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Masked Chafers
Cyclocephala
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names
Scientific Names
Preferred Names
Cyclocephala
Genus recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Cyclocephala
Genus recognized by
wikipedia русскую Википедию
,
wikipedia NO
,
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
,
Australia Species List
,
Belize Species List
,
Bolivia Species List
,
Brazil Species List
,
Canada Species List
,
Colombia Species List
,
Costa Rica Species List
,
Dominican Republic Species List
,
Ecuador Species List
,
El Salvador Species List
,
Guatemala Species List
,
Guyana Species List
,
Nicaragua Species List
,
Panama Species List
,
Peru Species List
,
Paraguay Species List
,
Puerto Rico Species List
,
Suriname Species List
,
Venezuela Species List
,
Argentina Species List
,
Mexico Species List
,
North Atlantic Species List
,
North Pacific Species List
,
NCBI
,
wikipedia emerging languages
,
United States Species List
,
wikidata_hierarchy tar gz
,
Paleobiology Database
,
wikipedia FR
,
wikipedia EN
,
Wikidata Common Names
,
Wikimedia Commons
,
Barcode of Life Data Systems
,
Global Biotic Interactions
,
Global Biotic Interactions
,
iNaturalist data coverage
,
Wikipedia (inferred records)
,
Dynamic Hierarchy Version 2.2.3
,
GGBN data coverage
, and
EOL Dynamic Hierarchy Version 2.2.4
Cyclocephala
Recognized by
IABIN
Cyclocephala Dejean 1821
Genus recognized by
GBIF classification
,
Catalogue of Life 26 March 2024
,
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
, and
GBIF data coverage
Alternative Names
Albridarollia ocellata
Burmeister 1847
Doubtful according to
GBIF classification
Cyclocephala frontalis
Burmeister 1847
Heterotypic synonym according to
GBIF classification
Cyclocephala pereirai
Endrödi 1969
Heterotypic synonym according to
GBIF classification
Cyclocephala pubescens
Burmeister 1847
Proparte synonym according to
GBIF classification
and
GBIF classification
Cyclocephala quisqueya
Joly T.
Doubtful according to
GBIF classification
(Possible variant of Cyclocephala quisqueya Joly, 1998)
Cyclocephala signatoides
Chapin 1932
Heterotypic synonym according to
GBIF classification
Cyclocephala spangleri
Joly T.
Doubtful according to
GBIF classification
(Possible variant of Cyclocephala spangleri Joly, 2000)
Cyclocephala vittoscutellaris
Endrödi 1966
Heterotypic synonym according to
GBIF classification
Melolontha elongata
Olivier 1789
Proparte synonym according to
GBIF classification
Melolontha signata
Fabricius 1781
Heterotypic synonym according to
GBIF classification
Melolontha signata
Olivier 1789
Heterotypic synonym according to
GBIF classification
Aclinidia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Aclinidia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Albridarollia Bolivar, Jimenez & Martinez 1963
Synonym according to
Catalogue of Life 26 March 2024
Albridarollia Bolívar, Pieltain, Jiménez-Asúa & Martínez 1963
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Aspidolea subgen. Aspidotites Höhne 1922
Synonym according to
Catalogue of Life 26 March 2024
Aspidolella Prell 1936
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Aspidolella Prell 1936
Synonym according to
Catalogue of Life 26 March 2024
Aspidolites Höhne 1922
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Cyclocephala
Harvest ancestor according to
EOL authors
,
GBIF national node type records France
,
GBIF national node type records Sweden
,
NMNH type specimens
,
iNaturalist
,
Harvard Museum of Comparative Zoology
,
Simpson et al 2020
,
TreatmentBank
,
Global Biotic Interactions
,
Global Biotic Interactions
, and
Flickr Group
Diapatalia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Diapatalia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Dichromina Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Dichromina Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Graphalia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Graphalia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Halotosia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Halotosia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Homochromina Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Homochromina Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Isocoryna Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Isocoryna Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Mononidia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Mononidia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Ochrosidia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Ochrosidia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Paraclinidia Martinez 1965
Synonym according to
Catalogue of Life 26 March 2024
Paraclinidia Martínez 1965
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Plagiosalia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Plagiosalia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Spilosota Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Spilosota Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Stigmalia Casey 1915
Invalid according to
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Stigmalia Casey 1915
Synonym according to
Catalogue of Life 26 March 2024
Surutoides Endrödi 1981
Synonym according to
Catalogue of Life 26 March 2024
Common Names
English
Masked Chafers
preferred
Recognized by
Wikidata Common Names
Arabic
جعال
preferred
Recognized by
Wikidata Common Names
Arz
جعال
preferred
Recognized by
Wikidata Common Names
Curated hierarchies for Cyclocephala
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Cyclocephala
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala acuta Arrow 1902
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1829
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala aulustjaorum Hielkema 2017
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala carinatipennis Martinez & Morón 1984
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala englemani (Ratcliffe 1977)
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavipennis Arrow 1914
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Maly 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala italoi Dupuis 1999
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longitarsis Dechambre 1999
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculata Burmeister 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melanopoda Ratcliffe 2008
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigra (Endrödi 1979)
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nigropicta Dechambre & Endrodi 1983
Cyclocephala niguasa Dechambre & Endrodi 1984
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala pan Ratcliffe 1992
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala parallela (Casey 1915)
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala perconfusa Dechambre 1992
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala pulchra Dechambre 1999
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1829
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1891
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santaritae Ratcliffe 1992
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala signatoides Höhne 1923
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Bréthes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1846
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala virgo Dechambre 1999
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
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Chalcasthenes
Chalcocrates
Chalepides
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Cryptodus
Dalgopus
Dasygnathus
Denhezia
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Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the
resource file
for a full list.
