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Creatures » … » Animal » … » Arthropods » … » Hexapods » Insects » Winged Insects » … » Endopterygotes » Beetles » … » Dynastidae

Heteronychus

Scientific Names

Preferred Names

Heteronychus
Genus recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Heteronychus
Genus recognized by wikipedia NO, wikipedia SV, EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, Australia Species List, Democratic Republic of the Congo Species List, New Zealand Species List, South Africa Species List, Tanzania Species List, Zambia Species List, Zimbabwe Species List, Indian Ocean Species List, South Pacific Species List, NCBI, wikidata_hierarchy tar gz, wikipedia FR, wikipedia EN, Wikidata Common Names, Wikimedia Commons, Barcode of Life Data Systems, Global Biotic Interactions, iNaturalist data coverage, Wikipedia (inferred records), Dynamic Hierarchy Version 2.2.3 - Test, and GGBN data coverage
Heteronychus
Recognized by OBIS environmental data records
Heteronychus Burmeister 1847
Genus recognized by GBIF classification and Mozambique Species List
Heteronychus Dejean 1833
Genus recognized by Catalogue of Life 26 March 2024 and GBIF data coverage
Heteronychus subgen. Heteronychus Dejean 1833
Subgenus recognized by Catalogue of Life 26 March 2024

Alternative Names

Heteronychus consimilis Jack 1923
Heterotypic synonym according to GBIF classification
Heteronychus sacchari Paoli 1934
Heterotypic synonym according to GBIF classification
Heteronychus
Harvest ancestor according to GBIF national node type records France, Mozambique Species List, iNaturalist, Simpson et al 2020, Global Biotic Interactions, Global Biotic Interactions, and Global Biotic Interactions

Common Names

Chinese

异爪犀金龟属 preferred
Recognized by Wikidata Common Names

Zh Cn

异爪犀金龟属 preferred
Recognized by Wikidata Common Names

Curated hierarchies for Heteronychus

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Heteronychus (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus atratus Klug 1855
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Pentodontini Mulsant 1842
Pentodontina Mulsant 1842
Heteronychus Dejean 1833 (this page)
Heteronychus subgen. Heteronychidius Paulian 1954
Heteronychus subgen. Heteronychus Dejean 1833
Aceratus Prell 1936
Adelaeus Ratcliffe 2014
Alissonotum Arrow 1908
Ampotis Dechambre & Madge 1979
Anoplognatho Rivers 1889
Anoronotum Arrow 1939
Aphonides Rivers 1889
Aphonodelus Kolbe 1905
Aphonus Le Conte 1856
Barutus Ratcliffe 1981
Bothynus Hope 1837
Calcitoryctes Krell 2011
Calicnemis Laporte 1832
Callistemonus Péringuey 1901
Carneiola Endrödi 1974
Carneodon Özdikmen 2009
Collagenus Ratcliffe & Hardy 2005
Coptognathus Burmeister 1847
Coscinocephalus Prell 1936
Dalgopus Dechambre 2001
Denhezia Dechambre 2006
Diloboderus Reiche 1859
Ebolowanius Dechambre 1979
Enarotadius Endrödi 1971
Eremobothynus Ohaus 1910
Eucopidocaulus Prell 1912
Euetheola Bates 1888
Euligyrus Casey 1915
Eutyctus Semenov 1890
Gillaspytes Howden 1980
Gorditus Ratcliffe 2010
Haplosoma Semenov 1890
Heteroconus Kolbe 1900
Heteroglobus Dupuis & Dechambre 2008
Heteroligus Kolbe 1900
Heterostriatus Dupuis 2006
Hiekeianus Endrödi 1978
Homoeomorphus Burmeister 1847
Hyboschema Péringuey 1901
Hylobothynus Ohaus 1910
Hyphoryctes Blackburn 1895
Ligyrus Burmeister 1847
Lonchotus Burmeister 1847
Marronus Coquerel 1866
Mellissius Wollaston 1869
Metanastes Arrow 1911
Microryctes Arrow 1908
Musurgus Vauloger 1898
Neocorynophyllus Carne 1985
Neometanastes Yamaya 2010
Neonastes Carne 1957
Neoryctes Arrow 1908
Nimbacola Paulian 1970
Orizabus Fairmaire 1878
Orsilochus Burmeister 1847
Oxyligyrus Arrow 1908
Papuana Arrow 1911
Paranodon Cockerell 1905
Pareteronychus Prell 1934
Parisomorphus Schaufuss 1890
Pentodina Endrödi 1968
Pentodon Hope 1837
Pentodontoschema Péringuey 1901
Pericoptus Burmeister 1847
Philoscaptus Brèthes 1919
Phyllognathus Eschscholtz 1830
Pimelopus Erichson 1842
Piscoperus Ratcliffe & Giraldo 2014
Podalgus Burmeister 1847
Prionoryctes Arrow 1911
Proculigyrus López-García & Deloya 2022
Pseudohomonyx Arrow 1908
Pycnoschema Thomson 1859
Saccharoscaptus Morón & Grossi 2015
Siralus Dechambre 1998
Temnorhynchus Hope 1837
Thronistes Burmeister 1847
Tomarus Erichson 1847
Tribialus Dupuis & Perrin 2022
Wernoryctes Dechambre 1999

