Smicridea (Smicridea) annulicornis (Blanchard ex Gay 1851)

Smicridea (Smicridea) annulicornis (Blanchard)

Hydropsyche annulicornis Blanchard, 1851:140.—Hagen, 1864:821.—Ulmer, 1904:24; 1905a:68; 1905b:18.

Smicridea annulicornis (Blanchard).—Ulmer, 1913:389–390, 406.—Fischer, 1963:131.

Smicridea (S.) annulicornis (Blanchard).—Flint, 1974b:88.

Rhyacophylax chilensis Navás, 1923:23; 1922 [1924]:367; 1928:126; 1930:361; 1932b:83; 1934a:28.—Lestage, 1925:41.—Fischer, 1963:136 [new synonymy].

Smicridea chilensis (Navás).—Schmid, 1949 [1950]:344–345 [sic Smicridia].—Flint, 1967:55.—Fischer, 1972:144.

Smicridea (S.) chilensis (Navás).—Flint, 1974b:88.

This, one of the most ubiquitous caddisflies in the Chilean Subregion, appears to be most closely related to S. decora. The two are usually easily distinguished by the darker color and smaller eyes of annulicornis. The deeply cupped lateral plates of the phallus that lack any mesobasal shoulder and the very small or lacking dorsolateral lobes are diagnostic in the male genitalia of annulicornis. The genitalia of the female are similar to that of decora, but may be recognized by the shape of the clasper receptacle. In decora the outer face is unmodified, but in annulicornis there is a distinct, oblique posteroventral fold overhanging the opening slightly and a similar, but smaller, one anteroventrally.

ADULT.—Length of forewing, 6–10 mm. Color brown, appendages slightly paler, antennae annulate; forewings pale brownish, variable, marked with dark brown flecks, often producing a distinct pattern, other times producing a more uniform speckling. Eye of male in frontal aspect with diameter slightly greater than half of interocular distance. Fifth sternum with anterolatereral processes slightly longer than sternum; with 2 pairs of internal sacs, anterior pair as long as its segment, posterior pair distinctly shorter than its segment.

Male Genitalia: Ninth segment with anterior margin nearly vertical. Tenth tergum elongate, tip divided dorsomesally; tergite with apex rounded in dorsal, and enlarged and upturned in lateral aspect. Clasper with basal segment inflated apicad; apical segment elongate, bluntly pointed. Phallus tubular, base at right angles to stem, width of basal opening slightly more than twice diameter of stem, apex distinctly enlarged; lateral plate 1½–2 times longer than broad, ventral margin well above ventral margin of stem, in ventral aspect with mesobasal shoulder lacking, deeply cupped, and with inner margins slightly concave but with ventral margin usually more lateral than dorsal; dorsolateral lobe very small, or sometimes apparently lacking.

Female Genitalia: Eighth sternite roughly rectangular, with posteromesal angle at about 90°; anterolateral angle produced, rounded. Ninth tergum strongly produced anteroventrally. Clasper receptacle deep, pocket-like, produced ventromesally; in dorsal aspect about ½ length of tergum, in lateral aspect with a narrow anterior overhang and a broader, oblique posteroventral overhang. Vaginal sclerite narrow, transversely oval, with a small posteromesal lobe bearing a small opening.

TYPE MATERIAL.—Hydropsyche annulicornis Blanchard, type in MNHNP labeled: “MUSEUM PARIS Chili Gay 15–43”; “15–43”; “TYPE”; “Smicridea annulicornis Blanch (Blanchard's Type von Hydropsyche annulicornis Blanch. Ulmer vid. 1913).”

Rhyacophylax chilensis Navás: Type in MZB labeled: “Marga Marga (Chile) 1. 1919”; “Rhyacophylax chilensis Nav. P. Navas S.J., det.”; “Typus.” The pin is bare except for a balsam mount on the pin with genitalia. However, both the original description and Schmid (1949) state that the type is a ! The genitalia are annulicornis as are most other specimens determined as chilensis by Navás, and other examples from Estero Marga Marga at Los Perales. Based on this evidence about the identity of the type, I make this synonymy.