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Cyclocephalini Castelnau 1840
Cyclocephala Dejean 1821
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arhuacana Neita-Moreno 2021
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1828
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brevis Höhne 1923
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala cristula Ratcliffe & Cave 2020
Cyclocephala cucullata Ratcliffe & Cave 2020
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dicula Dupuis 2021
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Malý 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala hylaea Ratcliffe & Paz 2022
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala marquezi Delgado & Mora-Aguilar 2020
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala mixteca Cruz-López & Osorio-Alcalá 2022
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octomaculata Dupuis 2021
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala parallela (Casey 1915)
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1828
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1892
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santandereana Neita-Moreno 2021
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala sculpta Dupuis 2021
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Brèthes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala ukuku Paz & Ratcliffe 2022
Cyclocephala ukukumateoi Paz & Ratcliffe 2022
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1847
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala vulcanorum Ratcliffe 2021
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Acrobolbia Ohaus 1912
Ancognatha Erichson 1847
Arriguttia Martinez 1960
Aspidolea Bates 1888
Augoderia Burmeister 1847
Chalepides Casey 1915
Dyscinetus Harold 1869
Erioscelis Burmeister 1847
Harposceles Burmeister 1847
Mimeoma Casey 1915
Parapucaya Prell 1934
Pucaya Ohaus 1910
Ruteloryctes Arrow 1908
Stenocrates Burmeister 1847
Surutu Martinez 1955
Dynamic Hierarchy Version 2.2.3
Cyclocephala
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1829
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala carinatipennis Martinez & Morón 1984
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala englemani (Ratcliffe 1977)
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Maly 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala parallela (Casey 1915)
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala pulchra Dechambre 1999
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1829
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1891
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santaritae Ratcliffe 1992
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Bréthes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1846
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala virgo Dechambre 1999
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Cyclocephala
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala acuta Arrow 1902
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1829
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala aulustjaorum Hielkema 2017
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala carinatipennis Martinez & Morón 1984
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala englemani (Ratcliffe 1977)
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavipennis Arrow 1914
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Maly 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala italoi Dupuis 1999
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longitarsis Dechambre 1999
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculata Burmeister 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melanopoda Ratcliffe 2008
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigra (Endrödi 1979)
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nigropicta Dechambre & Endrodi 1983
Cyclocephala niguasa Dechambre & Endrodi 1984
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala pan Ratcliffe 1992
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala parallela (Casey 1915)
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala perconfusa Dechambre 1992
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala pulchra Dechambre 1999
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1829
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1891
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santaritae Ratcliffe 1992
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala signatoides Höhne 1923
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Brethes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1846
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala virgo Dechambre 1999
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Aceratus
Adoryphorus
Aegopsis
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calcitoryctes
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
118 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy Version 2.2.4
Cyclocephala
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1829
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala carinatipennis Martinez & Morón 1984
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala englemani (Ratcliffe 1977)
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Maly 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala parallela (Casey 1915)