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Pentodontini Mulsant 1842
Pentodontina Mulsant 1842
Heteronychus Dejean 1833
Heteronychus subgen. Heteronychus Dejean 1833 (this page)
Heteronychus (Heteronychus) abyssinicus Jack 1923
Heteronychus (Heteronychus) amplipennis Benderitter 1915
Heteronychus (Heteronychus) amplus Kolbe 1900
Heteronychus (Heteronychus) andersoni Jack 1923
Heteronychus (Heteronychus) annulatus Bates 1891
Heteronychus (Heteronychus) approximans Kolbe 1900
Heteronychus (Heteronychus) arator (Fabricius 1775)
Heteronychus (Heteronychus) ascanius Kolbe 1900
Heteronychus (Heteronychus) atratus Klug 1855
Heteronychus (Heteronychus) basilewskyi Endrödi 1962
Heteronychus (Heteronychus) bituberculatus Kolbe 1900
Heteronychus (Heteronychus) brittoni Ferreira 1965
Heteronychus (Heteronychus) carvalhoi Ferreira 1970
Heteronychus (Heteronychus) citernii Paoli 1934
Heteronychus (Heteronychus) congoensis Kolbe 1900
Heteronychus (Heteronychus) consimilis Kolbe 1900
Heteronychus (Heteronychus) cordatus Endrödi 1974
Heteronychus (Heteronychus) costatus Lansberge 1886
Heteronychus (Heteronychus) cricetus (Hausmann 1807)
Heteronychus (Heteronychus) curtulus Fairmaire 1887
Heteronychus (Heteronychus) desaegeri Burgeon 1947
Heteronychus (Heteronychus) digitiformis Zhang 1983
Heteronychus (Heteronychus) fossor Reiche 1850
Heteronychus (Heteronychus) howdeni Yamaya 2016
Heteronychus (Heteronychus) impudens Jack 1923
Heteronychus (Heteronychus) infans Kolbe 1897
Heteronychus (Heteronychus) inoportunus Ferreira 1966
Heteronychus (Heteronychus) insignificus Jack 1923
Heteronychus (Heteronychus) intermedius Zhang 1983
Heteronychus (Heteronychus) jacki Arrow 1936
Heteronychus (Heteronychus) licas (Klug 1835)
Heteronychus (Heteronychus) lioderes Redtenbacher 1868
Heteronychus (Heteronychus) lusingae Endrödi 1974
Heteronychus (Heteronychus) minimus Dupuis 2012
Heteronychus (Heteronychus) mollis Endrödi 1961
Heteronychus (Heteronychus) monodi Paulian 1954
Heteronychus (Heteronychus) mosambicus Péringuey 1908
Heteronychus (Heteronychus) muticus Benderitter 1915
Heteronychus (Heteronychus) paolii Arrow 1936
Heteronychus (Heteronychus) parumpunctatus Burmeister 1847
Heteronychus (Heteronychus) parvus Burmeister 1847
Heteronychus (Heteronychus) pauperatus Péringuey 1901
Heteronychus (Heteronychus) plebejus (Klug 1834)
Heteronychus (Heteronychus) puerilis Kolbe 1900
Heteronychus (Heteronychus) puncticollis Jack 1923
Heteronychus (Heteronychus) punctolineatus Fairmaire 1893
Heteronychus (Heteronychus) pygidialis Kolbe 1900
Heteronychus (Heteronychus) rusticus (Klug 1834)
Heteronychus (Heteronychus) sabackyi Endrödi 1975
Heteronychus (Heteronychus) sacchari Arrow 1908
Heteronychus (Heteronychus) similis Dechambre 1992
Heteronychus (Heteronychus) simulans Jack 1923
Heteronychus (Heteronychus) sublaevis (Fairmaire 1891)
Heteronychus (Heteronychus) tenuestriatus Fairmaire 1893
Heteronychus (Heteronychus) tesari Endrödi 1968
Heteronychus (Heteronychus) tristis Boheman 1857
Heteronychus (Heteronychus) vixstriatus Jack 1923
Heteronychus (Heteronychus) wittei Janssens 1942
Heteronychus subgen. Heteronychidius Paulian 1954