MATERIAL EXAMINED.—ARGENTINA: PCIA. CHUBUT: Lago Puelo, [S] El Bolsón, 220 m, 26 Feb 1979, Nielsen et al., 2 (ZMC); Arroyo Golondrinas, 6 km N Lago Puelo, 8 Feb 1974, O.S. Flint, Jr., 2 (NMNH); Cholila, 26 Jan 1965, A. Kovacs, 1 , 1 (AMNH); Esquel, 1 Jan 1982, Nielsen et al., 2 , 3 (ZMC); El SagraRío Puerto, Lago Menéndez, [W] Esquel, 600 m, 2–4 Jan 1982, Nielsen et al., 1 (ZMC); Corcovado, [W] Tecka, 750 m, 17 Feb 1979, Nielsen et al., 3 (ZMC); 3 km N Trevellín, 28 Jan 1987, C.M. & O.S. Flint, Jr., 2 , 9 (NMNH).

PCIA. NEUQUÉN: San Martín de los Andes, 2–18 Nov 1959, J.E. Foerster, 3 , 3 (CNC); same, but Jan 1978, M. Gentili, 1 (NMNH); Puente Blanco, Cerro Chapelco, San Martín de los Andes, 25 Feb 1978, C.M. & O.S. Flint, Jr., 1 , 1 (NMNH); tributary Arroyo Trompul, W San Martín de los Andes, 23 Feb 1978, C.M. & O.S. Flint, Jr., 1 (NMNH); Arroyo Pilpil, near San Martín de los Andes, 22 Feb 1978 C.M. & O.S. Flint, Jr., 1 (NMNH); Lago Lacar, 5 km E Hua-Hum, 640 m, 25 Nov 1981, Nielsen et al., 4, 2 (ZMC); Río Quilquihue at Lago Lolog, 22–23 Jan 1974, O.S. Flint, Jr., 100s (NMNH); Río Quilquihue, Quilquihue, 26 Jan 1974, O.S. Flint, Jr., 100s (NMNH); Arroyo Rosales, near San Martín de los Andes, 22 Jan 1974, O.S. Flint, Jr., 1 , 1 (NMNH); Aluminé, 20 Jan 1949, F. Monros, 1 (CNC); same, but 14 Mar 1979, Nielsen et al., 4 , 5 (ZMC); Río Aluminé, 9 km N Aluminé, 27 Feb 1978, C.M. & O.S. Flint, Jr., 6, 10 (NMNH); same, but 5 km N Aluminé, 2, 2 (NMNH); same, but 5 km S Pilolil, 2 Feb 1987, 3 , 4 (NMNH); Río Litrán, 9 km N Lago Aluminé, 3 Feb 1987, C.M. & O.S. Flint, Jr., 5 , 6 (NMNH); Río Totoral, 24 km NW Villa La Angostura, 20 Feb 1978, C.M. & O.S. Flint, Jr., 5 , 11 (NMNH); same, but 31 Jan 1987, 2 (NMNH); Río Ruca Malén, 25 km N Villa La Angostura, 1 Feb 1987, C.M. & O.S. Flint, Jr., 1 , 4 (NMNH); Río Malleo, 22 km N Junín de los Andes, 2 Mar 1978, C.M. & O.S. Flint, Jr., 20 11 (NMNH); same, but 2 Feb 1987, 6 , 6 (NMNH); Arroyo del Gato, 8 km S Rahué, 2 Mar 1978, C.M. & O.S. Flint, Jr., 11, 1 (NMNH); Lago Rucachoroi, W Aluminé, 1–2 Mar 1978, C.M. & O.S. Flint, Jr., 1 (NMNH); same, but 15 Mar 1979, Nielsen et al., 1 (ZMC); Lago Moquehué at Arroyo Marcial, 28 Feb 1978, C.M. & O.S. Flint, Jr., 1 (NMNH); Río Agrio, N Zapala, 9–11 Dec 1983, L.E. Peña G., 26 , 12 (NMNH); 5 km SE Lago Huechulafquén, 26 Jan 1974, O.S. Flint, Jr., 5 , 1 (NMNH); Rodeo Grande, Lago Tromén, 900 m, 1 Jan 1979, Nielsen et al., 14 8 (ZMC); same, but 12 Mar 1979, 2 (ZMC); Río Nonthué, Estación Forestal Pucará, 28–31 Jan 1974, O.S. Flint, Jr., 24 , 43 (NMNH); Lago Lacar, Estación Forestal Pucará, 29–30 Jan 1974, O.S. Flint, Jr., 7 , 33 (NMNH); same, but 2 Dec 1978, Nielsen et al., 1 (ZMC); same, but 25–26 Dec 1978, 18 , 1 (ZMC); same, but 28–29 Nov 1981, 5 , 3 (ZMC) same, but 26–27 Dec 1981, 2 , 2 (ZMC); Pantano, near Estación Forestal Pucará, 29 Jan 1974, O.S. Flint, Jr., 3 (NMNH); Río Limay, Neuquén, 17–18 Feb 1978, C.M. & O.S. Flint, Jr., 5, 13 (NMNH); Río Limay, Confluencia, 3 Mar 1978, C.M. & O.S. Flint, Jr., 3 , 2 (NMNH); Piedra de Aguila, 18 Dec 1978, Nielsen et al., 13 , 10 (ZMC).

