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There is unpublished evidence that this fish species has been found in an additional river-watershed in northern Wisconsin and Michigan. An entrainment mortality study was conducted for Wisconsin Public Service (WPS), on the Menominee River, as part of their relicensing requirements for the Federal Energy Regulatory Commission (FERC). This study took place between May 1993 and April 1994. During the summer of 1993, 28 discreet samples, collected on 17 different sampling dates produced 104 specimens of this species (WPS 1994). After verification of the identification, several specimens were donated to the University of Michigan Museum of Zoology. Their accession number is 224173(Personal comm).

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Behavior

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Perception Channels: tactile ; chemical

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Conservation Status

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Reports from various agencies around the country indicate that the western sand darter is declining in abundance. Especially hard hit have been the populations in the central part of the range for the species. It is listed as rare in Illinois, depleted in Missouri, threatened in Iowa, and extirpated in Kansas. Wisconsin has placed the western sand darter in the Lake Michigan drainage on watch status. According to Cross (1967), fluctuating water levels and increasing siltation are especially detrimental to sand darters (Becker 1983). Stream degradation related to siltation is also cited as negatively impacting western sand darter populations (Page 1983).

US Federal List: no special status

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Benefits

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The western sand darter has no adverse affect on the human environment.

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Benefits

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There is little evidence that this species has a positive or negative benefit to humans. Because of its small, slender body it is not suitable as a bait fish (Becker 1983). However, the continued existence of this species will be directly influenced by humans. As has been stated elsewhere, the decline of the western sand darter has been directly linked to the degradation of aquatic habitats throughout the natural range of the sand darter-as the result of human activities. Perhaps the greatest benefit that Americans can derive from the western sand darter by its continued existence will be its contribution to the biodiversity of North America.

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Trophic Strategy

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The few studies that have been performed indicate that major food items include small or immature aquatic insects such as mayflies (Ephemeroptera) and midge larvae (Diptera:Chironomidae)

(Lutterbie 1976).

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Distribution

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Sporadic in the Mississippi drainage and adjacent Great Lakes drainage with small disjunct populations throughout this range. (Becker 1983)

It occurs as far south as the Naches and Sabine Rivers in eastern Texas (Williams 1975).

Biogeographic Regions: nearctic (Native )

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Habitat

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The western sand darter is found in medium to large rivers that have moderate to swift currents, primarily over extensive areas of sandy substrate. It is generally found in water that ranges from about .2-.9 meters in depth. Water preference of the western sand darter is for clear to slightly turbid water (Becker 1983).

Aquatic Biomes: benthic ; rivers and streams

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Life Expectancy

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Average lifespan
Status: captivity:
3 years.

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Morphology

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Ammocrypta clara is a small, very slender fish. Its average adult size ranges from 42-67mm (1.7-2.6in.). The word ammocrypta means "concealed in sand" and clara means "clear"-which is a reference to its clear or translucent flesh. As with almost all members of the family Percidae, the western sand darter has two dorsal fins. The first dorsal fin has 10-12 spines, and the second dorsal fin has 9-12 soft rays. The pectoral fins are fairly large and located directly behind the opercles; its pelvic fins are thoracic and located directly behind the head at the anterior end of the abdomen. Its fins range from clear to weakly pigmented along the spines and rays. Live specimens are very pallid-almost transparent. Preserved specimens are light tan or straw-colored. There is a row of twelve or more small dark saddles located down the center of the back. Along both sides of the fish are rows of 10-12 small oblong spots. The scales of the western sand darter are ctenoid. The breast and belly are completely devoid of scales; scales cover the dorsal surface from its median to the lateral line. The caudal peduncle is almost completely scaled. The caudal fin is slightly lobed (Becker 1983).

Until recently, the western sand darter was considered to be the same species as the eastern sand darter (Ammocrypta pellucida). The two species were separated based on scale counts, opercular structures and pigmentation. In contrast to the eastern form, the western sand darter has only 3-5 scale rows on the side of the body, a needlelike opercular spine and less pigmentation in the mid-dorsal and lateral blotches (Becker 1983).

For 47 western sand darters from central Wisconsin, the calculated lengths at the first three annuli were 43, 56 and 61 millimeters-almost identical to the calculated lengths for southern Wisconsin populations. The western sand darter reaches 71% of its total growth during its first year of life, and 91% during the second year. Females are generally larger and more numerous than males (Becker 1983).