Dynamic Hierarchy Version 2.2.3 - Test

Heteronychus (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus atratus Klug 1855
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Heteronychus (this page)
Heteronychus (Heteronychidius) minutus Burmeister 1847
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus atratus Klug 1855
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942
Aceratus
Adoryphorus
Aegopsis
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calcitoryctes
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
118 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Heteronychus Burmeister 1847 (this page)
Heteronychus abyssinicus Jack 1923
Heteronychus amplipennis Benderitter 1915
Heteronychus amplus Kolbe 1900
Heteronychus andersoni Jack 1923
Heteronychus annulatus Bates 1891
Heteronychus approximans Kolbe 1900
Heteronychus arator (Fabricius 1775)
Heteronychus ascanius Kolbe 1900
Heteronychus atratus Klug 1855
Heteronychus basilewskyi Endrödi 1962
Heteronychus bituberculatus Kolbe 1900
Heteronychus brittoni Ferreira 1965
Heteronychus carvalhoi Ferreira 1970
Heteronychus citernii Paoli 1934
Heteronychus congoensis Kolbe 1900
Heteronychus consimilis Kolbe 1900
Heteronychus cordatus Endrödi 1974
Heteronychus costatus Lansberge 1886
Heteronychus cricetus (Hausmann 1807)
Heteronychus curtulus Fairmaire 1887
Heteronychus desaegeri Burgeon 1947
Heteronychus digitiformis Zhang 1983
Heteronychus egenis
Heteronychus fossor Reiche 1847
Heteronychus howdeni Yamaya 2016
Heteronychus impudens Jack 1923
Heteronychus infans Kolbe 1897
Heteronychus inoportunus Ferreira 1966
Heteronychus insignificus Jack 1923
Heteronychus intermedius Zhang 1983
Heteronychus jacki Arrow 1936
Heteronychus krombeini Endrödi 1976
Heteronychus licas (Klug 1835)
Heteronychus lioderes Redtenbacher 1867
Heteronychus lusingae Endrödi 1974
Heteronychus minimus Dupuis 2012
Heteronychus minutus Burmeister 1847
Heteronychus mollis Endrödi 1961
Heteronychus monodi Paulian 1954
Heteronychus mosambicus Péringuey 1908
Heteronychus muticus Benderitter 1915
Heteronychus paolii Arrow 1936
Heteronychus parumpunctatus Burmeister 1847
Heteronychus parvus Burmeister 1847
Heteronychus pauperatus Péringuey 1901
Heteronychus plebejus (Klug 1832)
Heteronychus puerilis Kolbe 1900
Heteronychus puncticollis Jack 1923
Heteronychus punctolineatus Fairmaire 1893
Heteronychus pygidialis Kolbe 1900
Heteronychus rusticus (Klug 1832)
Heteronychus sabackyi Endrödi 1975
Heteronychus sacchari Arrow 1908
Heteronychus similis Dechambre 1992
Heteronychus simulans Jack 1923
Heteronychus sublaevis (Fairmaire 1891)
Heteronychus tenuestriatus Fairmaire 1893
Heteronychus tenvestriatus
Heteronychus tesari Endrödi 1968
