Neoserica

Body length: 10.7 mm, length of elytra: 7.8 mm, width: 6.1 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, nearly glabrous. Labroclypeus subrectangular, little wider than long, widest at base, lateral margins straight and weakly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins strongly reflexed; surface flat and shiny, basis with dull toment, punctation dense, behind the anterior margin with coarser punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a few minute and superficial punctures, with a long terminal seta. Frons dull, in posterior half weakly shiny, with fine and sparse punctures, beside the eyes and behind the frontoclypeal suture with a few erect setae. Eyes large, ratio diameter/interocular width: 0.75. Antenna with ten antennomeres, club with six antennomeres, strongly reflexed, twice as long as remaining antennomeres combined; antennomere 5 subequal to half of length of club, antennomere 4 strongly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum weakly produced medially, with a moderate median sinuation. Pronotum transverse, subrectangular, widest at base, lateral margins in basal half straight and nearly subparallel, evenly convex and strongly convergent in anterior half, anterior angles moderately sharp and moderately produced, posterior angles blunt, slightly rounded at the tip; anterior margin strongly convexly produced medially, with a medially incomplete marginal line; surface densely and finely punctate, with minute setae in punctures; setae of lateral border fine and sparse; hypomeron basally distinctly carinate, but carina not produced. Scutellum moderately wide and long, with fine, moderately dense punctures, smooth on basal midline, with only minute setae. Elytra oblong, widest in posterior third, striae weakly impressed, finely and moderately densely punctate, intervals weakly convex with punctures concentrated along the striae, odd intervals with a few fine, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura densely setose, apical border narrowly membraneous, with a fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and unevenly and not densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium weakly convex and dull, finely and densely punctate, without a smooth midline, with a few longer setae on posterior half. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, punctures and setae of anterior longitudinal row completely reduced, posterior margin in apical half ventrally smooth and not widened but slightly serrate in apical quarter, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/5.1, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length, but not very distinct; ventral edge finely serrated, with three robust setae of which the two distal ones are widely separated; medial face smooth, apex moderately distinctly concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few fine punctures; metatarsomeres with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 3I–K. Female unknown.

Body length: 14.4 mm, length of elytra: 10.3 mm, width: 6.9 mm. Body oblong, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.53. Antenna with ten antennomeres, club with six antennomeres, straight, 1.2 times as long as the remaining antennomeres combined; antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins weakly convex, convexly bent at middle, in basal half subparallel, in apical half strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with convex sides and with the apex slightly rounded, with fine, moderately dense punctures, with only minute setae. Elytra oblong, apex slightly truncate, widest at middle, striae weakly impressed, finely and moderately densely punctate, intervals nearly flat, with moderately dense evenly spaced, fine punctures, intervals with a few fine white setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.79. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few semi-erect setae on apical half. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.7, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures dorsally; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 1E–G. Female unknown.

Body length: 11.8 mm, length of elytra: 8.8 mm, width: 6.8 mm. Body oblong, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, densely covered with minute erect setae. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation moderately dense and superficial, punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye nearly as wide as long; ocular canthus short (length =1/4 of ocular diameter) and wide, densely and minutely setose. Frons dull, with fine and dense but superficial punctures each bearing a short to minute seta, beside eyes and behind the frontoclypeal suture with a few longer erect setae. Eyes small, ratio diameter/interocular width: 0.46. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomere 5 two thirds as long as the club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin weakly convex medially, marginal line widely interrupted medially; surface densely and finely punctate, with short to minute erect setae in punctures; setae of anterior and lateral border absent; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, subtriangular, with fine, moderately dense punctures, impunctate on basal midline, with minute erect setae in punctures. Elytra oblong, widest at posterior third, striae finely impressed, finely and moderately densely punctate, intervals nearly flat, with moderately dense evenly spaced, fine punctures each bearing a short fine erect setae, odd intervals with a few fine longer setae; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, densely covered with short setae; metacoxa glabrous, with a very few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, densely and finely setose, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, densely and finely setose, setae on apical half longer. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, anterior longitudinal row complete, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, impunctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 7E–G. Female unknown.

Lectotype. Body length: 13 mm, length of elytra: 9.6 mm, width: 7.2 mm. Body oblong, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.49. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than the remaining antennomeres combined; antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with convex sides and with the apex slightly rounded, with fine, moderately dense punctures, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, intervals nearly flat, with moderately dense evenly spaced, fine punctures, intervals with a few fine white setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.79. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few semi-erect setae beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.5, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 1A–C.

Body length: 12.3 mm, length of elytra: 9.6 mm, width: 7.4 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly sinuate medially, margins moderately reflexed; surface slightly convex and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.49. Antenna with ten antennomeres, club with six antennomeres, straight, 1.5 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 distinctly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, distinctly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals narrower and distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Pygidium weakly convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with a few semi-erect setae basally on sides, at apex with short, fine, moderately dense setae. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 1I–K.

Body length: 13.0 mm, length of elytra: 9.8 mm, width: 7.8 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface slightly convex and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and moderately long (length = 1/3 of ocular diameter), glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside the eyes and the frontoclypeal suture with a few erect setae. Eyes small, ratio diameter/interocular width: 0.48. Antenna with ten antennomeres, club with six antennomeres, straight, 1.2 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, strongly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, basally impunctate at middle, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals narrower and distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.77. Pygidium moderately convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with a few setae beside the margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.7, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, mesotarsomeres with a few very fine punctures; metatarsomeres lacking in holotype. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 2A–D. Female unknown.

Body length: 12.0 mm, length of elytra: 8.9 mm, width: 6.6 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly sinuate medially, margins moderately reflexed; surface slightly convex and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.52. Antenna with ten antennomeres, club with six antennomeres, straight, 1.2 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 distinctly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex in the basal half, in anterior third nearly straight, throughout strongly convergent anteriorly, anterior angles very sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals narrower and distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7. Pygidium weakly convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with a few semi-erect setae basally on sides, at apex with short, fine, moderately dense setae. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 2F–H. Female unknown.

Body length: 13.0 mm, length of elytra: 10.0 mm, width: 8.0 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface slightly convex and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and moderately long (length = 1/3 of ocular diameter), glabrous, with a robust terminal seta. Frons dull, with fine and dense punctures, beside the eyes and behind the frontoclypeal suture with a few erect setae. Eyes small, ratio diameter/interocular width: 0.48. Antenna with ten antennomeres, club with six antennomeres, straight, 1.1 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border abrased in holotype; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, basally impunctate at middle, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals narrower and distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura only sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.73. Pygidium moderately convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with numerous long setae on apex, otherwise with minute setae in punctures. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.8, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 2J–N.

