwright 1963
Onthophagus canelasensis Howden & Génier 2004
Onthophagus cartwrighti Howden 1973
Onthophagus cavernicollis Howden & Cartwright 1963
Onthophagus chevrolati Harold 1869
Onthophagus coahuilae Zunino & Halffter 1988
Onthophagus cochisus Brown 1927
Onthophagus concinnus Laporte 1840
Onthophagus coproides Horn 1881
Onthophagus cribricollis Horn 1881
Onthophagus cuevensis Howden 1973
Onthophagus cyanellus Bates 1887
Onthophagus cynomysi Brown 1927
Onthophagus depressus Harold 1871
Onthophagus fuscus Boucomont 1932
Onthophagus hecate (Panzer 1794)
Onthophagus hidalgus Zunino & Halffter 1988
Onthophagus hoepfneri Harold 1869
Onthophagus incensus Say 1835
Onthophagus knausi Brown 1927
Onthophagus knulli Howden & Cartwright 1963
Onthophagus landolti Harold 1880
Onthophagus lecontei Harold 1871
Onthophagus mcclevei Howden & Génier 2004
Onthophagus medorensis Brown 1929
Onthophagus mextexus Howden & Cartwright 1970
Onthophagus nuchicornis (Linnaeus 1758)
Onthophagus oklahomensis Brown 1927
Onthophagus orpheus (Panzer 1794)
Onthophagus polyphemi Hubbard 1894
Onthophagus pseudofuscus Zunino & Halffter 1988
Onthophagus schaefferi Howden & Cartwright 1963
Onthophagus striatulus (Palisot de Beauvois 1809)
Onthophagus subaeneus (Palisot de Beauvois 1811)
Onthophagus subopacus Robinson 1940
Onthophagus subtropicus Howden & Cartwright 1963
Onthophagus taurus (Schreber 1759)
Onthophagus tuberculifrons Harold 1871
Onthophagus velutinus Horn 1875
NCBI
root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Scarabaeinae incertae sedis
Onthophagus
Onthophagus pennsylvanicus
(this page)
Onthophagus alcyon
Onthophagus andalusicus
Onthophagus anthracinus
Onthophagus apterus
Onthophagus atripennis
Onthophagus babirussa
Onthophagus babirussoides
Onthophagus baolocensis
Onthophagus batesi
Onthophagus bicallosus
Onthophagus bidentatus
Onthophagus bivertex
Onthophagus bonasus
Onthophagus borneensis
Onthophagus capella
Onthophagus carbonarius
Onthophagus cervicapra
Onthophagus cervus
Onthophagus championi
Onthophagus clypeatus
Onthophagus coenobita
Onthophagus consentaneus
Onthophagus crassicollis
Onthophagus cuniculus
Onthophagus curvicornis
Onthophagus dama
Onthophagus damaki
Onthophagus diabolicus
Onthophagus dunningi
Onthophagus falculatus
Onthophagus fimetarius
Onthophagus fodiens
Onthophagus furcaticeps
Onthophagus furcatus
Onthophagus gibbulus
Onthophagus giraffa
Onthophagus glabratus
Onthophagus gracilipes
Onthophagus grossepunctatus
Onthophagus haematopus
Onthophagus hirtus
Onthophagus illyricus
Onthophagus incensus
Onthophagus interstitialis
Onthophagus japonicus
Onthophagus joannae
Onthophagus laminatus
Onthophagus laticollis
Onthophagus latigena
Onthophagus lemur
Onthophagus lenzi
Onthophagus longimanus
Onthophagus maki
Onthophagus maniti
Onthophagus marginalis
Onthophagus medius
Onthophagus melitaeus
Onthophagus merdarius
Onthophagus mije
Onthophagus minutus
Onthophagus mulgravei
Onthophagus muticus
Onthophagus nebulosus
Onthophagus nitidior
Onthophagus nutans
Onthophagus obscurior
Onthophagus ochreatus
Onthophagus ochromerus
Onthophagus ochropygus
Onthophagus opacicollis
Onthophagus orientalis
Onthophagus ovatus
Onthophagus pedator
Onthophagus phanaeides
Onthophagus piyawati
Onthophagus pullus
Onthophagus punctatus
Onthophagus quadripustulatus
Onthophagus rhinolophus
Onthophagus rorarius
Onthophagus rubicundulus
Onthophagus ruficapillus
Onthophagus rugicollis
Onthophagus schwaneri
Onthophagus semiareus
Onthophagus similis
Onthophagus sinicus
Onthophagus solivagus
Onthophagus stylocerus
Onthophagus taurinus
Onthophagus taurus
Onthophagus tragus
Onthophagus tricornis
Onthophagus vacca
Onthophagus variolaris
Onthophagus verticicornis
Onthophagus viduus
Onthophagus vinctus
Onthophagus vulpes
Onthophagus yukae
66 additional siblings truncated for brevity.