Heteronychus tristis Boheman 1857
Heteronychus vixstriatus Jack 1923
Heteronychus wittei Janssens 1942
Aceratus Prell 1936
Acrobolbia Ohaus 1912
Aegopsis Burmeister 1847
Agaocephala Le Peletier & Serville 1828
Allophileurinus Dupuis & Dechambre 2001
Amblyodus Westwood 1878
Amblyoproctus Kolbe 1910
Ancognatha Erichson 1847
Aphonodelus Kolbe 1905
Archophileurus Kolbe 1910
Aspidolea Bates 1888
Augosoma Burmeister 1841
Blabephorus Fairmaire 1898
Bothynus Hope 1837
Calicnemis Laporte de Castelnau 1832
Carneiola Endrödi 1974
Carneoryctes Özdikmen 2009
Cavonus Sharp 1875
Ceratophileurus Ohaus 1911
Ceratoryctoderus Arrow 1908
Chalcasthenes Arrow 1937
Chalcosoma Hope 1837
Cheiroplatys Kirby 1837
Chiliphileurus Endrodi 1977
Cnemidophileurus Kolbe 1910
Coenoryctoderus Prell 1933
Colacus Ohaus 1910
Collagenus Ratcliffe & Hardy 2005
Coptognathus Burmeister 1847
Coscinocephalus Prell 1936
Cryptodus Mac Leay 1819
Cyphonistes Burmeister 1847
Dalgopus Dechambre 2001
Dasygnathus Macleay 1819
Democrates Burmeister 1847
Denhezia Dechambre 2006
Dichodontus Burmeister 1847
Dyscinetus Harold 1869
Endroedianibe Chalumeau 1981
Enema Hope 1837
Eophileurus Arrow 1908
Epironastes Carne 1957
Erioscelis Burmeister 1847
Eupatorus Burmeister 1847
Eutyctus Semenov 1889
Gibboryctes Endrödi 1974
Golofa Hope 1837
Haplophileurus Kolbe 1910
Haplosoma Semenov 1889
Harposceles Burmeister 1847
Hatamus Sharp 1877
Hemicyrthus Reiche 1860
Hemiphileurus Kolbe 1910
Heteroglobus Dupuis & Dechambre 2008
Heterostriatus Dupuis 2006
Hexodon Olivier 1789
Homoeomorphus Burmeister 1847
Horridocalia Endrodi 1974
Hovophileurus Arrow 1911
Indieraligus Dechambre 1979
Irazua Ratcliffe 2003
Licnostrategus Prell 1933
Lonchotus Burmeister 1847
Lycomedes Brême 1845
Macrocyphonistes Ohaus 1910
Megaceras Hope 1837
Megasoma Kirby 1825
Mellissius Wollaston 1869
Microphileurus Kolbe 1910
Microryctes Arrow 1908
Musurgus Vauloger 1898
Neocorynophyllus Carne 1985
Neodasygnathus Carne 1957
Neohyphus Heller 1896
Novapus Sharp 1875
Onychionyx Arrow 1914
Oryctoderus Boisduval 1835
Oryctophileurus Kolbe 1878
Papuana Arrow 1911
Parapucaya Prell 1934
Parisomorphus Schaufuss 1890
Paroryctoderus Dechambre 1994
Pentodontoschema Péringuey 1901
Phileurus Latreille 1807
Philoscaptus Brèthes 1919
Platyphileurus Ohaus 1910
Prionoryctes Arrow 1911
Pseudocavonus Blackburn 1910
Pseudosyrichthus Péringuey 1901
Pucaya Ohaus 1910
Ruteloryctes Arrow 1908
Scapanes Burmeister 1847
Siralus Dechambre 1998
Syrichthodontus Péringuey 1901
Syrichthoschema Janssens 1942
Talautoclyster Yamaya 2001
Trichogomphus Burmeister 1847
Trioplus Burmeister 1847
Wernoryctes
Xyloryctes Hope 1837
116 additional siblings truncated for brevity.
See the resource file for a full list.

