Eunice guttata Baird 1869

Eunice guttata Baird, 1869

Eunice guttata Baird, 1869:350.

MATERIAL EXAMINED.—Holotype, BM(NH) ZB 1972.71, taken between Bombay and Singapore, Indian Ocean.

DESCRIPTION.—Holotype complete, of unknown sex, with 115 setigers; total length 52 mm; maximal width 3 mm at setiger 10; length through setiger 10, 8 mm.

Prostomium (Figure 53f) distinctly shorter and narrower than peristomium, as deep as of peristomium. Prostomial lobes frontally obliquely truncate, dorsally flattened; median sulcus deep. Eyes not observed. Antennae in shallow horseshoe, evenly spaced, similar in thickness. Ceratophores ring-shaped in all antennae, without articulations. Ceratostyles long, tapering, without articulations. A-I to posterior edge of peristomium; A-II to setiger 2; A-III to setiger 3. Peristomium slightly tapering anteriorly. Separation between rings distinct only dorsally; anterior ring of total peristomial length. Peristomial cirri to middle of anterior peristomial ring, tapering, without articulations.

Jaws not observed.

Branchiae (Figure 53g) present, pectinate, distinctly longer than notopodial cirri, not reduced in mid-body region, erect. Branchiae from setiger 6 to setiger 105. Branchiae present to near posterior end, present on more than 65% of total number of setigers. Where best developed branchiae distinctly pectinate with up to 10 filaments. Branchial stems nearly digitiform, longer than notopodial cirri. Filaments slender, digitiform, as long as or shorter than notopodial cirri.

Neuropodial acicular lobes conical or depressed conical; aciculae emerging at midline. Presetal lobes low, transverse folds. Anterior postsetal lobes rounded, flattened, as high as acicular lobe, becoming reduced to low, transverse folds in median and posterior setigers. Anterior ventral cirri tapering. Ventral cirri basally inflated from about setiger 20; ventral cirri remaining inflated through rest of body. Basal inflation very large, nearly spherical, very small tapering tips present in all setigers. Notopodial cirri basally inflated, distally tapering, without articulations.

Limbate setae tapering, gently geniculate, frayed. Pectinate setae (Figure 53i) numbering 5 or more in a fascicle. Shafts wide, flat. Blades tapering, flat. One marginal tooth clearly longer than other teeth, with ~14 teeth. Shafts of compound falcigers (Figure 53h) distally inflated, marginally smooth. Appendages very slender, tapering, with large heads, bidentate. Proximal teeth about as large as distal teeth, sharply directed laterally. Distal teeth flattened, nearly geniculate and directed laterally. Guards asymmetrically bluntly pointed, marginally smooth; mucros absent. Pseudocompound falcigers and compound spinigers absent. Aciculae numbering 1 to 3, dark, tapering, straight; cross-sections round. Notopodial aciculae paired, dark. Subacicular hooks (Figure 53j) dark, bidentate. Hooks first present from setiger 32, present in all setigers thereafter, paired in some setigers. Hooks smoothly tapering to small head. Proximal teeth larger than distal teeth; both teeth directed obliquely distally. Guards short and blunt.

UNKNOWN MORPHOLOGICAL FEATURES.—Jaw structure; pygidium and anal cirri.

EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—Mx III short and forming part of distal are with left Mx IV.

CHARACTERS USED IN PREPARATION OF KEY NOT SCORED.—Inappropriate Characters: 22, 56, 60. Unknown Characters: 13, 14, 39, 40, 42, 74, 78.

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—None.

MATERIAL EXAMINED.—One specimen, MNHN, Paris A.1(R.)-1868-no. 46a. Iles Chausey, Quatrefages.

COMMENTS ON MATERIAL EXAMINED.—The type has been lost; justification for using the current specimen to represent the species is given below. The jaws are half everted and the maxillary formula given is incomplete because detailing the formula would destroy the jaws completely.

DESCRIPTION.—Specimen incomplete, of unknown sex, with 63 setigers; length 22 mm; maximal width 2.5 mm at setiger 10; length through setiger 10, 5 mm. Body cylindrical.

Prostomium less than as thick as peristomium, narrow. Prostomium (Figure 54a) distinctly shorter and narrower than peristomium, less than as deep as peristomium. Prostomial lobes frontally obliquely truncate, dorsally flattened; median sulcus deep. Eyes at bases of A-I, faded. Antennae in shallow horseshoe, evenly spaced, similar in thickness. Ceratophores ring-shaped in all antennae, without articulations. Ceratostyles tapering, with up to 5 indistinct, cylindrical articulations in A-III. A-I to posterior peristomial ring; A-II to setiger 2; A-III to setiger 4. Peristomium cylindrical. Separation between rings distinct on all sides; anterior ring of total peristomial length. Peristomial cirri to middle of anterior peristomial ring, slender and tapering, without articulations.

Incomplete maxillary formula 1+1, ?+?, 6+0, 3+8, 1+1. Mx VI missing. Mx III long, but part of distal are with left Mx IV.

Branchiae (Figure 54d) present, pectinate, distinctly longer than notopodial cirri, not reduced in mid-body region, erect. Branchiae from setiger 4 to end of fragment. First 3 branchiae single filaments; all others pectinate with maximally 10 filaments at about setiger 15; last branchia present, on setiger 63, with 2 filaments only. Stems tapering, shorter than notopodial cirri. Filaments long, digitiform, as long as notopodial cirri where best developed.

Anterior neuropodial acicular lobes truncate; posterior acicular lobes triangular; aciculae emerging at midline. Pre-and postsetal lobes low, transverse folds. First 5 ventral cirri thick, tapering. Ventral cirri basally inflated from setiger 6 through about setiger 35; inflated base ovate; narrow tips tapering. Posterior ventral cirri broadly transversely attached; enfolding lower edge of parapodia, nearly scoop-shaped. Anterior notopodial cirri basally slightly inflated, retaining same size in all setigers, but less distinctly inflated in posterior setigers, without articulations.

Limbate setae slender, marginally smooth. Pectinate setae tapering, flat. One marginal tooth longer than other teeth, with ~12 teeth. Shafts of compound falcigers (Figure 54c) distally tapering, marginally smooth. Appendages relatively short, tapering with large, distinct heads, bidentate. Proximal teeth as large as distal teeth, triangular, directed laterally. Distal teeth distinctly bent, sharply tapering. Guards asymmetrically bluntly pointed, marginally smooth; mucros absent. Pseudocompound falcigers and compound spinigers absent. Aciculae paired; anterior aciculae honey colored, becoming darker brown in posterior setigers, distally tapering to sharp points, straight; cross-sections round. Separation between core and sheath indistinct in both aciculae and subacicular hooks. Subacicular hooks (Figure 54b) light brown, bidentate. Hooks first present from setiger 28, present in all setigers thereafter, always single (except for replacements). Hooks with distinct neck. Proximal teeth large, curved, directed laterally. Distal teeth bluntly triangular, nearly erect.

UNKNOWN MORPHOLOGICAL FEATURES.—Pygidium and anal cirri.

EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—None.

CHARACTERS USED IN PREPARATION OF KEY NOT SCORED.—Inappropriate Characters: 56, 58, 59. Unknown Characters: 1,2, 36–38,40.

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—37,1; 38,1.