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala pulchra Dechambre 1999
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1829
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1891
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santaritae Ratcliffe 1992
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Bréthes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1846
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala virgo Dechambre 1999
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the
resource file
for a full list.
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Cyclocephala Dejean 1821
(this page)
Cyclocephala abrelata Ratcliffe & Cave 2002
Cyclocephala acoma Ratcliffe 2008
Cyclocephala acuta Arrow 1902
Cyclocephala aequatoria Endrödi 1963
Cyclocephala affinis Endrödi 1966
Cyclocephala alazonia Ratcliffe 2003
Cyclocephala alexi Ratcliffe & Delgado 1990
Cyclocephala almitana Dechambre 1992
Cyclocephala altamontana Dechambre 1999
Cyclocephala alutacea Höhne 1923
Cyclocephala alvarengai Dechambre 1980
Cyclocephala amazona (Linnaeus 1767)
Cyclocephala amblyopsis Bates 1888
Cyclocephala ampliata Bates 1888
Cyclocephala amplitarsis Ratcliffe 1992
Cyclocephala anibali Joly 2009
Cyclocephala antoinei Dupuis 2018
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrodi 1966
Cyclocephala arnaudi Dechambre 1980
Cyclocephala arrowiana Martinez 1967
Cyclocephala atricapilla Mannerheim 1829
Cyclocephala atriceps (Casey 1915)
Cyclocephala atricolor Chapin 1932
Cyclocephala atripes Bates 1888
Cyclocephala aulustjaorum Hielkema 2017
Cyclocephala barrerai Martinez 1969
Cyclocephala batesi Delgado-Castillo & Castaneda 1994
Cyclocephala bella Endrödi 1969
Cyclocephala berti Delgado-Castillo 1992
Cyclocephala bicolor Castelnau 1840
Cyclocephala bicolorata Endrödi 1964
Cyclocephala bimaculata Dechambre 1999
Cyclocephala binotata Dechambre 1999
Cyclocephala bleuzeni Dechambre 1995
Cyclocephala boliviana Dechambre 1992
Cyclocephala bollei Dechambre & Endrodi 1984
Cyclocephala borburatae Endrödi 1980
Cyclocephala borealis Arrow 1911
Cyclocephala boucheri Dechambre 1997
Cyclocephala boulardi Dechambre 1979
Cyclocephala brasiliana Endrödi 1966
Cyclocephala brevipennis Endrödi 1985
Cyclocephala brittoni Endrödi 1964
Cyclocephala burmeisteri Endrödi 1964
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala camachicola Ohaus 1910
Cyclocephala capitata Höhne 1923
Cyclocephala carbonaria Arrow 1911
Cyclocephala cardini Chapin 1935
Cyclocephala carinatipennis Martinez & Morón 1984
Cyclocephala cartwighti Endrödi 1964
Cyclocephala cartwrighti Endrödi 1964
Cyclocephala casanova Ratcliffe & Cave 2009
Cyclocephala castanea (Olivier 1789)
Cyclocephala castaniella Bates 1888
Cyclocephala caussaneli Dechambre 1999
Cyclocephala cearae Höhne 1923
Cyclocephala celata Dechambre 1980
Cyclocephala cerea Burmeister 1847
Cyclocephala chalumeani Martínez
Cyclocephala chalumeaui Martinez 1978
Cyclocephala chera Ratcliffe 2008
Cyclocephala chiquitita Ratcliffe 2008
Cyclocephala colasi Endrödi 1964
Cyclocephala collaris Burmeister 1847
Cyclocephala comata Bates 1888
Cyclocephala compacta Ratcliffe 2008
Cyclocephala complanata Burmeister 1847
Cyclocephala concolor Burmeister 1847
Cyclocephala confusa Endrödi 1966
Cyclocephala conspicua Sharp 1877
Cyclocephala contraria Kirsch 1873
Cyclocephala coriacea Dechambre 1992
Cyclocephala couturieri Dechambre 1999
Cyclocephala crassa Endrödi 1967
Cyclocephala crepuscularis Martinez 1954
Cyclocephala cribrata Burmeister 1847
Cyclocephala curta Bates 1888
Cyclocephala dalensi Ponchel 2009
Cyclocephala deceptor (Casey 1915)
Cyclocephala dechambrei Dupuis 2018
Cyclocephala decorella Endrödi 1966
Cyclocephala defecta Endrödi 1970
Cyclocephala deltoides Ratcliffe 1992
Cyclocephala dichroa Dechambre 1992
Cyclocephala dilatata (Prell 1934)
Cyclocephala diluta Erichson 1847
Cyclocephala discicollis Arrow 1902
Cyclocephala discolor (Herbst 1790)
Cyclocephala dispar (Herbst 1790)
Cyclocephala distincta Burmeister 1847
Cyclocephala divaricata Joly 2005
Cyclocephala dolichotarsa Ratcliffe & Cave 2008
Cyclocephala dominicana Endrödi 1985
Cyclocephala duodecimpunctata Endrödi 1966
Cyclocephala dupuisi Ratcliffe 2014
Cyclocephala durantonorum Dechambre 1999
Cyclocephala dyscinetoides Dechambre 1999
Cyclocephala emarginata Endrödi 1966
Cyclocephala endrodii Martinez 1965
Cyclocephala endroedyyoungai Endrödi 1964
Cyclocephala englemani (Ratcliffe 1977)
Cyclocephala enigma Ratcliffe 2003
Cyclocephala epistomalis Bates 1888
Cyclocephala ergastuli Dechambre 1997
Cyclocephala erotylina Arrow 1914
Cyclocephala everardoi Grossi, Santos & Almeida 2016
Cyclocephala fankhaeneli Endrödi 1964
Cyclocephala fasciolata Bates 1888
Cyclocephala figurata Burmeister 1847
Cyclocephala flavipennis Arrow 1914
Cyclocephala flavoscutellaris Höhne 1923
Cyclocephala flora Arrow 1911
Cyclocephala forcipulata Howden & Endrodi 1966
Cyclocephala forsteri Endrödi 1963
Cyclocephala freudei Endrödi 1963
Cyclocephala freyi Endrödi 1964
Cyclocephala frontalis Chevrolat 1844
Cyclocephala fulgurata Burmeister 1847
Cyclocephala fulvipennis Burmeister 1847
Cyclocephala gabaldoni Martinez & Martinez 1981
Cyclocephala genieri Joly 2010
Cyclocephala gigantea Dupuis 1999
Cyclocephala goetzi Endrödi 1966
Cyclocephala gravis Bates 1888
Cyclocephala gregaria Heyne & Taschenberg 1908