Body length: 14.0 mm, length of elytra: 10.5 mm, width: 8.2 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins nearly straight and convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly sinuate medially, margins moderately reflexed; surface slightly convex and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and moderately long (length = 1/3 of ocular diameter), glabrous, with one or two robust terminal setae. Frons dull, with fine and dense punctures, beside the eyes and behind the frontoclypeal suture with a few erect setae. Eyes small, ratio diameter/interocular width: 0.47. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomere 5 distinctly shorter than the club, antennomere 4 slightly widened but not transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest just before base, lateral margins evenly convex and convergent anteriorly, slightly convergent also towards the strongly rounded posterior angles, anterior angles sharp and distinctly produced; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals narrower and slightly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura only sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.84. Pygidium apically strongly convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with numerous long setae on apex, otherwise with minute setae in punctures. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 3A–C. Female unknown.

Body length: 11 mm, length of elytra: 7.6 mm, width: 5.2 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, nearly glabrous. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles weakly rounded, anterior margin weakly sinuate medially, margins strongly reflexed; surface flat and shiny, basis without dull toment, punctation dense, behind the anterior margin with coarser punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus long and slender, glabrous, with a few minute and superficial punctures, with a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes and behind the frontoclypeal suture with a few erect setae. Eyes large, ratio diameter/interocular width: 0.69. Antenna with ten antennomeres, club with six antennomeres, moderately reflexed, twice as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 strongly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum weakly produced medially, with a moderate median sinuation. Pronotum transverse, subrectangular, widest at base, lateral margins in basal half straight and nearly subparallel, evenly convex and strongly convergent in anterior half, anterior angles sharp and strongly produced, posterior angles right-angled, slightly rounded at the tip; anterior margin strongly convexly produced medially, marginal line incomplete medially; surface densely and finely punctate, with minute setae in punctures; setae of lateral border nearly absent; hypomeron basally distinctly carinate, but carina not produced. Scutellum moderately wide and short, with fine, moderately dense punctures, smooth on basal midline, with only minute setae. Elytra oblong, widest shortly behind middle, striae weakly impressed, finely and moderately densely punctate, intervals weakly convex with punctures concentrated along the striae, odd intervals with a few fine, white, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura densely setose, apical border narrowly membraneous, with a fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and unevenly and not densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.47. Pygidium weakly convex and dull, finely and densely punctate, without a smooth midline, with a few longer setae on posterior half. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, punctures and setae of anterior longitudinal row completely reduced, posterior margin in apical half ventrally smooth and not widened but slightly serrate in apical quarter, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/4.3, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length, but not very distinct; ventral edge finely serrated, with three robust setae of which the two distal ones are widely separated; medial face smooth, apex moderately distinctly concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few fine punctures; metatarsomeres with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 3E–G. Female unknown.

Body length: 11 mm, length of elytra: 7.6 mm, width: 5.2 mm. Body oblong, dark brown, ventral surface and antennal yellowish brown, anterior labroclypeus shiny, dorsal surface moderately shiny, frons dull, with numerous long, yellow, semi-erect setae. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins strongly reflexed; surface flat and shiny, basis without dull toment, finely and densely punctate, with numerous coarser punctures interspersed each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus long and slender, impunctate and glabrous, with a long terminal seta. Frons dull, finely and densely punctate, with numerous coarser punctures interspersed each bearing a long erect seta. Eyes moderately large, ratio diameter/interocular width: 0.69. Antenna with ten antennomeres, club with five antennomeres, straight, as long as remaining antennomeres combined; antennomere 6 subequal to two thirds of length of club, antennomere 5 transverse. Mentum elevated and slightly flattened anteriorly. Labrum weakly produced medially, with a moderate median sinuation. Pronotum transverse, subrectangular, widest just before base, lateral margins evenly convex and strongly convergent towards sharp and moderately produced anterior angles, posterior angles blunt, rounded at the tip; anterior margin strongly convexly produced medially, marginal line robust and complete medially; surface densely and finely punctate, with coarse punctures irregularly interspersed each bearing a long semi-erect seta, fine punctures minutely setose; setae of lateral and anterior border dense; hypomeron basally distinctly carinate, but carina not produced. Scutellum moderately wide and short, with fine, dense punctures, widely smooth on basal midline, with only minute setae. Elytra oblong, widest at middle, striae weakly impressed, finely and moderately densely punctate, intervals weakly convex with fine and moderately dense punctures concentrated along striae, all intervals with sparse semi-erect long setae, otherwise punctures only with very minute setae; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura densely setose, apical border chitinous, without fringe of microtrichomes (visible at 100× magnification). Ventral surface shiny, coarsely densely punctate and finely setose; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and densely punctuate, with minute setae in punctures, each sternite with a transverse row of coarse punctures each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.47. Pygidium weakly convex and moderately shiny, coarsely and densely punctate, on basal portion also with finer punctures interspersed, without a smooth midline, with dense, long setae. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur shiny, anterior margin acute, behind anterior edge without serrated line, punctures and setae of anterior longitudinal row complete, posterior margin in apical half ventrally smooth and neither widened nor serrate in apical quarter, posterior margin dorsally distinctly serrated, on its basal portion with a very few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/4.3, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, finely and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust setae of which the two distal ones are widely separated; medial face smooth, apex moderately distinctly concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few fine punctures; metatarsomeres with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 7I–K. Female unknown.

Body length: 7.6 mm, length of elytra: 5.2 mm, width: 4.0 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 4L). Labroclypeus subtrapezoidal, widest at base; lateral margins convex and convergent towards moderately rounded anterior angles; anterior margin distinctly sinuate medially; margins moderately reflexed; surface weakly convex medially, coarsely and finely but densely punctate, densely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 1.5 times as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with a few long seta. Frons with coarse and dense punctures mixed with sparse, fine ones, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.71. Antenna with ten antennomeres, club with four antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex and moderately convergent towards moderately produced and sharp anterior angles; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with dense, long setae; hypomeron carinate at base. Scutellum small, with fine, dense punctures and dense, fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals flat, odd ones slightly convex; intervals with fine, dense punctures, punctures on odd intervals concentrated along striae, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.74. Pygidium moderately convex and shiny, finely and densely punctate, without smooth midline; with dense, long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with sparse, fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/3.33; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; basally with a few strong short single setae in coarse puncture with serrated borders; lateral face moderately densely and coarsely punctate, shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4I–K.