PCIA. Río NEGRO: El Bolsón, 7 Jan–13 Feb 1961, Topal, 17, 13 (HNHM & NMNH); same, but 7 Nov 1961, l (HNHM); Cascada Mallín Ahogado, [N] El Bolsón, 9 Feb 1974, O.S. Flint, Jr., 7 , 4 (NMNH); General Fernández Oro, Jan–Feb 1976, S. Coscarón, 18 , 19 (AMNH & NMNH); Río Negro, General Fernández Oro, 17 Feb 1978, C.M. & O.S. Flint, Jr., 13 , 5 (NMNH); Lago Pellegrini, NE Cipolletti, 17 Feb 1978, C.M. & O.S. Flint, Jr., 1 (NMNH); Colonia Suiza, Bariloche, 810 m, 20 Nov 78, Nielsen et al., 1 (ZMC); Ñirihuau, [E] Bariloche, 9 Dec 1978, Nielsen et al., 1 (ZMC); same, but 2 Jan 1979, 1 , 1 (ZMC); Río Ñirihuau, Estación Ñirihuau, 11 Feb 1974, O.S. Flint, Jr., 3 , 3 (NMNH); Puerto Blest, Lago Nahuel Huapí, 18–23 Dec 1978, Nielsen et al., 2 5 (ZMC); 5 km S Río Villegas, 7 Feb 1974, O.S. Flint, Jr., 1 (NMNH).

CHILE: PCIA. ACONCAGUA: N El Tártaro, Putaendo, 5–6 Feb 1984, L.E. Peña G., 5 , 5 (NMNH); Curimon (W) Los Andes, 700 m, 28 Mar 1979, Nielsen et al., 1 , 3 (ZMC).

PCIA. AISÉN: Lago Risopatrón, 17 km N Puyuhuapi, 24 Jan 1987, C.M. & O.S. Flint, Jr., 3 , 11 (NMNH); Río Cisnes, 10 km E Puerto Cisnes, 25 Jan 1987, C.M. & O.S. Flint, Jr., 1 , 1 (NMNH).

PCIA. ARAUCO: Estero Peral, Contulmo, 1–2 Jan 1966, Flint & Cekalovic, 21 , 30 (NMNH); Puente Trongol [∼12 km S Curanilahue], 1 Jan 1966, Flint & Cekalovic, 1 (NMNH); same, but 15–16 Oct 1969, Flint & Barria, 1 (NMNH).

PCIA. BÍO-BÍO: Santa Bárbara, 6 Feb 1958, L.E. Peña G., 1 (CNC); Río Queuco, E Santa Bárbara, 17–18 Mar 1984, L.E. Peña G., 58 , 10 (NMNH); Estero Huequecura, 25 km E Santa Bárbara, 24 Jan 1978, C.M. & O.S. Flint, Jr., 34 , 32 (NMNH); El Abanico, 1100 m, 17–19 Mar 1984, L.E. Peña G., 4 (NMNH); 5 km W Tucapel, 28 Dec 1950, Ross & Michelbacher, 87 , 10 (CAS); Salto de Laja, 17–18 Apr 1966, L.E. Peña G., 10 , 11 (NMNH); ∼4 km N Salto de Laja, ∼200 m, 12 Jan 1982, D.R. Davis, 3 , 7 (NMNH); Rt. 5, 2 km N Río Laja, 16 Jan 1987, C.M. & O.S. Flint, Jr., 4 (NMNH).