Cyclocephala guaguarum Dechambre & Endrodi 1984
Cyclocephala guianae Endrödi 1969
Cyclocephala guttata Bates 1888
Cyclocephala guycolasi Dechambre 1992
Cyclocephala halffteriana Martinez 1968
Cyclocephala hardyi Endrödi 1975
Cyclocephala hartmannorum Maly 2006
Cyclocephala helavai Endrödi 1975
Cyclocephala herteli Endrödi 1964
Cyclocephala hiekei Endrödi 1964
Cyclocephala hirsuta Höhne 1923
Cyclocephala hirta Le Conte 1861
Cyclocephala histrionica Burmeister 1847
Cyclocephala holmbergi Martinez 1968
Cyclocephala howdenannae Endrödi 1975
Cyclocephala huamilule Romero & Morón 2017
Cyclocephala huesingi Endrödi 1964
Cyclocephala iani Ratcliffe 1992
Cyclocephala immaculata (Olivier 1789)
Cyclocephala inca Endrödi 1966
Cyclocephala insulicola Arrow 1937
Cyclocephala isabellina Höhne 1923
Cyclocephala isthmiensis Ratcliffe 1992
Cyclocephala italoi Dupuis 1999
Cyclocephala jalapensis Casey 1915
Cyclocephala jauffreti Dechambre 1979
Cyclocephala kahanoffae Martinez 1975
Cyclocephala kaszabi Endrödi 1964
Cyclocephala kechua (Martinez 1957)
Cyclocephala krombeini Endrödi 1979
Cyclocephala kuntzeniana Höhne 1923
Cyclocephala labidion Ratcliffe 2003
Cyclocephala lachaumei Dechambre 1992
Cyclocephala laevis Arrow 1937
Cyclocephala lamarcki Dechambre 1999
Cyclocephala laminata Burmeister 1847
Cyclocephala larssoni Endrödi 1964
Cyclocephala latericia Höhne 1923
Cyclocephala latipennis Arrow 1911
Cyclocephala latreillei Dechambre 1999
Cyclocephala lecourti Dechambre 1992
Cyclocephala letiranti Young 1992
Cyclocephala lichyi Dechambre 1980
Cyclocephala ligyrina Bates 1888
Cyclocephala lineata Dupuis 2008
Cyclocephala lineigera Höhne 1923
Cyclocephala liomorpha Arrow 1911
Cyclocephala literata Burmeister 1847
Cyclocephala lizeri Martinez 1964
Cyclocephala longa Endrödi 1963
Cyclocephala longicollis Burmeister 1847
Cyclocephala longimana Dechambre 1980
Cyclocephala longitarsis Dechambre 1999
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala lutea Endrödi 1966
Cyclocephala machadoi Grossi, Santos & Almeida 2016
Cyclocephala macrophylla Erichson 1847
Cyclocephala maculata Burmeister 1847
Cyclocephala maculiventris Höhne 1923
Cyclocephala mafaffa Burmeister 1847
Cyclocephala magdalenae Young & Le Tirant 2005
Cyclocephala malleri Martinez 1968
Cyclocephala malyi Dupuis 2014
Cyclocephala mannheimsi Endrödi 1964
Cyclocephala marginalis Kirsch 1871
Cyclocephala marginicollis Arrow 1902
Cyclocephala marianista Dechambre & Endrodi 1984
Cyclocephala marqueti Dechambre 1997
Cyclocephala martinezi Endrödi 1964
Cyclocephala marylizae Ratcliffe 2003
Cyclocephala mathani Dechambre 1982
Cyclocephala mechae Martinez 1978
Cyclocephala mecynotarsis Höhne 1923
Cyclocephala megalophylla Endrödi 1966
Cyclocephala melane Bates 1888
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala melanopoda Ratcliffe 2008
Cyclocephala melolonthida Ratcliffe & Cave 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado 2012
Cyclocephala metrica Steinheil 1872
Cyclocephala miamiensis Howden & Endrodi 1966
Cyclocephala minuchae Joly 2003
Cyclocephala minuta Burmeister 1847
Cyclocephala modesta Burmeister 1847
Cyclocephala molesta Endrödi 1969
Cyclocephala monacha Ratcliffe 2008
Cyclocephala monzoni Ratcliffe & Cave 2009
Cyclocephala moreti Dechambre 1992
Cyclocephala morphoidina Prell 1937
Cyclocephala multiplex Casey 1915
Cyclocephala munda Kirsch 1871
Cyclocephala mustacha Ratcliffe 2003
Cyclocephala mutata Harold 1869
Cyclocephala nana Dechambre 1999
Cyclocephala nicolasi Dupuis 2018
Cyclocephala nigerrima Bates 1888
Cyclocephala nigra (Endrödi 1979)
Cyclocephala nigritarsis Ratcliffe 1992
Cyclocephala nigrobasalis Höhne 1923
Cyclocephala nigropicta Dechambre & Endrodi 1983
Cyclocephala niguasa Dechambre & Endrodi 1984
Cyclocephala nike Ratcliffe 1992
Cyclocephala nodanotherwon Ratcliffe 1992
Cyclocephala notata (Illiger 1806)
Cyclocephala obscura Endrödi 1966
Cyclocephala occipitalis Fairmaire 1892
Cyclocephala ocellata Burmeister 1847
Cyclocephala ochracea Prell 1937
Cyclocephala octopunctata Burmeister 1847
Cyclocephala ohausiana Höhne 1923
Cyclocephala olivieri Arrow 1911
Cyclocephala ovulum Bates 1888
Cyclocephala pan Ratcliffe 1992
Cyclocephala panthera Dechambre 1979
Cyclocephala paraflora Martinez 1978
Cyclocephala paraguayensis Arrow 1903
Cyclocephala pardolocarnoi Dechambre 1995
Cyclocephala pasadenae (Casey 1915)
Cyclocephala perconfusa Dechambre 1992
Cyclocephala pereirai (Martinez 1960)
Cyclocephala perforata Arrow 1913
Cyclocephala perplexa Ratcliffe 2008
Cyclocephala peruana Endrödi 1966
Cyclocephala pichinchana Dechambre 1992
Cyclocephala picipes (Olivier 1789)
Cyclocephala picta Burmeister 1847
Cyclocephala pilosa Dupuis 2006
Cyclocephala pokornyi Dupuis 2014
Cyclocephala pompanoni Dechambre 1979
Cyclocephala poncheli Dechambre & Duranton 2005
Cyclocephala porioni Dechambre 1979
Cyclocephala prelli Endrödi 1967
Cyclocephala prolongata Arrow 1902
Cyclocephala proxima Dechambre 1997
Cyclocephala pseudoconfusa Ratcliffe 1992
Cyclocephala pseudomelanocephala Dupuis 1996
Cyclocephala puberula Le Conte 1863
Cyclocephala pugnax Arrow 1914
Cyclocephala pulchra Dechambre 1999
Cyclocephala puncticollis Endrödi 1966
Cyclocephala putrida Burmeister 1847
Cyclocephala pygidialis Joly 2000
Cyclocephala pygidiata Dupuis 1999
Cyclocephala quadripunctata Höhne 1923
Cyclocephala quatuordecimpunctata Mannerheim 1829
Cyclocephala quercina Burmeister 1847
