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Heteronychus atratus Klug 1855

Scientific Names

Preferred Names

Heteronychus atratus Klug 1855
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus atratus
Species recognized by Australia Species List, Tanzania Species List, and Indian Ocean Species List
Heteronychus atratus Klug 1855
Species recognized by EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, and GBIF classification
Heteronychus dissidens
Species recognized by Zimbabwe Species List and Indian Ocean Species List
Heteronychus (Heteronychus) atratus Klug 1855
Species recognized by Catalogue of Life 26 March 2024
Heteronychus atratus
Species recognized by wikidata_hierarchy tar gz
Heteronychus atratus
Recognized by BHL data coverage
Heteronychus atratus Klug 1855
Species recognized by GBIF data coverage, Dynamic Hierarchy Version 2.2.3, and EOL Dynamic Hierarchy Version 2.2.4

Alternative Names

Heteronychus bucobensis Kolbe 1914
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus densatifrons Fairmaire 1893
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus dissidens Péringuey 1901
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus indotatus Péringuey 1901
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus obtusifrons Fairmaire 1893
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus transvaalicus Kolbe 1900
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus viator Péringuey 1901
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Heteronychus bucobensis Kolbe 1914
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus densatifrons Fairmaire 1893
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus dissidens Péringuey 1901
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus indotatus Péringuey 1901
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus obtusifrons Fairmaire 1893
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus transvaalicus Kolbe 1900
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus viator Péringuey 1901
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Heteronychus bucobensis Kolbe 1914
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus bucobensis Kolbe 1914
Synonym according to Catalogue of Life 26 March 2024
Heteronychus densatifrons Fairmaire 1893
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus densatifrons Fairmaire 1893
Synonym according to Catalogue of Life 26 March 2024
Heteronychus dissidens Péringuey 1901
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus dissidens Péringuey 1901
Synonym according to Catalogue of Life 26 March 2024
Heteronychus indotatus Péringuey 1901
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus indotatus Péringuey 1901
Synonym according to Catalogue of Life 26 March 2024
Heteronychus obtusifrons Fairmaire 1893
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus obtusifrons Fairmaire 1893
Synonym according to Catalogue of Life 26 March 2024
Heteronychus transvaalicus Kolbe 1900
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus transvaalicus Kolbe 1900
Synonym according to Catalogue of Life 26 March 2024
Heteronychus viator Péringuey 1901
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Heteronychus viator Péringuey 1901
Synonym according to Catalogue of Life 26 March 2024

Common Names

There are no common names associated with this taxon.