PCIA. CACHAPOAL: La Leonera [15 km E Codegua], 26–28 Dec 1954, L.E. Peña G., 19 , 42 (CNC); same, but 12–13 Feb 1986, 5 , 8 (NMNH); Graneros, 4 Mar 1962, L.E. Peña G., 1 (CNC); Río Peuco, Pilay, ∼45 km S Santiago, 800 m, 23–25 Nov 1981, Davis & Peña, 18 24 (NMNH); Río Claro, 5 km N Rengo, 300 m, 23 Jan 1978, C.M. & O.S. Flint, Jr., 1 , 2 (NMNH); Palmar de Cocolán, NE Las Cabras, 15–16 Dec 1986, L.E. Peña G., 2 (NMNH).

PCIA. CAUQUENES: Near coastal stream, 17.5 km S Curanipe, 50 m, 25 Jan 1979, Davis et al., 8, 2 (NMNH); W Cauquenes, 3 Oct 1983, L.E. Peña G., 1 , 4 (NMNH); same, but 25 Mar 1984, 1, 1 (NMNH); Cayunanquil, W Cauquenes, 400 m, 23–31 Jan 1981, L.E. Peña G., 4 (NMNH); Alto Tregualemu, ∼20 km SE Chovellén, 500 m, 26–27 Jan 1979, Davis et al., 31 6 (NMNH); Tregualemu, 14 Mar 1967, T. Cekalovic K., 11 6 (NMNH).

PCIA. CAUTÍN: Cerro Nielol, Temuco, 200 m, 9 Feb 1979, Davis et al., 33 30 (NMNH); 20 km E Temuco, 8 Jan 1951, Ross & Michelbacher, 34, 28 (CAS); Allipén, Feb 1972, L.E. Peña G., 3 , 7 (NMNH); Río Cautín, Cajón, 3 Jan 1966, Flint & Cekalovic, 4 , 2 (NMNH); same, but 24 Oct 1969, Flint & Barria, 6 , 1 (NMNH); Río Cautín, Temuco, 30 Dec 1967, T. Cekalovic K., 1 , 2 (NMNH); Huife, ∼40 km E Pucón, 18 Jan 1987, C.M. & O.S. Flint, Jr., 7 , 9 (NMNH); near Pucón [Río Minetué, ∼12 km E Pucón], 4 Jan 1966, Flint & Cekalovic, 7 , 6 (NMNH); 10 mi [∼16 km] NE Pucón, 12 Jan 1951, Ross & Michelbacher, 6 (CAS); Villarrica, 4–5 Jan 1966, Flint & Cekalovic, 2 (NMNH); 30 km NE Villarrica, 1–30 Jan 1965, L.E. Peña G., 11 6 (NMNH); Fundo El Coigue, 27 km NE Villarrica, 500 m, 28 Feb–3 Mar 1979, Davis et al., 100s (NMNH); Fundo Neltume, 2 km N Villarrica, 200 m, 27 Feb 1979, Davis et al., 4, 3 (NMNH); Río Toltén, Teodoro Schmidt, 16–17 Mar 1987, L.E. Peña G., 1 (NMNH).

PCIA. CHACABUCO: Cuesta La Dormida, W Tiltil, 600–800 m, 13–18 Nov 1982, L.E. Peña G., 1 (NMNH).

PCIA. CHILOÉ: Lago Tepuhueco, 1–3 Mar 1984, L.E. Peña G., 1 (NMNH); same, but 11–15 Dec 1985, 2, 4 (NMNH); Puntra, ∼30 air km S Ancud, 50 m, 15 Dec 1985, L.E. Peña G., 18 , 28 (NMNH); same, but 21–22 Dec 1981, Davis & Peña, 3 7 (NMNH); Huequetrumao, 22 km N Quellón, 26–28 Dec 1981, L.E. Peña G., 1 (NMNH); Río Butalcura, 21 Oct 1969, Flint & Barria, 35 (NMNH); Río Cude, Pudguapi, 20 Jan 1987, C.M. & O.S. Flint, Jr., 2 , 4 (NMNH).