See the
resource file
for a full list.
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Onthophagini
Onthophagus
Onthophagus pennsylvanicus
(this page)
Onthophagus angularis
Onthophagus asper
Onthophagus aspericollis
Onthophagus asperipennis
Onthophagus aspernatus
Onthophagus asperrimus
Onthophagus asperulus
Onthophagus astigma
Onthophagus athiensis
Onthophagus atricolor
Onthophagus atridorsis
Onthophagus atroaereus
Onthophagus atrofasciatus
Onthophagus atronitidus
Onthophagus atrosericeus
Onthophagus aureofuscus
Onthophagus aureopilosus
Onthophagus auriculatus
Onthophagus aurifex
Onthophagus australis
Onthophagus axillaris
Onthophagus aztecus
Onthophagus comperei
Onthophagus compositus
Onthophagus costatus
Onthophagus costiger
Onthophagus creber
Onthophagus cribellum
Onthophagus cribratus
Onthophagus cribripennis
Onthophagus curvispina
Onthophagus cyaneiceps
Onthophagus cyanellus
Onthophagus cyaneoniger
Onthophagus cyanochlorus
Onthophagus cyclographus
Onthophagus cynomysi
Onthophagus cyobioides
Onthophagus dama
Onthophagus dandalu
Onthophagus dapcauensis
Onthophagus dapitanensis
Onthophagus decedens
Onthophagus digitifer
Onthophagus discrepans
Onthophagus endroedii
Onthophagus epilamprus
Onthophagus frontalis
Onthophagus gaillardi
Onthophagus gajo
Onthophagus galeatus
Onthophagus ganalensis
Onthophagus gandju
Onthophagus gangeticus
Onthophagus gangulu
Onthophagus gazellinus
Onthophagus geelongensis
Onthophagus gemellatus
Onthophagus gemmatus
Onthophagus gestroi
Onthophagus gibber
Onthophagus gibberosus
Onthophagus gibbidorsis
Onthophagus gibbus
Onthophagus gidju
Onthophagus giraffa
Onthophagus glabratus
Onthophagus glasunovi
Onthophagus gonopygus
Onthophagus gracilipes
Onthophagus humboldti
Onthophagus illyricus
Onthophagus indosinicus
Onthophagus indutus
Onthophagus inelegans
Onthophagus ineptus
Onthophagus inermiceps
Onthophagus inermicollis
Onthophagus inflaticollis
Onthophagus infucatus
Onthophagus inops
Onthophagus laborans
Onthophagus mocquerysi
Onthophagus modestus
Onthophagus noctis
Onthophagus pilauco
Onthophagus pluton
Onthophagus quadripustulatus
Onthophagus queenslandicus
Onthophagus quinquedens
Onthophagus quinquetuberculatus
Onthophagus ragazzii
Onthophagus rotundicollis
Onthophagus schultzei
Onthophagus sideki
Onthophagus simplex
Onthophagus simpliciceps
Onthophagus simulator
Onthophagus sinuosicollis
Onthophagus sinuosus
2369 additional siblings truncated for brevity.
See the
resource file
for a full list.