Cyclocephala quisqueya Joly 1998
Cyclocephala rangelana Chapin 1935
Cyclocephala ratcliffei Endrödi 1977
Cyclocephala recta Dupuis 2008
Cyclocephala regularis Casey 1915
Cyclocephala robusta Le Conte 1863
Cyclocephala rogerpauli Moore, Branham & Cave 2018
Cyclocephala rogezi Dechambre 1992
Cyclocephala rondoniana Ratcliffe 1992
Cyclocephala rorulenta Höhne 1923
Cyclocephala rotundipenis Dupuis 2009
Cyclocephala rubescens Bates 1891
Cyclocephala rufa Endrödi 1967
Cyclocephala rufescens Endrödi 1967
Cyclocephala ruficollis Burmeister 1847
Cyclocephala rufonigra Demay 1838
Cyclocephala rufovaria Arrow 1911
Cyclocephala rustica (Olivier 1789)
Cyclocephala saltini Ratcliffe 2008
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala santaritae Ratcliffe 1992
Cyclocephala sarahae Ratcliffe 1992
Cyclocephala sardadebiae Dechambre & Duranton 2005
Cyclocephala sarpedon Ratcliffe 1992
Cyclocephala scarabaeina (Gyllenhal 1817)
Cyclocephala schmitzorum Ratcliffe 1992
Cyclocephala seditiosa Le Conte 1863
Cyclocephala setosa Burmeister 1847
Cyclocephala sexpunctata Castelnau 1840
Cyclocephala signaticollis Burmeister 1847
Cyclocephala signatoides Höhne 1923
Cyclocephala similis Dechambre 1980
Cyclocephala simillima Dechambre 1999
Cyclocephala simulatrix Höhne 1923
Cyclocephala sinaloae Howden & Endrodi 1966
Cyclocephala sinuosa Höhne 1923
Cyclocephala spangleri Joly 2000
Cyclocephala sparsa Arrow 1902
Cyclocephala spilopyga Erichson 1847
Cyclocephala stictica Burmeister 1847
Cyclocephala stockwelli Ratcliffe 2003
Cyclocephala striata Endrödi 1963
Cyclocephala subsignata Burmeister 1847
Cyclocephala supernana Dechambre 1999
Cyclocephala suturalis Ohaus 1911
Cyclocephala sylviae Dechambre 1995
Cyclocephala testacea Burmeister 1847
Cyclocephala tetrica Burmeister 1847
Cyclocephala tidula Dechambre 1999
Cyclocephala toulgoeti Dechambre 1992
Cyclocephala tridentata (Fabricius 1801)
Cyclocephala tronchonii Martinez 1975
Cyclocephala tucumana Bréthes 1904
Cyclocephala tutilina Burmeister 1847
Cyclocephala tylifera Höhne 1923
Cyclocephala unamas Ratcliffe 2003
Cyclocephala undata (Olivier 1789)
Cyclocephala unidentata Endrödi 1980
Cyclocephala variabilis Burmeister 1847
Cyclocephala varians Burmeister 1847
Cyclocephala variipenis Dechambre 1999
Cyclocephala variolosa Burmeister 1847
Cyclocephala verticalis Burmeister 1847
Cyclocephala vestita Höhne 1923
Cyclocephala vidanoi Dechambre 1992
Cyclocephala villosa Blanchard 1846
Cyclocephala vincentiae Arrow 1900
Cyclocephala vinosa Arrow 1937
Cyclocephala virgo Dechambre 1999
Cyclocephala viridis Dechambre 1982
Cyclocephala vittoscutellaris Prell 1937
Cyclocephala wandae Hardy 1974
Cyclocephala weidneri Endrödi 1964
Cyclocephala williami Ratcliffe 1992
Cyclocephala zischkai Martinez 1965
Cyclocephala zodion Ratcliffe 1992
Cyclocephala zurstrasseni Endrödi 1964
Aceratus Prell 1936
Acrobolbia Ohaus 1912
Aegopsis Burmeister 1847
Agaocephala Le Peletier & Serville 1828
Allophileurinus Dupuis & Dechambre 2001
Amblyodus Westwood 1878
Amblyoproctus Kolbe 1910
Ancognatha Erichson 1847
Aphonodelus Kolbe 1905
Archophileurus Kolbe 1910
Aspidolea Bates 1888
Augosoma Burmeister 1841
Blabephorus Fairmaire 1898
Bothynus Hope 1837
Calicnemis Laporte de Castelnau 1832
Carneiola Endrödi 1974
Carneoryctes Özdikmen 2009
Cavonus Sharp 1875
Ceratophileurus Ohaus 1911
Ceratoryctoderus Arrow 1908
Chalcasthenes Arrow 1937
Chalcosoma Hope 1837
Cheiroplatys Kirby 1837
Chiliphileurus Endrodi 1977
Cnemidophileurus Kolbe 1910
Coenoryctoderus Prell 1933
Colacus Ohaus 1910
Collagenus Ratcliffe & Hardy 2005
Coptognathus Burmeister 1847
Coscinocephalus Prell 1936
Cryptodus Mac Leay 1819
Cyphonistes Burmeister 1847
Dalgopus Dechambre 2001
Dasygnathus Macleay 1819
Democrates Burmeister 1847
Denhezia Dechambre 2006
Dichodontus Burmeister 1847
Dyscinetus Harold 1869
Endroedianibe Chalumeau 1981
Enema Hope 1837
Eophileurus Arrow 1908
Epironastes Carne 1957
Erioscelis Burmeister 1847
Eupatorus Burmeister 1847
Eutyctus Semenov 1889
Gibboryctes Endrödi 1974
Golofa Hope 1837
Haplophileurus Kolbe 1910
Haplosoma Semenov 1889
Harposceles Burmeister 1847
Hatamus Sharp 1877
Hemicyrthus Reiche 1860
Hemiphileurus Kolbe 1910
Heteroglobus Dupuis & Dechambre 2008
Heterostriatus Dupuis 2006
Hexodon Olivier 1789
Homoeomorphus Burmeister 1847
Horridocalia Endrodi 1974
Hovophileurus Arrow 1911
Indieraligus Dechambre 1979
Irazua Ratcliffe 2003
Licnostrategus Prell 1933
Lonchotus Burmeister 1847
Lycomedes Brême 1845
Macrocyphonistes Ohaus 1910
Megaceras Hope 1837
Megasoma Kirby 1825
Mellissius Wollaston 1869
Microphileurus Kolbe 1910
Microryctes Arrow 1908
Musurgus Vauloger 1898
Neocorynophyllus Carne 1985
Neodasygnathus Carne 1957
Neohyphus Heller 1896
Novapus Sharp 1875
Onychionyx Arrow 1914
Oryctoderus Boisduval 1835
Oryctophileurus Kolbe 1878
Papuana Arrow 1911
Parapucaya Prell 1934
Parisomorphus Schaufuss 1890
Paroryctoderus Dechambre 1994
Pentodontoschema Péringuey 1901
Phileurus Latreille 1807
Philoscaptus Brèthes 1919
Platyphileurus Ohaus 1910
Prionoryctes Arrow 1911
Pseudocavonus Blackburn 1910
Pseudosyrichthus Péringuey 1901
Pucaya Ohaus 1910
Ruteloryctes Arrow 1908
Scapanes Burmeister 1847
Siralus Dechambre 1998
Syrichthodontus Péringuey 1901
Syrichthoschema Janssens 1942
Talautoclyster Yamaya 2001
Trichogomphus Burmeister 1847
Trioplus Burmeister 1847
Wernoryctes
Xyloryctes Hope 1837
116 additional siblings truncated for brevity.