PCIA. CHOAPA: Hacienda Illapel, 29–30 Oct 1954, L.E. Peña G., 14 , 2 (CNC); Río Choapa, Salamanca, 6 Feb 1986, L.E. Peña G., 2 , 10 (NMNH); El Naranjo, S Caimanes, 7 Feb 1986, L.E. Peña G., 1 (NMNH); Cuesta Cavilolén, NE Los Vilos, 5 Feb 1986, L.E. Peña G., 3 (NMNH); Ñague, 11 km N Los Vilos, 20 m, 4–5 Nov 1981, D.R. Davis, 2 , 1 (NMNH).

PCIA. CONCEPCIÓN: Concepción, 26 Nov–10 Dec 1959, light trap, 3 (NMNH); same, but 29 Jan 1967, L.E. Peña G., 6 , 4 (NMNH); 20 km E Concepción, 19–20 Mar 1984, L.E. Peña G., 17 , 6 (NMNH); ∼20 km S Coelemu, ∼50 m, 28 Jan 1979, Davis et al., 21 , 9 (NMNH).

PCIA. COPIAPÓ: Río Jorquera, Cordillera Copiapó, 27 Oct 1967, L.E. Peña G., 8 (NMNH); La Junta, E Copiapó, 2 Oct 1980, L.E. Peña G., 26 , 9 (NMNH).

PCIA. CORDILLERA: El Manzano, 9 Feb 1950, L.E. Peña G., 6 , 4 (CNC); E1 Alfalfal, Cordillera Santiago, 3 Dec 1963, L.E. Peña G., 6 (NMNH); Los Maitenes, 2 Mar 1964, L.E. Peña G., 29 , 18 (NMNH); same, but 30 Oct 1964, 32 , 2 (NMNH).

PCIA. CURICO: Estero Potrero Grande, 3 km E Potrero Grande, 8 Feb 1987, C.M. & O.S. Flint, Jr., 9 , 1 (NMNH); Las Tablas, E Curico [13 km E Potrero Grande], 26 Mar 1984, L.E. Peña G., 3 , 2 (NMNH); same, but Feb 1985, 2 (NMNH); Río Teno, ∼40 km E Curico, 800 m, 25–28 Nov 1981, Davis & Peña, 2 (NMNH); El Coigo, 1 Mar 1968, Flint & Peña, 100s (NMNH); Río Los Morongos, Bajo Los Morongos, 653 m, 21–22 Nov 1972, Pescador & Barria, 1 , 1 (NMNH).

PCIA. ELQUI: Rivadavia, 16 May 1952, L.E. Peña G., 1 , 3 (CNC); Quebrada Seca, 16 May 1952, L.E. Peña G., 1 , 2 (CNC); Vicuña, near irrigation ditch, 30 Nov 1976, Gurney & Barria, 7 , 14 (NMNH); 22 mi [∼36 km] S La Sirena, 9 Dec 1950, Ross & Michelbacher, 1 (CAS).

PCIA. HUASCO: 3 km S Freirina, 13–14 Nov 1981, D. & M. Davis, 1 (NMNH); El Tránsito, E Vallenar, 25 Oct 1980, L.E. Peña G., 6 , 21 (NMNH); Río Conai, Cordillera Vallenar, 2 Dec 1967, L.E. Peña G., 9 , 12 (NMNH); Quebrada Pinte, Cordillera Vallenar, 2 Dec 1967, L.E. Peña G., 100s (NMNH).

PCIA. LIMARI: 5 mi [∼8 km] W La Junta, 7 Dec 1950, Ross & Michelbacher, 2 , 1 (CAS).

PCIA. LINARES: Hacienda San Manuel [∼31 km SE Parral], 14 Jan 1953, L.E. Peña G., 8 (CNC); Estero de Leiva [near Hacienda San Manuel], 8 Jan 1953, L.E. Peña G., 8 , 2 (CNC); Rinconada de Parral [36°04′S; 71°46′W], 19–20 Mar 1952, L.E. Peña G., 1 , 2 (CNC); Tranque de Bullileo, 10–12 Jan 1979, Davis et al., 9 , 11 (NMNH); Fundo Malcho, Jan 1958, L.E. Peña G., 1 (CNC); Puente Malcho, near Río Longavi, 600 m, 13–15 Jan 1979, Davis et al., 35 , 21 (NMNH); Río Ancoa, 35 km E Linares, 320 m, 23 Jan 1978, C.M. & O.S. Flint, Jr., 6 , 22 (NMNH).

