Comprehensive Description
provided by Smithsonian Contributions to Zoology
Jerildaria joubiphoxus
DESCRIPTION (female-like).—Head about 18 percent of total body length, greatest width about 64 percent of length; rostrum unconstricted, broad, short, almost reaching apex of article 2 on antenna 1. Eyes medium, largely occluded with pigment. Article 1 on peduncle of antenna 1 about 1.2 times as long as wide, about 2.1 times as wide as article 2, apicoventral margin with about 8 setules, strongly produced dorsal apex with 4 setules; article 2 about 0.85 times as long as article 1, with apicoventral row of 3–4 setae; primary flagellum with 5 articles, about 0.75 times as long as peduncle, bearing several short aesthetascs; accessory flagellum elongate, with 4 articles. Spine formula on article 4 of antenna 2 = 2–4–4 or 2–4–5, one spine of proximal group thin, all others thick, dorsal margin with notch bearing one seta and one spine, ventral margin with 3 groups of 1–2 medium setae, 2 ventrodistal short thick spines and 2 middistal long spines; article 5 about 0.8 times as long as article 4, facial spine formula = 2–2, dorsal margin naked, ventral margin with one set of setae, ventrodistal medium seta, setule and spine; flagellum about as long as article 5 of peduncle, with 4–5 articles. Mandibles with medium palpar hump; right incisor with 3 teeth; left incisor with 2 weak branches; right lacinia mobilis bifid, distal branch little longer than proximal, flabellate, broad, denticulate, with one facial hump, proximal branch simple, pointed; left lacinia mobilis with 4 teeth plus weak accessory teeth, middle teeth shortened; right rakers 2; left rakers 3; molars triturative; palp article 1 short, article 2 with 2–3 long inner apical setae and one other shorter inner basal seta, article 3 about 0.85 times as long as article 2, oblique apex with 8 spine-seta, basofacial formula = 0. Inner plate of maxilla 1 ordinary, naked, palp with 2 apical setae, one apicomedial and one apicolateral setae. Plates of maxilla 2 extending subequally, equally wide, outer with one apicolateral seta, inner with 11 medial setae. Inner plate of maxilliped with 3 thin apical spines and setae, 2–3 mediofacial setae, outer plate with 5–6 medial and apical spines, basalmost thin, medial margin of article 2 moderately setose, article 3 not protuberant, with 5 facial setae, no lateral setae, nail of article 4 medium, with 2 accessory setules. Coxa 1 expanded distally, anterior margin straight; main ventral setae of coxae 1–4 = (4–5)–(4–5)–(4)–(1–2), posteriormost seta of coxae 1–3 shortest; anterior and posterior margins of coxa 4 strongly divergent, posterior margin very oblique, almost straight, posterodorsal corner subsharp, posterodorsal margin L-shaped, width–length ratio of coxa 4 = 1:1. Long posterior setae on article 2 of gnathopods 1–2 and pereopods 1–2 = 0–3–(2–3)–3, short anteriors = 2–2–(2–3)–2, no others; gnathopods with stout sixth article, that of gnathopod 2 larger; width ratios of articles 5–6 on gnathopods 1–2 = 20:70 and 21:73, length ratios = 88:62 and 64:80; palmar humps small, palms almost transverse; article 5 of gnathopod 1 elongate, elliptical, thin, posterior margin weakly rounded, long; article 5 of gnathopod 2 elongate, elliptical, thin, posterior margin rounded, short. Pereopods 1–2 similar; posterior margin of article 4 on pereopod 2 slightly more setose than on pereopod 1, facial setae formula on article 4 = 1–1, on article 5 = 0–0; main spine of article 5 extending to M. 70 on article 6, article 5 with no proximoposterior spines; spine formula of article 6 = 2 + 2 plus middistal spine plus posterior spine, some spines short; acclivity on inner margin on dactyls of pereopods 1–2 weak, emergent setule long, midfacial pluseta very large, highly proximal, nail weakly distinct. Coxae 5–7 posteroventral setule formula = 1–1–2. Articles 4–5 of pereopods 3–4 narrow, facial spine rows sparse, pereopod 3 especially small, facial ridge formula on article 2 of pereopods 3–5 = 0–1–1; width ratios of articles 2, 4, 5, 6 of pereopod 3 = 44:21:19:11, of pereopod 4 = 66:27:21:12, of pereopod 5 = 90:20:18:10, length ratios of pereopod 3 = 52:19:29:30, of pereopod 4 = 82:50:45:34, of pereopod 5 = 110:24:30:28; article 2 of pereopod 5 not reaching middle of article 5; medial apex of article 6 very finely combed. Posteroventral corner of epimeron 1 rounded, posterior margin weakly convex, margin above corner with setule, anteroventral margin naked, posteroventral face with one or no long seta; posteroventral corner of epimeron 2 rounded–quadrate, posterior margin straight, bearing one setule, facial setae = 4, posteriormost pair set vertically; posteroventral corner of epimeron 3 weakly protuberant, with setule sinus, posterior margin straight, with 2 long setae set in notches, occasional additional setule notch, ventral margin and face naked; epimera 1–2 with large seta on posterodorsal margin set in weak notch. Urosomite 1 with lateral setule at base of uropod 1, long ventral seta, articulation line complete; urosomite 3 weakly protuberant dorsally. Rami of uropods 1–2 with fused enlarged apical nails, outer ramus of uropod 1 with 2 dorsal spines, inner with 2, outer ramus of uropod 2 with one dorsal spine, inner naked; peduncle of uropod 1 with one small apicolateral spine, medially with 3 marginal spines, apicalmost ordinary; peduncle of uropod 2 with 4 dorsal spines, medially with one small apical spine; apicolateral corners on peduncles of uropods 1–2 lacking comb. Peduncle of uropod 3 with 2 ventral spines, dorsally with one lateral setule, one medial spine; rami feminine, inner extending to M. 33 on article 1 of outer ramus, apex with one short seta, article 2 of outer ramus short, 0.10, bearing 2 long setae, apicomedial margin of article 1 with sets of 2 and 3 setae, lateral margin with 4 acclivities, setal formula = 1–1–2–3–3. Telson ordinary, length–width ratio = 35:31, almost fully cleft, each apex wide, truncate, lateral acclivity weak, bearing lateral setule, spine next medial much longer, medially with short seta, occasionally with extra lateral short seta, midlateral setules ordinary to large. Cuticle with ordinary sparse bulbar setules of varying sizes mixed with pipes, emergent setules occasionally branched.