See the
resource file
for a full list.
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Cyclocephalini Laporte 1840
Cyclocephala Dejean 1821
(this page)
Cyclocephala aravaipensis Ratcliffe 1992
Cyclocephala arenosa Howden & Endrödi 1966
Cyclocephala barrerai Martínez 1969
Cyclocephala borealis Arrow 1911
Cyclocephala caelestis Delgado & Ratcliffe 1990
Cyclocephala comata Bates 1888
Cyclocephala deceptor (Casey 1915)
Cyclocephala forcipulata Howden & Endrödi 1966
Cyclocephala freudei Endrödi 1963
Cyclocephala hirta Le Conte 1861
Cyclocephala knobelae (Brown 1934)
Cyclocephala longula Le Conte 1863
Cyclocephala lunulata Burmeister 1847
Cyclocephala lurida Bland 1863
Cyclocephala mafaffa Burmeister 1847
Cyclocephala melanocephala (Fabricius 1775)
Cyclocephala miamiensis Howden & Endrödi 1966
Cyclocephala nigricollis Burmeister 1847
Cyclocephala parallela (Casey 1915)
Cyclocephala pasadenae (Casey 1915)
Cyclocephala pilosicollis Saylor 1936
Cyclocephala puberula Le Conte 1863
Cyclocephala regularis Casey 1915
Cyclocephala sanguinicollis Burmeister 1847
Cyclocephala seditiosa Le Conte 1863
Cyclocephala sinaloae Howden & Endrödi 1966
Cyclocephala sororia Bates 1888
Cyclocephala sparsa Arrow 1902
Cyclocephala testacea Burmeister 1847
Cyclocephala variabilis Burmeister 1847
Cyclocephala wandae Hardy 1974
Ancognatha Erichson 1847
Dyscinetus Harold 1869
NCBI
root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Cyclocephala
(this page)
Cyclocephala aequatoria
Cyclocephala almitana
Cyclocephala amazona
Cyclocephala amblyopsis
Cyclocephala atripes
Cyclocephala borealis
Cyclocephala brevis
Cyclocephala carbonaria
Cyclocephala concolor
Cyclocephala confusa
Cyclocephala conspicua
Cyclocephala deceptor
Cyclocephala discolor
Cyclocephala erotylina
Cyclocephala fasciolata
Cyclocephala freyi
Cyclocephala gravis
Cyclocephala inca
Cyclocephala insulicola
Cyclocephala kaszabi
Cyclocephala ligyrina
Cyclocephala longula
Cyclocephala lunulata
Cyclocephala lurida
Cyclocephala mafaffa
Cyclocephala melanocephala
Cyclocephala multiplex
Cyclocephala nigerrima
Cyclocephala nigritarsis
Cyclocephala pan
Cyclocephala porioni
Cyclocephala sanguinicollis
Cyclocephala sexpunctata
Cyclocephala signaticollis
Cyclocephala sparsa
Cyclocephala stictica
Cyclocephala stockwelli
Cyclocephala tridentata
Cyclocephala tridentata tridentata × Cyclocephala insulicola
Cyclocephala tutilina
Cyclocephala weidneri
Cyclocephala williami
Cyclocephala zodion
unclassified Cyclocephala
Agaocephala
Alissonotum
Allomyrina
Amblyoproctus
Ancognatha
Aspidolea
Augosoma
Beckius
Bothynus
Calicnemis
Carneodon
Chalcosoma
Cheiroplatys
Coelosis
Cryptodus
Cyphonistes
Diloboderus
Dynastes
Dyscinetus
Enema
Eophileurus
Euetheola
Eupatorus
Eutyctus
Golofa
Haploscapanes
Hemiphileurus
Heterogomphus
Hexodon
Homophileurus
Lycomedes
Marronus
Megaceras
Megasoma
Metanastes
Mimeoma
Neocorynophyllus
Novapus
Orizabus
Oryctes
Oryctoderus
Pachyoryctes
Papuana
Parapucaya
Peltonotus
Pentodon
Phileurus
Phyllognathus
Pimelopus
Pucaya
Pycnoschema
Semanopterus
Spodistes
Stenocrates
Strategus
Syrictes
Temnorhynchus
Tomarus
Trichogomphus
Trissodon
Trypoxylus
unclassified Dynastinae
Xyloryctes
Xylotrupes
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Cyclocephalini
Cyclocephala
(this page)
Cyclocephala abrelata
Cyclocephala acoma
Cyclocephala acuta
Cyclocephala aequatoria
Cyclocephala alazonia
Cyclocephala alexi
Cyclocephala almitana
Cyclocephala altamontana
Cyclocephala alutacea
Cyclocephala alvarengai
Cyclocephala amazona
Cyclocephala amblyopsis
Cyclocephala ampliata
Cyclocephala amplitarsis
Cyclocephala anibali
Cyclocephala antoinei
Cyclocephala aravaipensis
Cyclocephala arenosa
Cyclocephala arnaudi
Cyclocephala arrowiana
Cyclocephala atricapilla
Cyclocephala atriceps
Cyclocephala atricolor
Cyclocephala atripes
Cyclocephala barrerai
Cyclocephala batesi
Cyclocephala bella