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Rutelinae
Heteronychus (this page)
Heteronychus arator
Heteronychus lioderes
unclassified Heteronychus
Adoretus
Anisoplia
Anomala
Anomalorhina
Anoplognathus
Anoplostethus
Anticheira
Blithopertha
Blitopertha
Brachysternus
Callistethus
Calloodes
Chaetopteroplia
Chlorota
Chrysina
Cotalpa
Exomala
Hylamorpha
Isoplia
Lepadoretus
Mimadoretus
Mimela
Oryctomorphus
Paracotalpa
Paraschizognathus
Parastasia
Pelidnota
Phalangogonia
Phyllopertha
Platycoelia
Plusiotis
Popillia
Proagopertha
Prodoretus
Repsimus
Rutela
Schizognathus
Spodochlamys
Strigoderma
Trigonostomum
Trioplognathus
unclassified Rutelinae

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Pentodontini
Heteronychus (this page)
Heteronychidius
Heteronychus abyssinicus
Heteronychus amplipennis
Heteronychus amploides
Heteronychus amplus
Heteronychus andersoni
Heteronychus annulatus
Heteronychus approximans
Heteronychus arator
Heteronychus ascanius
Heteronychus atratus
Heteronychus basilewskyi
Heteronychus bituberculatus
Heteronychus brittoni
Heteronychus carvalhoi
Heteronychus citernii
Heteronychus congoensis
Heteronychus consimilis
Heteronychus cordatus
Heteronychus costatus
Heteronychus cricetus
Heteronychus curtulus
Heteronychus desaegeri
Heteronychus digitiformis
Heteronychus fossor
Heteronychus howdeni
Heteronychus humilis
Heteronychus impudens
Heteronychus infans
Heteronychus inoportunus
Heteronychus insignificus
Heteronychus intermedius
Heteronychus jacki
Heteronychus krombeini
Heteronychus licas
Heteronychus lioderes
Heteronychus lusingae
Heteronychus minimus
Heteronychus minutus
Heteronychus mollis
Heteronychus monodi
Heteronychus mosambicus
Heteronychus muticus
Heteronychus paolii
Heteronychus parumpunctatus
Heteronychus parvus
Heteronychus pauperatus
Heteronychus plebejus
Heteronychus puerilis
Heteronychus puncticollis
Heteronychus punctolineatus
Heteronychus pygidialis
Heteronychus rusticus
Heteronychus sabackyi
Heteronychus sacchari
Heteronychus similis
Heteronychus simulans
Heteronychus sublaevis
Heteronychus tenuestriatus
Heteronychus tesari
Heteronychus tristis
Heteronychus vixstriatus
Heteronychus wittei
Seidlitz
Ampotis
Anaronotum
Anoplognatho
Australocus
Barutus
Callistemonus
Callistemonus intrusus
Cheiroplatina
Coptognatus
Dasygnathus
Dipelicina
Dipelicus
Eutheola
Heikeianus
Heterostriatus
Hylobothynus
Idioschema karruense
Marronus
Microryctes
Neobothynus
Neoryctes
Nimbacola
Orizabus
Oxyligyrus
Papuana
Parabothynus
Paranodon
Pentodon
Pentodontina
Pericoptus
Piscoperus
Podalgus
Pseudocavonus
Pseudoryctina
Pycnoschema
Temnorrhynchus
Tomarus
Wernoryctes
Wernoryctes werneri
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