PCIA. LLANQUIHUE: Río Maullín [near Llanquihue], 6 Jan 1966, Flint & Cekalovic, 100s (NMNH); Salto Chamiza, Correntoso, 100 m, 19 Jan 1987, C.M. & O.S. Flint, Jr., 13 , 40 (NMNH); El Chingue, N Correntoso (S Volcán Calbuco), 300 m, 20–25 Jan 1980, L.E. Peña G., 29 , 62 (NMNH); Hornohuinco, near Correntoso, 7–11 Feb 1985, L.E. Peña G., 2 (NMNH); Lago Chapo, E Puerto Montt, 20–28 Dec 1985, L.E. Peña G., 11 , 43 (NMNH); Petrohué, Lago Todos Los Santos, 600 m, 1–3 Jan 1982, Davis & Peña, 9 , 4 (NMNH); same, but 28–29 Mar 1968, L.E. Peña G., 2 , 3 (NMNH); same, but Jan 1969, 10 , 13 (NMNH).

PCIA. LOS ANDES: Los Andes, 12 Nov 1957, J. Illies, 1 (NMNH).

PCIA. MAIPO: Aculeo, 18 Dec 1986, L.E. Peña G., 1 (NMNH); Las Canchas, Aculeo, 8–11 Dec 1983, Yrarrázaval, 1 (NMNH); Quebrada El Cepillo, E slope coastal range, SW Aculeo, 26–28 Mar 1981, L.E. Peña G., 2 (NMNH).

PCIA. MALLECO: Angol, 13 Mar 1925, D.S. Bullock, 1 (NMNH); same, but 1 Feb 1929, 2 (NMNH); same, but 26 Mar 1936, 1 (NMNH); same, but 6 Jan 1937, 2 (NMNH); same, but 17 Feb 1956, L.E. Peña G., 7 , 2 (CNC); Vegas Blancas, 27 km W Angol, 700 m, 17 Jan 1987, C.M. & O.S. Flint, Jr., 1 (NMNH); Río Manzanares [∼10 km W Purén], 2 Jan 1966, Flint & Cekalovic, 1 , 1 (NMNH); same, but 19 Oct 1969, Flint & Barria, 1 , 7 (NMNH); Victoria, Nov 1979, L.E. Peña G., 1 , 1 (NMNH); 7 km N Victoria, 3 Jan 1966, Flint & Cekalovic, 1 , 1 (NMNH); Tolhuaca, 11 Jan 1962, L.E. Peña G., 2 (CNC); same, but 15–20 Jan 1959, 5 , 1 (CNC); Río Blanco, E Curacautín, 5–27 Mar 1950, L.E. Peña G., 7 , 3 (CNC); same, but Feb 1964, L.E. Peña G., 2 (NMNH); Curacautín, 15 Dec 1949, L.E. Peña G., 1 (CNC); E Lonquimay, 1000 m, 21–23 Dec 1976, L.E. Peña G., 31 , 30 (NMNH); Río Bío-Bío, 17 km E Lonquimay, 4 Feb 1987, C.M. & O.S. Flint, Jr., 3 , 6 (NMNH); Pino Hachado, E Lonquimay, 1600 m, 17–20 Feb 1980, L.E. Peña G., 6 , 25 (NMNH); same, but 6–10 Jan 1959, 5 , 1 (CNC); Laguna Icalma, Cordillera Lonquimay, 2 Jan 1968, L.E. Peña G., 6 , 4 (NMNH); Rucanuco [outlet to Lago Icalma], Cordillera Lonquimay, 9–13 Jan 1959, L.E. Peña G., 2 (CNC); same, but 4 Jan 1968, 100s (NMNH); Liucura, 9–11 Jan 1959, L.E. Peña G., 2 , 2 (CNC); Marimenuco, 10–13 Jan 1959, L.E. Peña G., 1 (CNC); same, but 10–13 Jan 1964, 6 , 2 (NMNH); El Arco, 2–6 Jan 1959, L.E. Peña G., 5 , 3 (CNC); Cordillera Las Raíces, W Lonquimay, 1700 m, Feb 1980, L.E. Peña G., 1 , 2 (NMNH); same, but 1650 m, 7–8 Feb 1979, Davis et al., 2 , 4 (NMNH); 10 mi [∼16 km] N Perquenco, 6 Jan 1951, Ross & Michelbacher, 1 (CAS).