Cyclocephala berti
Cyclocephala bicolor
Cyclocephala bicolorata
Cyclocephala bimaculata
Cyclocephala binotata
Cyclocephala bleuzeni
Cyclocephala boliviana
Cyclocephala bollei
Cyclocephala borburatae
Cyclocephala borealis
Cyclocephala boucheri
Cyclocephala boulardi
Cyclocephala brasiliana
Cyclocephala brevipennis
Cyclocephala brevis
Cyclocephala brittoni
Cyclocephala burmeisteri
Cyclocephala caelestis
Cyclocephala camachicola
Cyclocephala capitata
Cyclocephala carbonaria
Cyclocephala cardini
Cyclocephala carinatipennis
Cyclocephala carlsoni
Cyclocephala cartwrighti
Cyclocephala casanova
Cyclocephala castanea
Cyclocephala castaniella
Cyclocephala caussaneli
Cyclocephala cearae
Cyclocephala celata
Cyclocephala cerea
Cyclocephala chalumeaui
Cyclocephala chera
Cyclocephala chiquita
Cyclocephala colasi
Cyclocephala collaris
Cyclocephala comata
Cyclocephala compacta
Cyclocephala complanata
Cyclocephala concolor
Cyclocephala confusa
Cyclocephala conspicua
Cyclocephala contraria
Cyclocephala coriacea
Cyclocephala couturieri
Cyclocephala crassa
Cyclocephala crepuscularis
Cyclocephala cribrata
Cyclocephala cubana
Cyclocephala curta
Cyclocephala dalensi
Cyclocephala danforthi
Cyclocephala deceptor
Cyclocephala dechambrei
Cyclocephala decorella
Cyclocephala defecta
Cyclocephala deltoides
Cyclocephala dichroa
Cyclocephala dilatata
Cyclocephala diluta
Cyclocephala discicollis
Cyclocephala discolor
Cyclocephala dispar
Cyclocephala distincta
Cyclocephala divaricata
Cyclocephala dolichotarsa
Cyclocephala dominicana
Cyclocephala duodecimpunctata
Cyclocephala dupuisi
Cyclocephala durantonorum
Cyclocephala dyscinetoides
Cyclocephala emarginata
Cyclocephala endroedii
Cyclocephala endroedy
Cyclocephala endroedy-youngai
Cyclocephala englemani
Cyclocephala enigma
Cyclocephala epistomalis
Cyclocephala ergastuli
Cyclocephala erotylina
Cyclocephala everardoi
Cyclocephala falsa
Cyclocephala fankhaeneli
Cyclocephala fasciolata
Cyclocephala ferruginea
Cyclocephala figurata
Cyclocephala flavipennis
Cyclocephala flavoscutellaris
Cyclocephala flora
Cyclocephala forcipulata
Cyclocephala forsteri
Cyclocephala freudei
Cyclocephala freyi
Cyclocephala frontalis
Cyclocephala fulgurata
Cyclocephala fulvipennis
Cyclocephala fusiformis
Cyclocephala gabaldoni
Cyclocephala genieri
Cyclocephala gigantea
Cyclocephala goetzi
Cyclocephala gravis
Cyclocephala gregaria
Cyclocephala guaguarum
Cyclocephala guianae
Cyclocephala guttata
Cyclocephala guycolasi
Cyclocephala halffteriana
Cyclocephala hardyi
Cyclocephala hartmannorum
Cyclocephala helavai
Cyclocephala herteli
Cyclocephala hiekei
Cyclocephala hielkemaorum
Cyclocephala hirsuta
Cyclocephala hirta
Cyclocephala histrionica
Cyclocephala holmbergi
Cyclocephala howdenannae
Cyclocephala husingi
Cyclocephala hylaea
Cyclocephala iani
Cyclocephala immaculata
Cyclocephala inca
Cyclocephala insulicola
Cyclocephala isabellina
Cyclocephala isthmiensis
Cyclocephala italoi
Cyclocephala jalapensis
Cyclocephala jauffreti
Cyclocephala kahanoffae
Cyclocephala kaszabi
Cyclocephala kechua
Cyclocephala knobelae
Cyclocephala krombeini
Cyclocephala kuntzeniana
Cyclocephala labidion
Cyclocephala lachaumei
Cyclocephala laevis
Cyclocephala lamarcki
Cyclocephala laminata
Cyclocephala larssoni
Cyclocephala latericia
Cyclocephala latipennis
Cyclocephala latreillei
Cyclocephala lecourti
Cyclocephala letiranti
Cyclocephala lichyi
Cyclocephala ligyrina
Cyclocephala lineata
Cyclocephala lineigera
Cyclocephala liomorpha
Cyclocephala literata
Cyclocephala lizeri
Cyclocephala longa
Cyclocephala longicollis
Cyclocephala longimana
Cyclocephala longitarsis
Cyclocephala longula
Cyclocephala lunulata
Cyclocephala lurida
Cyclocephala lutea
Cyclocephala machadoi
Cyclocephala macrophylla
Cyclocephala maculata
Cyclocephala maculiventris
Cyclocephala mafaffa
Cyclocephala magdalenae
Cyclocephala malleri
Cyclocephala malyi