Range mass: 1.5 (high) g.

Other Physical Features: bilateral symmetry

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Reproduction

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Little is known about the spawning requirements or the reproductive behavior of the western sand darter. It has been suggested that spawning may occur in mid-summer, judging by increased activity then and gravid females (Lutterbie 1976). By examining the ovaries and development of breeding tubercules, Williams(1975) placed the height of the breeding season in July and early August. Williams found females with mature ova in late August. In Wisconsin, western sand darters spawn from late June through July (Lutterbie 1976).

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Cowton, S. 2001. "Ammocrypta clara" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Ammocrypta_clara.html
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Stanley F. Cowton, Eastern Michigan University
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Cynthia Sims Parr, University of Michigan-Ann Arbor
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Biology

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Inhabits sandy runs of medium to large rivers (Ref. 5723) and streams (Ref. 10294). Feeds on aquatic insects (Ref. 79012).
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Western sand darter

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The western sand darter (Ammocrypta clara) is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the central United States.

Distribution

The western sand darter occurs in river systems from Lake Michigan to Texas, including several sections of the Mississippi Basin.[1] Its range extends as far east as the Elk River in West Virginia.[3]

Distribution map

Description

This species is up to 7.1 centimeters in length.[2] It is slender and nearly cylindrical in shape. It is pale, translucent silvery white with yellowish coloration along the back. It is distinguished from other sand darters the lack of dark bands or blotches, and by a spine on its operculum.[4]

Biology

This fish lives in medium and large rivers, over sandy and gravel substrates. It requires loose substrate, because it spends much of its time buried in the sand with just its head protruding.[4] This behavior helps it reach cooler temperatures.[5]

It feeds on invertebrates, especially the larvae of aquatic insects.[4]

It spawns in summer, starting in June in northern regions and May farther south.[4] Females produce an average of 57 eggs at a time, with larger females producing more eggs.[6]

Taxonomy

The Western sand darter was first formally described in 1885 by the American ichthyologists David Starr Jordan (1851-1931) and Seth Eugene Meek (1859-1914) with the type locality given as the Des Moines River at Ottumwa, Iowa.[7] This species forms a clade with the naked sand darter (A. beanii) the Florida sand darter (A. bifascia).[8]

Conservation

This is considered to be a vulnerable species because it has a fragmented distribution and its habitat is degraded in many areas. Increased silt and pollution in river systems reduces the quality of its habitat. The Mississippi River and associated streams and tributaries are heavily channelized and partitioned by locks and dams, eliminating sites where the fish might live.[1]

References

  1. ^ a b c NatureServe (2014). "Ammocrypta clara". IUCN Red List of Threatened Species. 2014: e.T202428A2744621. doi:10.2305/IUCN.UK.2014-3.RLTS.T202428A2744621.en. Retrieved 19 November 2021.
  2. ^ a b Froese, Rainer; Pauly, Daniel (eds.) (2019). "Ammocrypta clara" in FishBase. December 2019 version.
  3. ^ Dan A. Cincotta & Stuart A. Welsh (2010). "Discovery of Ammocrypta clara (Western Sand Darter) in the Upper Ohio River of West Virginia". The American Midland Naturalist. 163 (2): 318–325. doi:10.1674/0003-0031-163.2.318. JSTOR 40730928. S2CID 85926557.
  4. ^ a b c d "Ammocrypta clara Western Sand Darter". NatureServe explorer. NatureServe. Retrieved 20 September 2020.
  5. ^ Becker, George. "Fishes of Wisconsin". University of Wisconsin Press. Retrieved 23 April 2017.
  6. ^ Driver, L. J. & Adams, G. L. (2013). "Life history and spawning behavior of the western sand darter (Ammocrypta clara) in Northeast Arkansas" (PDF). The American Midland Naturalist. 170 (2): 199–212. doi:10.1674/0003-0031-170.2.199. JSTOR 23525570. S2CID 85675303.
  7. ^ Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Ammocrypta clara". Catalog of Fishes. California Academy of Sciences. Retrieved 20 September 2020.
  8. ^ James D. Williams (1975). "Systematics of the Percid Fishes of the Subgenus Ammocrypta, Genus Ammocrypta, with Descriptions of Two New Species". Bulletin of the Alabama Museum of Natural History (1): 1–56.

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Western sand darter: Brief Summary

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The western sand darter (Ammocrypta clara) is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the central United States.

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