Body length: 7.2 mm, length of elytra: 5.3 mm, width: 4.3 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 1H). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards moderately rounded anterior angles; anterior margin shallowly sinuate medially; margins moderately reflexed; surface moderately elevated medially, coarsely and densely punctate, densely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 2.5 times as wide as long. Ocular canthus moderately long, impunctate, with one or two single setae. Frons with coarse and moderately dense punctures, with dense setae being bent posteriorly. Eyes moderately large, ratio diameter/interocular width: 0.6. Antenna with ten antennomeres, club with four antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex and convergent anteriorly; anterior angles distinctly produced and sharp; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae, on basal midline punctures less dense. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.35. Pygidium moderately convex and shiny, finely and densely punctate, without smooth midline; shortly and densely setose, with sparse long and erect setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/3.2; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; basally with a few strong short single setae; lateral face densely and coarsely punctate, sparsely and shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Female unknown. Aedeagus: Fig. 1E–G.

Body length: 7.5 mm, length of elytra: 5.0 mm, width: 3.8 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 1L). Labroclypeus short and subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards strongly rounded anterior angles; anterior margin shallowly sinuate medially; margins moderately reflexed; surface moderately elevated medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 2.5 times as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with one or two single setae. Frons with coarse and moderately dense punctures, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.76. Antenna with ten antennomeres, club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles moderately produced and sharp; posterior angles blunt, rounded at tip; anterior margin with robust, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.59. Pygidium moderately convex and shiny, finely and densely punctate, without smooth midline; with dense, moderately long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with dense and thick, evenly long setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/3.0; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a continuously serrated line and some single punctures each bearing a short seta; lateral face densely and coarsely punctate, densely and shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 1I–K.

Body length: 7.2 mm, length of elytra: 5.2 mm, width: 3.9 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (pilosity partly abraded; Fig. 2D). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards moderately rounded anterior angles; anterior margin shallowly sinuate medially; margins moderately reflexed; surface moderately elevated medially, coarsely and finely, densely punctate, densely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye three times as wide as long. Ocular canthus moderately long, impunctate, with two or three setae. Frons with coarse and dense punctures, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.75. Antenna with ten antennomeres, club with four antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins weakly convex and convergent anteriorly; anterior angles distinctly produced and moderately sharp; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.73. Pygidium strongly convex and shiny, finely and densely punctate, without smooth midline; shortly and densely setose, with sparse long and erect setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened. Metatibia wide and moderately long, widest at middle; ratio of width/length: 1/2.7; dorsal margin sharply carinate, with two groups of spines; basal group at middle, apical group at three quarters of metatibial length; in basal half with a few strong and short single setae in coarse punctures beside a undulated serrated line; lateral face densely and coarsely punctate, densely and shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Meso- and metatarsomeres of holotype also missing. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Female unknown. Aedeagus: Fig. 2A–C.

Body length: 7.5 mm, length of elytra: 5.7 mm, width: 4.2 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 2H). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards strongly rounded anterior angles; anterior margin distinctly sinuate medially; margins moderately reflexed; surface moderately elevated medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 2.5 times as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with one or two single setae. Frons with coarse and moderately dense punctures, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.73. Antenna with ten antennomeres, club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles moderately produced and sharp; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.62. Pygidium weakly convex and shiny, finely and densely punctate, without smooth midline; with dense, long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/2.7; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a continuously serrated line and some single punctures each bearing a short seta; lateral face densely and coarsely punctate, densely and shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; metatarsomeres 2-5 and dorsal tibial spur of holotype also missing. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 2E–G.

Body length: 6.7 mm, length of elytra: 4.8 mm, width: 3.8 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 3D). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards strongly rounded anterior angles; anterior margin distinctly sinuate medially; margins moderately reflexed; surface moderately elevated medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye twice as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with two long setae. Frons with coarse and moderately dense punctures, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.74. Antenna with ten antennomeres, club with four antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles moderately produced and sharp; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, around midline punctures very dense, partly fusing with each other transversely, densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.57. Pygidium weakly convex and shiny, finely and densely punctate, without smooth midline; with dense, long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/2.7; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a undulated, nearly continuously serrated line and beside it some single punctures each bearing a short seta; lateral face moderately densely and coarsely punctate, shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 3A–C.

Body length: 7.3 mm, length of elytra: 5.2 mm, width: 3.9 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine and short, semi-erect setae (in part abraded; Fig. 3H). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards moderately rounded anterior angles; anterior margin shallowly sinuate medially; margins moderately reflexed; surface convexly elevated medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 2.5 times as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with a single short terminal seta. Frons with coarse and sparse punctures, with numerous setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.78. Antenna with ten antennomeres, club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins nearly straight and convergent, slightly convex anteriorly and moderately convergent towards moderately produced and sharp anterior angles; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, except on disc (probably abraded) densely setose; anterior and lateral borders with sparse but longer setae; hypomeron carinate at base. Scutellum with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals nearly flat, odd ones slightly convex; intervals with fine, dense punctures, punctures on odd intervals concentrated along striae, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.57. Pygidium strongly convex and shiny, finely and densely punctate, without smooth midline; with dense, long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/2.7; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a undulated, nearly continuously serrated line and beside it single coarse punctures each bearing a short robust seta; lateral face moderately densely and coarsely punctate, shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres missing in holo- and paratype. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 3E–G.

Body length: 6.2 mm, length of elytra: 4.5 mm, width: 3.2 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, densely covered with fine, semi-erect setae (Fig. 4D). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards moderately rounded anterior angles; anterior margin shallowly sinuate medially; margins moderately reflexed; surface moderately convex medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye 2.5 times as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, with a single short terminal seta. Frons with coarse and dense punctures, with dense setae being bent posteriorly. Eyes large, ratio diameter/interocular width: 0.71. Antenna with ten antennomeres, club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at posterior third, lateral margins evenly convex, moderately convergent posteriorly and towards moderately produced and sharp anterior angles; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, densely setose; anterior and lateral borders with dense, long setae; hypomeron carinate at base. Scutellum small, with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals flat, odd ones slightly convex; intervals with fine, dense punctures, punctures on odd intervals concentrated along striae, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.55. Pygidium moderately convex and shiny, finely and densely punctate, without smooth midline; with dense, long setae on disc and beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with sparse, fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/3.0; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a slightly undulated, nearly continuously serrated line and beside it single coarse punctures each bearing a short robust seta; lateral face moderately densely and coarsely punctate, shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres with a strongly serrated ventral ridge; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Female unknown. Aedeagus: Fig. 4A–C.