Cyclocephala mannheimsi
Cyclocephala marginalis
Cyclocephala marginicollis
Cyclocephala marianista
Cyclocephala marqueti
Cyclocephala martinezi
Cyclocephala marylizae
Cyclocephala mateoi
Cyclocephala mathani
Cyclocephala mecynotarsis
Cyclocephala megalophylla
Cyclocephala meinanderi
Cyclocephala melanae
Cyclocephala melanocephala
Cyclocephala melanopoda
Cyclocephala melolonthida
Cyclocephala mesophylla
Cyclocephala metrica
Cyclocephala miamiensis
Cyclocephala minuchae
Cyclocephala minuta
Cyclocephala modesta
Cyclocephala molesta
Cyclocephala monacha
Cyclocephala monzoni
Cyclocephala moreti
Cyclocephala morphoidina
Cyclocephala multiplex
Cyclocephala munda
Cyclocephala mustacha
Cyclocephala mutata
Cyclocephala nana
Cyclocephala nicolasi
Cyclocephala nigerrima
Cyclocephala nigra
Cyclocephala nigricollis
Cyclocephala nigritarsis
Cyclocephala nigrobasalis
Cyclocephala nigropicta
Cyclocephala niguasa
Cyclocephala nike
Cyclocephala nodanotherwon
Cyclocephala notata
Cyclocephala obscura
Cyclocephala occipitalis
Cyclocephala ocellata
Cyclocephala ochracea
Cyclocephala octopunctata
Cyclocephala ohausiana
Cyclocephala olivieri
Cyclocephala ovulum
Cyclocephala pan
Cyclocephala panthera
Cyclocephala paraflora
Cyclocephala paraguayensis
Cyclocephala parallela
Cyclocephala pardolocarnoi
Cyclocephala pasadenae
Cyclocephala perconfusa
Cyclocephala pereirai
Cyclocephala perforata
Cyclocephala perplexa
Cyclocephala peruana
Cyclocephala pichinchana
Cyclocephala picipes
Cyclocephala picopijola
Cyclocephala picta
Cyclocephala pilosa
Cyclocephala pilosicollis
Cyclocephala pokornyi
Cyclocephala pompanoni
Cyclocephala poncheli
Cyclocephala porioni
Cyclocephala prelli
Cyclocephala prolongata
Cyclocephala proxima
Cyclocephala pseudoconfusa
Cyclocephala pseudomelanocephala
Cyclocephala puberula
Cyclocephala pugnax
Cyclocephala pulchra
Cyclocephala puncticollis
Cyclocephala putrida
Cyclocephala pygidialis
Cyclocephala pygidiata
Cyclocephala quadripunctata
Cyclocephala quatuordecimpunctata
Cyclocephala quercina
Cyclocephala quisqueya
Cyclocephala rangelana
Cyclocephala recta
Cyclocephala regularis
Cyclocephala robusta
Cyclocephala rogerpauli
Cyclocephala rogezi
Cyclocephala rondoniana
Cyclocephala rorulenta
Cyclocephala rotundipenis
Cyclocephala rubescens
Cyclocephala rufa
Cyclocephala rufescens
Cyclocephala ruficollis
Cyclocephala rufonigra
Cyclocephala rufovaria
Cyclocephala rustica
Cyclocephala saltini
Cyclocephala sanguinicollis
Cyclocephala santaritae
Cyclocephala sarahae
Cyclocephala sardadebiae
Cyclocephala sarpedon
Cyclocephala scarabaeina
Cyclocephala schmitzorum
Cyclocephala seditiosa
Cyclocephala setosa
Cyclocephala sexpunctata
Cyclocephala signaticollis
Cyclocephala signatoides
Cyclocephala similis
Cyclocephala simillima
Cyclocephala simulatrix
Cyclocephala sinaloae
Cyclocephala sinuosa
Cyclocephala sororia
Cyclocephala stictica
Cyclocephala stockwelli
Cyclocephala striata
Cyclocephala subsignata
Cyclocephala supernana
Cyclocephala suturalis
Cyclocephala sylviae
Cyclocephala tarsalis
Cyclocephala testacea
Cyclocephala tetrica
Cyclocephala tidula
Cyclocephala toulgoeti
Cyclocephala tridentata
Cyclocephala tronchonii
Cyclocephala tucumana
Cyclocephala tutilina
Cyclocephala tylifera
Cyclocephala ukuku
Cyclocephala unamas
Cyclocephala undata
Cyclocephala unidentata
Cyclocephala variabilis
Cyclocephala varians
Cyclocephala variipenis
Cyclocephala variolosa
Cyclocephala verticalis
Cyclocephala vestita
Cyclocephala vidanoi
Cyclocephala vidua
Cyclocephala villosa
Cyclocephala vincentiae
Cyclocephala vinosa
Cyclocephala virgo
Cyclocephala viridis
Cyclocephala vittoscutellaris
Cyclocephala wandae
Cyclocephala warneri
Cyclocephala weidneri
Cyclocephala williami
Cyclocephala zischkai
Cyclocephala zodion
Cyclocephala zurstrasseni
Ancognatha
Chalepides
Dyscinetus
Harposcelis
Harposcelis paradoxus
Peltonotus
Ruteloryctes
Stenocrates