OBSERVATIONS.—Epistome protruding in front of upper lip, both parts fully articulate, upper lip with weak midhump ventrally, bearing minute sharp cusps; inner lobes of lower lip fused for most of length, marked medially with longtitudinal plaque, each outer lobe with cone. Neither specimen of this species shows any mark of sex except for one possible rudimentary penial process on specimen “w”.
ILLUSTRATIONS.—Outline of epistome shown on lateral view of head; left eye of holotype displaced medially in dorsal view of head; left mandible and article 3 of antenna 1 not flattened; apically rounded spines on uropods 1–2 blunted by damage.
HOLOTYPE.—AM, sex unknown “b,” 5.50 mm.
TYPE-LOCALITY.—SBS Long Reef, 27 Apr 1972, Long Reef, New South Wales, N of North Head, Sydney, Australia, 43.3 m.
VOUCHER MATERIAL.—Type-Locality, sex unknown “w,” 5.46 mm (illus.), only known specimen in addition to holotype.
DISTRIBUTION.—New South Wales, Long Reef, Sydney, 43 m.
Leptophoxoides J. L. Barnard
Leptophoxoides J. L. Barnard, 1962:51.
DIAGNOSIS.—Eyes present. Flagella of antennae 1–2 reduced in female. Article 2 of antenna 1 especially shortened, ventral setae if present confined apically. Article 1 of antenna 2 not ensiform; [? article 3 with 2 setules]; facial spines on article 4 in one main row; article 5 especially short. Right mandibular incisor with 3 teeth; molar strongly triturative, not bearing fuzz; palpar hump small. Palp of maxilla 1 uniarticulate; inner plate naked. Setation of maxilla 2 weak. Inner plate of maxillipeds ordinary; apex of palp article 3 strongly protuberant, dactyl elongate, apical nail distinct, medium. Gnathopods dissimilar, gnathopod 2 strongly enlarged; article 5 of gnathopods 1–2 very short, cryptic, with eusirid attachment, palms oblique, hands of gnathopods 1–2 ordinary to broadened, poorly setose anteriorly. Article 5 of pereopods 1–2 setose posteroproximally. Article 2 of pereopod 3 of broad form, articles 4–5 of pereopods 3–4 narrow, article 2 of pereopods 3–4 not setose posteriorly, pereopod 5 ordinary, article 2 naked ventrally, article 3 ordinary, [? dactyl normal]. Epimera 1–2 lacking long posterior setae, without midfacial setae above ventral facial ridge, epimeron 3 of rounded classification and lacking long setae. Urosomite 1 generally naked; urosomite 3 without dorsal hook or special process. Peduncle of uropod 1 normally elongate, without apicoventral spike, without special enlarged apicolateral–medial spine, only peduncular apices of uropod 1 combed, inner ramus of uropod 1 with marginal spines in one row, no rami continuously spinose to apex, inner ramus of uropod 2 ordinary. Uropod 3 ordinary; elongate article 2 of outer ramus carrying one medium apical seta. Telson elongate, with only one apical spine on each lobe plus setules, without special dorsal and lateral spines or setae.
DESCRIPTION.—Rostrum fully developed. [Fuzz on article 1 of antenna 1 in male unknown, calceoli on male antennae 1–2 unknown. Prebuccal parts unknown.] Right lacinia mobilis [? bifid or simple], thin; article 1 of mandibular palp short, palp medium to thick, article 2 without outer setae, apex of article 3 oblique. Lower lip lacking cones. Outer plate of maxilla 1 with 7 spines, no spine especially thickened. Inner plates of maxilliped especially thin, poorly armed. Coxae 2–4 without special anterodorsal humps. All posterior spines on article 6 of pereopods 1–2 thin and seta-like, [midapical spine or seta unknown]. Article 2 of pereopod 5 without facial setae. Peduncle of uropod 1 with dorsolateral spines widely spread, medial spines confined apically; peduncle of uropod 2 with only one medial spine or setule confined apically. Peduncle of uropod 3 lacking extra subapical setae or spines. Telson with ordinary pair of midlateral or dorsal setules on each side.
TYPE-SPECIES.—Leptophoxoides molaris J. L. Barnard, 1962 (monotypy).
COMPOSITION.—Unique.