Body length: 6.6 mm, length of elytra: 5.1 mm, width: 4.0 mm. Body oblong, reddish brown, antennal club yellowish brown, dorsal surface shiny, elytra densely covered with fine, semi-erect setae; setae on head and pronotum abraded in type specimens (Fig. 4H). Labroclypeus subtrapezoidal, widest at base; lateral margins weakly convex and moderately convergent towards moderately rounded anterior angles; anterior margin distinctly sinuate medially; margins moderately reflexed; surface moderately convex medially, coarsely and finely but densely punctate, sparsely setose. Frontoclypeal suture finely incised, weakly elevated and moderately angled medially. Smooth area anterior to eye twice as wide as long. Ocular canthus narrow and moderately long, sparsely punctate, glabrous. Frons with coarse and dense punctures, with a few moderately long setae beside eyes. Eyes moderately large, ratio diameter/interocular width: 0.68. Antenna with ten antennomeres, club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum short and almost straight anteriorly, with a transverse rim of very dense, short and robust setae. Pronotum widest at base, lateral margins evenly convex, moderately convergent towards moderately produced and sharp anterior angles; posterior angles blunt, rounded at tip; anterior margin with fine, complete marginal line, weakly produced medially; surface densely and finely punctate, sparsely setose; anterior and lateral borders with sparse, long setae; hypomeron carinate at base. Scutellum small, with fine, dense punctures and a few fine setae. Elytra oblong, widest behind middle, striae weakly impressed, finely and densely punctate; intervals flat, odd ones slightly convex; intervals with fine, dense punctures, punctures on odd intervals concentrated along striae, densely covered with fine, moderately long setae. Epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border with a wide membranous rim of microtrichomes (visible at magnification 100×). Ventral surface shiny, finely and densely punctate. Metasternum with short, fine setae. Metacoxa glabrous, with a few single setae laterally. Abdominal sternites finely and densely punctate, finely setose, with a transverse row of coarse punctures each bearing a long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.61. Pygidium moderately convex and shiny, shortly and densely punctate, without smooth midline; with dense, long setae beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and densely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior margin of metafemur smooth, dorsally and ventrally, in apical half moderately widened, dorsal posterior margin with sparse, fine setae. Metatibia wide and moderately long, widest at two thirds of metatibial length; ratio of width/length: 1/2.85; dorsal margin sharply carinate, with two groups of spines; basal group shortly behind middle of metatibial length, apical group at three quarters of metatibial length; in basal half with a slightly undulated, nearly continuously serrated line and beside it single coarse punctures each bearing a short robust seta; lateral face moderately densely and coarsely punctate, shortly setose; ventral edge finely serrated, with four robust equidistant setae; medial face impunctate; apex weakly truncate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae; not carinate laterally, impunctate dorsally; metatarsomeres missing in holotype. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Female unknown. Aedeagus: Fig. 4E–G.

Body length: 13.5 mm, length of elytra: 10.0 mm, width: 8.3 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.5. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than the remaining antennomeres combined; antennomere 5 subequal to two thirds of length of club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins straight in the basal half and slightly weakly convergent anteriorly, convex and strongly convergent in anterior half, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum wide and moderately long, triangular with nearly straight sides, apex slightly rounded, with fine, moderately dense punctures, basally smooth at middle, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals slightly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border narrowly membraneous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.73. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few semi-erect setae basally on sides and along the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and moderately long, widest at apex, ratio of width/length: 1/3.4, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5A–D.

Body length: 12.2 mm, length of elytra: 8.8 mm, width: 7.3 mm. Body oblong, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, partially dull toment lost and moderately shiny, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with a transverse row of coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.53. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomere 5 half as long as the club, antennomere 6 three quarters as long as club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately long, triangular with convex sides and with the apex slightly rounded, with fine, moderately dense punctures, with only minute setae. Elytra oblong, widest shortly behind middle, striae weakly impressed, finely and moderately densely punctate, intervals nearly flat, with moderately dense evenly spaced, fine punctures, intervals with a few fine white setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.59. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few semi-erect setae beside the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/4.0, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5I–K.

Body length: 11.7 mm, length of elytra: 8.1 mm, width: 6.6 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, nearly glabrous. Labroclypeus subrectangular, little wider than long, widest at base, lateral margins straight and weakly convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly sinuate medially, margins strongly reflexed; surface flat and shiny, basis with dull toment, punctation dense, behind the anterior margin with coarser punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a few minute and superficial punctures, with a long terminal seta. Frons dull, in posterior half weakly shiny, with fine and sparse punctures, beside the eyes and behind the frontoclypeal suture with a few erect setae. Eyes large, ratio diameter/interocular width: 0.76. Antenna with ten antennomeres, club with six antennomeres, moderately reflexed, 1.5 times as long as remaining antennomeres combined; antennomere 5 subequal to half of length of club, antennomere 4 moderately transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum weakly produced medially, with a moderate median sinuation. Pronotum transverse, subrectangular, widest just before base, lateral margins in basal half straight and nearly subparallel, slightly narrowed towards the base, evenly convex and strongly convergent in anterior half, anterior angles moderately sharp and moderately produced, posterior angles blunt, slightly rounded at the tip; anterior margin strongly convexly produced medially, with a medially incomplete marginal line; surface densely and finely punctate, with minute setae in punctures; setae of lateral border fine and sparse; hypomeron basally distinctly carinate, but carina not produced. Scutellum moderately wide and long, with fine, moderately dense punctures, smooth on basal midline, with only minute setae. Elytra oblong, widest in posterior third, striae weakly impressed, finely and moderately densely punctate, intervals weakly convex with punctures concentrated along the striae, odd intervals with fine, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura densely setose, apical border narrowly membraneous, with a fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and unevenly and not densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium weakly convex and dull, finely and densely punctate, without a smooth midline, with a few longer setae between the minute setae. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, punctures and setae of anterior longitudinal row completely reduced, posterior margin in apical half ventrally smooth and not widened and smooth in apical quarter, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/5.1, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length, but not very distinct; ventral edge finely serrated, with three robust setae of which the two distal ones are widely separated; medial face smooth, apex moderately distinctly concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with dense, fine punctures; metatarsomeres with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 7A–C. Female unknown.

Body length: 12.7 mm, length of elytra: 9.2 mm, width: 7.6 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.52. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than the remaining antennomeres combined; antennomere 5 subequal to two thirds of length of club, antennomere 4 slightly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex, in the basal half only weakly convergent, strongly convergent in anterior half, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum wide and moderately long, triangular with nearly straight sides, apex slightly rounded, with fine, moderately dense punctures, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals slightly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.9. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few semi-erect setae basally on sides, at apex with short, fine, very dense setae. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and moderately long, widest at apex, ratio of width/length: 1/3.3, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6A–C.