Curated hierarchies for Heteronychus atratus Klug 1855

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Heteronychus
Heteronychus atratus Klug 1855 (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Pentodontini Mulsant 1842
Pentodontina Mulsant 1842
Heteronychus Dejean 1833
Heteronychus subgen. Heteronychus Dejean 1833
Heteronychus (Heteronychus) atratus Klug 1855 (this page)
Heteronychus (Heteronychus) abyssinicus Jack 1923
Heteronychus (Heteronychus) amplipennis Benderitter 1915
Heteronychus (Heteronychus) amplus Kolbe 1900
Heteronychus (Heteronychus) andersoni Jack 1923
Heteronychus (Heteronychus) annulatus Bates 1891
Heteronychus (Heteronychus) approximans Kolbe 1900
Heteronychus (Heteronychus) arator (Fabricius 1775)
Heteronychus (Heteronychus) ascanius Kolbe 1900
Heteronychus (Heteronychus) basilewskyi Endrödi 1962
Heteronychus (Heteronychus) bituberculatus Kolbe 1900
Heteronychus (Heteronychus) brittoni Ferreira 1965
Heteronychus (Heteronychus) carvalhoi Ferreira 1970
Heteronychus (Heteronychus) citernii Paoli 1934
Heteronychus (Heteronychus) congoensis Kolbe 1900
Heteronychus (Heteronychus) consimilis Kolbe 1900
Heteronychus (Heteronychus) cordatus Endrödi 1974
Heteronychus (Heteronychus) costatus Lansberge 1886
Heteronychus (Heteronychus) cricetus (Hausmann 1807)
Heteronychus (Heteronychus) curtulus Fairmaire 1887
Heteronychus (Heteronychus) desaegeri Burgeon 1947
Heteronychus (Heteronychus) digitiformis Zhang 1983
Heteronychus (Heteronychus) fossor Reiche 1850
Heteronychus (Heteronychus) howdeni Yamaya 2016
Heteronychus (Heteronychus) impudens Jack 1923
Heteronychus (Heteronychus) infans Kolbe 1897
Heteronychus (Heteronychus) inoportunus Ferreira 1966
Heteronychus (Heteronychus) insignificus Jack 1923
Heteronychus (Heteronychus) intermedius Zhang 1983
Heteronychus (Heteronychus) jacki Arrow 1936
Heteronychus (Heteronychus) licas (Klug 1835)
Heteronychus (Heteronychus) lioderes Redtenbacher 1868
Heteronychus (Heteronychus) lusingae Endrödi 1974
Heteronychus (Heteronychus) minimus Dupuis 2012
Heteronychus (Heteronychus) mollis Endrödi 1961
Heteronychus (Heteronychus) monodi Paulian 1954
Heteronychus (Heteronychus) mosambicus Péringuey 1908
Heteronychus (Heteronychus) muticus Benderitter 1915
Heteronychus (Heteronychus) paolii Arrow 1936
Heteronychus (Heteronychus) parumpunctatus Burmeister 1847
Heteronychus (Heteronychus) parvus Burmeister 1847
Heteronychus (Heteronychus) pauperatus Péringuey 1901
Heteronychus (Heteronychus) plebejus (Klug 1834)
Heteronychus (Heteronychus) puerilis Kolbe 1900
Heteronychus (Heteronychus) puncticollis Jack 1923
Heteronychus (Heteronychus) punctolineatus Fairmaire 1893
Heteronychus (Heteronychus) pygidialis Kolbe 1900
Heteronychus (Heteronychus) rusticus (Klug 1834)
Heteronychus (Heteronychus) sabackyi Endrödi 1975
Heteronychus (Heteronychus) sacchari Arrow 1908
Heteronychus (Heteronychus) similis Dechambre 1992
Heteronychus (Heteronychus) simulans Jack 1923
Heteronychus (Heteronychus) sublaevis (Fairmaire 1891)
Heteronychus (Heteronychus) tenuestriatus Fairmaire 1893
Heteronychus (Heteronychus) tesari Endrödi 1968
Heteronychus (Heteronychus) tristis Boheman 1857
Heteronychus (Heteronychus) vixstriatus Jack 1923
Heteronychus (Heteronychus) wittei Janssens 1942

Dynamic Hierarchy Version 2.2.3

Heteronychus atratus Klug 1855 (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Heteronychus
Heteronychus atratus Klug 1855 (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942

EOL Dynamic Hierarchy Version 2.2.4

Heteronychus atratus Klug 1855 (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Heteronychus Burmeister 1847
Heteronychus atratus Klug 1855 (this page)
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus egenis
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus minutus Burmeister 1847
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tenvestriatus
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Pentodontini
Heteronychus
Heteronychus atratus (this page)
Heteronychidius
Heteronychus abyssinicus
Heteronychus amplipennis
Heteronychus amploides
Heteronychus amplus
Heteronychus andersoni
Heteronychus annulatus
Heteronychus approximans
Heteronychus arator
Heteronychus ascanius
Heteronychus basilewskyi
Heteronychus bituberculatus
Heteronychus brittoni
Heteronychus carvalhoi
Heteronychus citernii
Heteronychus congoensis
Heteronychus consimilis
Heteronychus cordatus
Heteronychus costatus
Heteronychus cricetus
Heteronychus curtulus
Heteronychus desaegeri
Heteronychus digitiformis
Heteronychus fossor
Heteronychus howdeni
Heteronychus humilis
Heteronychus impudens
Heteronychus infans
Heteronychus inoportunus
Heteronychus insignificus
Heteronychus intermedius
Heteronychus jacki
Heteronychus krombeini
Heteronychus licas
Heteronychus lioderes
Heteronychus lusingae
Heteronychus minimus
Heteronychus minutus
Heteronychus mollis
Heteronychus monodi
Heteronychus mosambicus
Heteronychus muticus
Heteronychus paolii
Heteronychus parumpunctatus
Heteronychus parvus
Heteronychus pauperatus
Heteronychus plebejus
Heteronychus puerilis
Heteronychus puncticollis
Heteronychus punctolineatus
Heteronychus pygidialis
Heteronychus rusticus
Heteronychus sabackyi
Heteronychus sacchari
Heteronychus similis
Heteronychus simulans
Heteronychus sublaevis
Heteronychus tenuestriatus
Heteronychus tesari
Heteronychus tristis
Heteronychus vixstriatus
Heteronychus wittei
Seidlitz
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