Body length: 13.0 mm, length of elytra: 9.9 mm, width: 7.7 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.64. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than the remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 strongly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and moderately convergent anteriorly, in anterior half more strongly convergent, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately wide and long, triangular with nearly straight sides, apex slightly rounded, with fine, moderately dense punctures, smooth along the middle, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals slightly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border narrowly membraneous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few short setae on sides and along the apical margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and moderately long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5E–G.

Body length: 12.0 mm, length of elytra: 9.6 mm, width: 7.6 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and moderately convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface convexly elevated at middle and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and long (length = 1/3 of ocular diameter), glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside the eyes a with a few erect setae. Eyes small, ratio diameter/interocular width: 0.48. Antenna with ten antennomeres, club with seven antennomeres, straight, 1.2 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 more than half as long as the club, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, strongly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, impunctate along the middle, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.77. Pygidium moderately convex and dull, densely punctate, fine punctures mixed with coarser ones, without smooth midline, with a few setae beside apical margin. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/3.7, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6E–H.

Body length: 10.6 mm, length of elytra: 7.8 mm, width: 6.1 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus slightly subrectangular, distinctly wider than long, widest at base, lateral margins nearly straight and weakly convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface convexly elevated at middle and shiny, basis with dull toment, punctation moderately dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and long (length = 1/3 of ocular diameter), glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside the eyes a with a few erect setae. Eyes small, ratio diameter/interocular width: 0.48. Antenna with ten antennomeres, club with seven antennomeres, straight, nearly twice as long as remaining antennomeres combined; antennomere 5 equal to length of club, antennomere 4 subequal to three quarter of length of club, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, strongly rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, with nearly invisible fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.77. Pygidium weakly convex and dull, densely punctate, fine punctures mixed with coarser ones, with a narrow smooth midline, nearly glabrous, only with minute setae in punctures. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia moderately slender and long, widest at apex, ratio of width/length: 1/3.7, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4I–K.

Body length: 14.5 mm, length of elytra: 10.7 mm, width: 8.3 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes small, ratio diameter/interocular width: 0.48. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomere 5 subequal to half of length of club, antennomere 4 strongly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and moderately convergent anteriorly, in anterior half more strongly convergent, anterior angles sharp and distinctly produced, posterior angles blunt, slightly rounded at the tip; anterior margin nearly straight, with a distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum moderately wide and long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, smooth along the middle, with only minute setae. Elytra oblong, apex slightly truncate and slightly concavely sinuated before the apical angle, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals slightly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the blunt external apical angle of elytra, epipleura only sparsely setose, apical border narrowly membraneous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.54. Pygidium weakly convex and dull, coarsely and densely punctate, with a very narrow smooth midline, with a few longer setae on sides and along the apical margin, otherwise only with minute setae in punctures. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row with very sparse and short setae only, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and moderately long, widest at apex, ratio of width/length: 1/4.0, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with four robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4E–G. Female unknown.

Body length: 11.8 mm, length of elytra: 8.2 mm, width: 6.5 mm. Body oblong, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, opaque toment on elytra and pronotum less thick, with a light trace of shine, sparsely setose. Labroclypeus subrectangular, distinctly wider than long, widest at base, lateral margins moderately convex and very little convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface convexly elevated at middle and shiny, basis with dull toment, punctation moderately dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture distinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately wide and long (length = 1/3 of ocular diameter), glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside the eyes a with a few erect setae, otherwise with minute setae only. Eyes small, ratio diameter/interocular width: 0.5. Antenna with ten antennomeres, club with seven antennomeres, straight, 1.2 times as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 0.7 times as long as the club, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a moderate median sinuation. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins weakly convex and in basal half weakly convergent anteriorly, at middle more convex and in anterior half weakly convex and strongly convergent, anterior angles moderately sharp and distinctly produced, posterior angles blunt, moderately rounded at the tip; anterior margin nearly straight, with a fine and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of lateral border sparse; hypomeron basally distinctly carinate, but carina only weakly produced. Scutellum long, triangular with nearly straight sides, apex slightly rounded, with fine, dense punctures, impunctate along the middle, with only minute setae. Elytra oblong, widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, odd intervals distinctly convex with punctures concentrated along the striae, others evenly punctate and nearly flat, odd intervals with white scale-like, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, very narrow behind the middle, ending at the moderately rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, with only a very fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short setae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.8. Pygidium moderately convex and dull, densely punctate, fine punctures mixed with coarser ones, with a narrow smooth midline, with a few longer setae on disc and beside the apical margin. Legs moderately slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row nearly completely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia moderately slender and long, widest at apex, ratio of width/length: 1/3.2, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6J–L.

Body length: 10.5 mm, length of elytra: 7.5 mm, width: 4.9 mm. Body oblong and slender, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, nearly glabrous. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins straight and moderately convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, margins strongly reflexed; surface flat and shiny, basis with dull toment, punctation dense, behind anterior margin with a transversal row of coarser punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a few minute and superficial punctures and a long terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with a few erect setae. Eyes large, ratio diameter/interocular width: 0.92. Antenna with ten antennomeres, club with six antennomeres, strongly reflexed, twice as long as remaining antennomeres combined; antennomere 5 subequal to length of club, antennomere 4 strongly transverse, antennomere 3 half as long as pedicellus. Mentum elevated and slightly flattened anteriorly. Labrum weakly produced medially, with a moderate median sinuation. Pronotum transverse, subrectangular, widest at base, lateral margins in basal half straight and nearly subparallel and slightly concavely sinuate, evenly convex and strongly convergent in anterior half, anterior angles sharp and distinctly produced, posterior angles right-angled, slightly rounded at the tip; anterior margin strongly convexly produced medially, with a medially incomplete marginal line; surface densely and finely punctate, with minute setae in punctures; setae of lateral border fine and sparse; hypomeron basally distinctly carinate, but carina not produced. Scutellum moderately wide and long, with fine, moderately dense punctures, smooth on basal midline, with only minute setae. Elytra oblong and very slender, widest just before posterior third, striae weakly impressed, finely and moderately densely punctate, intervals weakly convex with punctures concentrated along the striae, odd intervals with a few fine, adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura densely setose, apical border narrowly membraneous, with a fine fringe of microtrichomes (visible at 100× magnification). Ventral surface dull, coarsely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few short setae laterally, posterior margin straight; abdominal sternites finely and unevenly and not densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.38. Pygidium weakly convex and dull, finely and densely punctate, without a smooth midline, with a few longer setae on posterior half. Legs slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, punctures and setae of anterior longitudinal row completely reduced, posterior margin in apical half ventrally smooth and not widened but slightly serrate in apical quarter, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ratio of width/length: 1/4.7, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length, but nearly longitudinally convex and very indistinct; ventral edge finely serrated, with three robust setae of which the two distal ones are widely separated; medial face smooth, apex moderately distinctly concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few fine punctures; metatarsomeres with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4A–C. Female unknown.

Holotype: ♂ “N-Thailand 25.-29.5.1990 Doi Inthanon leg. Malicky” (ZSMC). Paratype. 1 ♂ “N-Thailand 27.III.1990 Doi Inthang [sic, = Doi Inthanon] lg. Malicky” (ZFMK), 1 ♂ “N. Thailand Doi Pui, near Chiang Mai; 09.V.1985; leg. H. Nara” (ZFMK).

Holotype: ♂ “N-Vietnam Tam Dao, Vinh Phu Prov. 21°27'18"N, 105°38'58"E 1050–1200m 2.–6.VI.1999 leg. Fabrizi, Jäger, Ahrens” (ZFMK). Paratypes: 1 ♀ “N-Vietnam Tam Dao, Vinh Phu Prov. 21°27'18"N, 105°38'58"E 1050–1200m 2.–6.VI.1999 leg. Fabrizi, Jäger, Ahrens” (ZFMK), 1 ♂ “Vietnam N. 15.5.–16.6. 75 km NW from Hanoi Tam Dao E. Jendek leg.” (CA), 1 ♂ “N. Vietnam/ Tonkin/ Tamdao/ pr. Vinhphu/ 2.–11.6.1985 Vit Kubáň leg./ VS 50” (CPPB).

Holotype ♂ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500-1900m, 17.5.-3.6.2007, leg. C. Holzschuh” (ZFMK). Paratypes: 2 ♂♂ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500-1900m, 17.5.-3.6.2007, leg. Vit Kubáň” (NMPC), 1 ♂, 1 ♀ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500-1900m, 17.5.-3.6.2007, leg. C. Holzschuh” (ZFMK).

Holotype: ♂ “Myanmar N (Burma) 25km E Putao, H-800m Nan Sa Bon vill. 06–09.05.1998 leg. S. Murzin & V. Siniaev/ Coll. Takeshi Itoh Osaka (Japan)” (ZFMK). Paratype: 1 ♀ “Myanmar N (Burma) 25km E Putao, H-800m Nan Sa Bon vill. 06–09.05.1998 leg. S. Murzin & V. Siniaev/ Coll. Takeshi Itoh Osaka (Japan)” (ZFMK).

Holotype: ♂ “China, Yunnan prov.; Gaoligongshan mts.; 90km W of Baoshan; S. Bečvář leg.; 26–29.v.1995/ Coll. P. Pacholátko Invt. No.” (CPPB). Paratypes: 5 ♂♂ “China, Yunnan prov.; Gaoligongshan mts.; 90km W of Baoshan; S. Bečvář leg.; 26–29.v.1995/ Coll. P. Pacholátko Invt. No.” (ZFMK), 32 ♂♂ “China, W Yunnan prov., mts. 60Km E Tengchong, 2300m 14.–19.v.2006, S. Murzin & I. Shokhin leg.” (CP, ZFMK).

Holotype: ♂ “Yingjiang, Yunnan, 13.IV.1980, 1300m, leg. Li Hongxing” (IZAS). Paratypes: 1 ♂ “Yingjiang, Yunnan, 13.IV.1980, 1700m, leg. Gao Ping” (ZFMK), 1 ♂ “Yingjiang, Yunnan, 13.IV.1980, 300m, leg. Song Shimei” (IZAS).

Holotype ♂ “Myanmar (Burma), Province Kachin State Kanphant/ Grenze to China, 29–30.05.2006, N26°08'51.2', E, 098°34'58.2" Nachtfang Leg. M. Langer, S. Naumann, & S. Loeffler/ Coll. M. Langer” (ZFMK). Paratypes. 1 ♂ “Myanmar (Burma), Province Kachin State Kanphant/ Grenze to China, 29–30.05.2006, 26°08'51.2'N, 098°34'58.2"E Nachtfang Leg. M. Langer, S. Naumann, & S. Loeffler/ Coll. M. Langer” (ZFMK), 1 ♂ “Myanmar (Burma), Kachin State, Hpiman; 21.–23.V.2002; leg. A. Azuma” (ZFMK), 1 ♂ “[China] Yunnan, Pianma, 2011-V-10, N: 26.018 E: 98.625, 1970m” (IZAS).

Holotype: ♂ “Myanmar N (Burma) 25km E Putao, H-800m Nan Sa Bon vill. 06–09.05.1998 leg. S. Murzin & V. Siniaev” (ZFMK). Paratypes: 1 ♀ “Myanmar N (Burma) 25km E Putao, H-800m Nan Sa Bon vill. 06-09.05.1998 leg. S. Murzin & V. Siniaev” (ZFMK), 4 ♂♂ “Myanmar N (Burma) 65 km NE Putao, 1250 m Zi Yar Dam vill. 18/21.05.1998 leg. S. Murzin & V. Sinaev” (ZFMK).

Lectotype (here designated): ♂ “Tonkin Montes Mauson April, Mai 2-3000’ H. Fruhstorfer / abnormis Mos. [handwritten Moser]” (ZMHB). Paralectotypes: 3 ♂♂, 3 ♀♀ “Tonkin Montes Mauson April, Mai 2-3000’ H. Fruhstorfer” (ZMHB).

Holotype: ♂ “China, W-Jiangxi Jingang Shan- Ciping 2–14.VI.1994 E. Jendek & O. Šauša leg./ Coll. P. Pacholátko Invt. No./ CS10” (CPPB). Paratypes: 2 ♂♂ “China, W-Jiangxi Jingang Shan- Ciping 2-14.VI.1994 E. Jendek & O. Šauša leg.” (CP, ZFMK), 1 ♂ “Tongzhong Forestry Farm, Fangcheng, Guangxi, 9.IV.2002, light trap, leg. Xue Huaijun” (NKUT).

Holotype: ♂ “China: E-Yunnan Damaidi 2500m, Guangnan near Vietnam VII-2003 leg. Li et al.” (ZFMK). Paratypes: 1 ♂ “China, SE Yunnan, Xichou – E env., 1400-1700m, 13.–18.5.95 23°22-26'[N]/ 104°41-49'[E] L.+R. Businský lgt.” (CPPB), 1 ♂, 1 ♀ “CH, Guizhou prov., ~650m, Jiangkou (ca 50km SW), 27°32.83'N, 108°36.45'E, Shidu vill. env., 29.vi.-6.vii.2001, C. Holzschuh leg.” (CPPB).

Holotype: ♂ “Burma (Myanmar) SW Shan state Taunggyi J. Rejsek 1.–18.6.1997/ Coll. Dirk Ahrens” (ZFMK).

Holotype: ♂ “N Vietnam (Tonkin) pr. Vinh Phu 1990 Tam Dao 6.–9.v. P. Pacholátko leg./ Coll. P. Pacholátko Invt. No./ VS74” (CPPB). Paratypes: 3 ♂♂ “Vietnam N (Sa Pa) Lao Cai Prov., 250km from Hanoi bearing 31°, Sa Pa vill. Env. Hoang Lien Son Nat. Res. 27.5.–3.6.1998 1250m leg. A. Napolov” (CNAR, ZFMK), 1 ♂ “Vietnam N (Sa Pa) Lao Cai Prov., 250km from Hanoi bearing 31°, Sa Pa vill. Env. Hoang Lien Son Nat. Res. 21.–23.6.1998 1250m leg. A. Napolov” (CNAR).

Holotype: ♂ “N. Vietnam (Tonkin) Tamdao 12.-24.5.1989 Pacholátko Leg./ Coll. P. Pacholátko Invt. No./ VS 45” (CPPB). Paratype: 1 ♂ “Mt. Wuyanling, Zhejiang, 3.VIII.2007, light trap, leg. Zhu Weibing” (NKUT).

Holotype: ♂ “N. Thailand, Doi Pui, near Chiang Mai; 09.V.1085; leg. H. Nara” (ZFMK). Paratypes: 1 ♂ “[BMNH] 703021 Laos, Phongsaly, Phongsaly env., 21°41.2'N, 102°6-8'E, leg. P. Pacholátko/ DA21/ =246 Sericini Asia” (ZFMK), 1 ♂ “Thai Chiang Mai prov. 18°49’N, 98°54'E 1600m, Doi Pui mt., 2.-6.v. Vit Kubáň leg., 1996/ Coll. P. Pacholátko Invt. No./ TS114” (CPPB), 8 ♂♂, 1 ♀ “N. Thailand, Doi Pui, near Chiang Mai; 09.V.1085; leg. H. Nara” (ZFMK), 39 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54''-104°00'03"E, 1480–1510m Phou Pane Mt., 22.IV.-14.V.2008 Vit Kubáň leg./ 875 Sericini Asia spec.” (ZFMK, NMPC), 4 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"–104°00'03"E, 1480–1510m Phou Pane Mt., 22.4.–14.5.2008 Vit Kubáň leg./ 875 Sericini Asia spec.” (ZFMK, NMPC), 1 ♂ „NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20°12'N, 104°01'E; 14.iv.–15.v.2012; 1300–1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012/13“ (ZFMK), 2 ♂♂ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500–1900m, 17.5.–3.6.2007, leg. C. Holzschuh” (ZFMK), 4 ♂♂ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500–1900m, 17.5.–3.6.2007, leg. Vit Kuban” (ZFMK), 8 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"-104°00'03"E, 1480–1550m Phou Pane Mt., 9.–16.vi.2009, David Hauck leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 16 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"-104°00'03"E, 1480–1550m Phou Pane Mt., 1.–16.vi.2009, Zdenek Kraus leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 11 ♂♂ “Laos-NE, Houa Phan prov., ~20°13'N, 104°00'E, Phou Pane Mt., 1.–16.vi.2009, 1350–1500 m, M. Brancucci leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 1 ♂ “Laos-NE, Houa Phan prov., ~20°12–13.5'N, 103°59.5'–104°01'E, Ban Saluei - Phou Pane Mt., 10.-16.vi.2009, 1340–1870 m, M. Brancucci & local collectors leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 3 ♂♂ “Laos-NE, Houa Phan prov., ~20°12–13.5'N, 103°59.5'–104°01'E, Ban Sauei - Phou Pane Mt., 1340–1870 m, 15.iv–15.v.2008, Lao collectors leg.” (NHMB), 30 ♂♂ “Laos-NE, Houa Phan prov., 20°11–13'N, 103°59’-104°01'E, Ban Sauei - Phou Pane Mt., 9.–17.vi.2009, 1300–1900 m, Michael Geiser leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 1 ♂ “Laos-NE, Xieng Khouang prov., 19°38.20'N, 103°20.20'E, Phonsavan (30 km NE); Phou Sane Mt., 1420 m, 10.–30.v.2009, D. Hauck leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 11 ♂♂ “Laos-NE, Xieng Khouang prov., 19°38.20'N, 103°20.20'E, Phonsavan (30 km NE): Phou Sane Mt., 1420 m, 10.–30.v.2009, Z. Kraus leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 9 ♂♂ “Laos-NE, Xieng Khouang prov., 19°37–38'N, 103°20'E, 30 km NE Phonsavan: Ban Na Lam - Phou Sane Mt., 1300–1500 m, 10.-30.v.2009, M. Brancucci leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 15 ♂♂ “Laos-NE, Xieng Khouang prov., 19°37–38'N, 103°20'E, 30 km NE Phonsavan: Ban Na Lam - Phou Sane Mt., 1300–1500 m, 10.–30.v.2009, M. Geiser leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB).

Lectotype: ♀ “56942/ inclinata Type Brsk/ Conchin Morsb./ inclinata Brsk.* Hinter-Ind.” (ZMHB).

Syntypes: 1 ♂ “N. E. Burma R. Malaise B. M. 1945-71/ N. E. Burma Kambaiti 7000 ft. 22/4/1934 R. Malaise/ Co-Type/ Serica ponderosa co-type Arrow“ (BMNH), 1 ♂ “N. E. Burma R. Malaise B. M. 1945-71/ N. E. Burma Kambaiti 7000 ft. 22/6/1934 R. Malaise/ Co-Type/ Serica ponderosa co-type Arrow“ (BMNH), 1 ♂ “N.E. Burma Kambaiti, 2000 m 29/5. 1934 Malaise/ Typus/ Serica ponderosa n.sp. Arrow/ 3136 E91/Naturhistoriska Riksmuseet Stockholm Loan no 559/94” (NHRS), 1 ♂ “N.E. Burma Kambaiti, 7000 ft. 4-8/6. 1934 R. Malaise/ Serica ponderosa co-type Arrow/ ponderosa Arrow/ 3137 E91/Naturhistoriska Riksmuseet Stockholm Loan no 560/94” (NHRS).

Holotype ♂ “Myanmar (Burma), Province Chin/ Chin Hills Umg. Kanpetlet Natmatoung N.P. (NF), 23.VI.2008 leg. Michael Langer/ E 093°57'N, 21°13' H= ca. 1500m” (ZFMK). Paratypes: 3 ♂♂ “Myanmar (Burma), Province Chin/ Chin Hills Umg. Kanpetlet Natmatoung N.P. (NF), 23.VI.2008 leg. Michael Langer/ E 093°57'N, 21°13' H= ca. 1500m” (ZFMK).

Holotype: ♂ “Cambodia Cardamom Mts., Tumpor area, 12°22'N, 103°02'E, 1250m, Submontane forest, 27.ii-5.iii.2000, leg. M. Nuss” (SMTD). Paratypes: Cambodia: 1 ♂ Cambodia Cardamom Mts., Tumpor area, 12°22'N, 103°02'E, 1250m, Submontane forest, 27.ii-5.iii.2000, leg. M. Nuss” (ZFMK). Thailand: 1 ♂ “N-Thailand V.1990 Doi Inthanon lg. Malicky” (ZSMC), 1 ♂ “Thai 24–29.IV.1993 Doi Suithep Pacholátko & Dembicky leg./ TS73” (CPPB), 1 ♂ “Thai, 17–24.V.1991 Chiang Dao 1000m 19°25'N, 98°52'E Vit Kuban leg./ Thailand ‘91 “Thanon Thong Cha” D. Král & V. Kubáň” (ZFMK). Laos: 1 ♂ “Laos-NE Hua Phan prov., 20°12'N, 104°01'E, Phu Phan Mt., 1500–1900m, 17.5.–3.6.2007, leg. Vit Kuban” (ZFMK), 4 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"-104°00'03"E, 1480–1510m Phou Pane Mt., 22.4.–14.5.2008 Vit Kubáň leg. (NMPC, ZFMK), 2 ♂♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"-104°00'03"E, 1480–1510m Phou Pane Mt., 22.IV.–14.V.2008 Vit Kuban leg. (ZFMK), 1 ♂ “Laos-N, Louang Namtha circ., 04.05.1996 I. Pljushtch lg.” (ZFMK), 1 ♂ “Laos-NE, Houa Phan prov., ~20°12–13.5'N, 103°59.5'–104°01'E, Ban Sauei - Phou Pane Mt., 10.–16.vi.2009, 1340–1870 m, M. Brancucci & local collectors leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 2 ♂♂ “Laos-NE, Houa Phan prov., ~20°12–13.5'N, 103°59.5'–104°01'E, Ban Sauei - Phou Pane Mt., 1340–1870 m, 15.iv-15.v.2008, Lao collectors leg.” (NHMB), 1 ♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"–104°00'03"E, 1480–1550m Phou Pane Mt., 1.-16.vi.2009, Zdenek Kraus leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 1 ♂ “Laos-NE, Houa Phan prov., 20°11–13'N, 103°59'-104°01'E, Ban Sauei - Phou Pane Mt., 9.–17.vi.2009, 1300–1900 m, Michael Geiser leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 2 ♂♂ “Laos-NE, Xieng Khouang prov., 19°38.20'N, 103°20.20'E, Phonsavan (30 km NE); Phou Sane Mt., 1420 m, 10.–30.v.2009, D. Hauck leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 1 ♂ “Laos-NE, Houa Phan prov., 20°13'09–19"N, 103°59'54"–104°00'03"E, 1480–1550m Phou Pane Mt., 9.-16.vi.2009, David Hauck leg./ NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 2 ♂♂ “Laos-NE, Xieng Khouang prov., 19°38.20'N, 103°20.20'E, Phonsavan (30 km NE): Phou Sane Mt., 1420 m, 10.-30.v.2009, Z. Kraus leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 3 ♂♂ “Laos-NE, Xieng Khouang prov., 19°37–38'N, 103°20'E, 30 km NE Phonsavan: Ban Na Lam - Phou Sane Mt., 1300–1500 m, 10.-30.v.2009, M. Geiser leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň” (NHMB), 12 ♂♂, 2 ♀♀ “Laos, Attapeau prov.; Annam Highlands Mts Dong Amphan; NBCA, ca. 1160 m Nong Fa (crater lake) env., 15°05.9'N, 107°25.6'E St. Jakl lgt, 30.4.–6.5.2010” (ZFMK), 9 ♂♂, 4 ♀♀ “Laos, Attapeau prov.; Annam Highlands Mts. Dong Amphan NBCA, ca. 1160 m Nong Fa (crater lake) env., 15°05.9'N, 107°25.6'E Jiří Hájek leg. 30.iv.-6.v.2010” (NMPC). China: 3 ♂♂ “Mengla, Yunan, 21.IV.1982, leg. Jiang Shengqiao” (IZAS), 1 ♂, 2 ♀♀ “Caiyanghe, Pu'er, Yunnan, 28.VII.2007, leg. Ren Guodong, Hou Wenjun, Li Yalin”(HBUM), 1 ♂ “Mengla, Xishuangbanna, Yunnan, 4.IV.1982, leg. Wang Sumei, Zhou Jingruo” (NUYS), 1 ♂ “Mengla, Yunnan, 20.IV.1982, 670m, light trap, leg. Yu Peiyu” (IZAS).

Holotype ♂ [China] “Dongqiong, Chayu, Xizang, 16.VII.1973, 1850–2500m, leg. Huang Fusheng” (IZAS).

Holotype: ♂ “Defu, Napo, Guangxi, 19.VI.2000, 1350m, leg. Yao Jian” (IZAS). Paratypes: 1 ♂ “Defu, Napo, Guangxi, 19.VI.2000, 1350m, leg. Zhu Chaodong “ (ZFMK), 1 ♂ “Mengla, Yunnan, 21.IV.1982, leg. Jiang Shengqiao” (IZAS).

Holotype: ♂ “Laos, 21°09'N, 101°19'E, Louangnamtha pr. Namtha-> MuangSing, 5.–31.v.1997, 900-1200m Vit Kubáň leg./ LS19” (CPPB).

Holotype: ♂ “China: N-Yunnan, Baiyungshan (Bai Railing Mts.) 2400m, Yong Ren, VII-2003 leg. Ying et al.” (ZFMK).