PCIA. MELIPILLA: Bollenar, N Melipilla, 15 Feb 1986, L.E. Peña G., 1 , 1 (NMNH).

PCIA. ÑUBLE: Puente Espinal, near Santa Clara, 24 Oct 1969, Flint & Barria, 5 , 1 (NMNH); 50 km E San Carlos, 26 Dec 1950, Ross & Michelbacher, 15 , 9 (CAS); Río Pinto, E Chillán, 24 Oct 1969, Flint & Barria, 30 , 37 (NMNH); Recinto, 30 Nov 1951, L.E. Peña G., 4 , 2 (CNC); same, but 4–6 Mar 1968, Flint & Peña, 5 , 2 (NMNH); same, but 800 m, 22–23 Jan 1979, Davis et al., 24 , 15 (NMNH); Río Chillán, near Recinto, 6 Mar 1968, Flint & Peña, 100s (NMNH); Atacalco, near Recinto, 700 m, 28 Nov 1951, L.E. Peña G., 16 (CNC); same, but 22 Jan 1955, 1 (CNC); same, but 17–18 Dec 1976, 45 , 9 (NMNH); Los Cipreces [N Recinto], 25 Nov 1951, L.E. Peña G., 1 , 2 (CNC); Los Pellines [near Los Cipreces], 2 Mar 1952, L.E. Peña G., 6 , 3 (CNC); same, but 2 Dec 1951, 22 (CNC); Cueva de los Pincheira, near Recinto, 17 Dec 1976, Gurney & Barria, 1 (NMNH); Las Trancas, 21 km E Recinto, 20 Jan 1953, L.E. Peña G., 1 (CNC); same, but 6–11 Feb 1966, 1 (NMNH); same, but 14–15 Dec 1976, 11 , 3 (NMNH); same, but 17–20 Dec 1983, 4 , 4 (NMNH); same, but 20–23 Mar 1984, 1 (NMNH); same, but 1260 m, 23–30 Feb 1956, 1 (CNC); same, but 2 Mar 1968, Flint & Peña, 3 , 1 (NMNH); same, but near high waterfall, 1300 m, 16–19 Jan 1979, Davis et al., 48 , 41 (NMNH); same, but Shangri-La, SW side Volcán Chillán, 1600 m, 19–21 Jan 1979, Davis et al., 30 , 27 (NMNH); same, but 14 Dec 1983, L.E. Peña G., 5 , 5 (NMNH); Río Niblinto, E Coihueco, 19–20 Jan 1968, L.E. Peña G., 100s (NMNH); Fundo El Roble, E Coihueco, 17 Jan 1968, L.E. Peña G., 58 , 28 (NMNH).

PCIA. OSORNO: Parque Nacional Puyehue, Anticura, 28–31 Dec 1985, L.E. Peña G., 3 , 3 (NMNH); same, but 1–5 Jan 1986, 20 , 11 (NMNH); same, but 350 m, 18 Dec 1981, Nielsen et al., 16 , 1 (ZMC); same, but Río Anticura, 31 Jan–13 Feb 1978, C.M. & O.S. Flint, Jr., 100s (NMNH); same, but 3 km E Anticura, 3 Feb 1978, 37 , 35 (NMNH); same, but Río Golgol, 2 Feb 1978, 30 , 35 (NMNH); same, but Salto del Indio, 1–12 Feb 1978, 1 , 38 (NMNH); same, but Río Pescadero, 7 Feb 1978, 3 (NMNH); same, but Lago el Toro, 7–8 Feb 1978, 5 , 2 (NMNH); same, but Aguas Calientes, 6 Feb 1978, 2 , 47 (NMNH); same, but 450 m, 12 Nov 1981, Nielsen et al., 1 (ZMC); same, but Aguas Calientes to 2 km S, 600 m, 10–22 Feb 1979, Davis et al., 3 , 5 (NMNH); same, but 12– 20 Dec 1981, 1 , 4 (NMNH); same, but Río Chanleufú, 1 km S Aguas Calientes, 8–9 Feb 1978, C.M. & O.S. Flint, Jr., 1 , 12 (NMNH); 10 km E Puyehue, 24 Jan 1951, Ross & Michelbacher, 4 , 2 (CAS); Tril-Tril, S Pucatrihue, 1–10 Feb 1980, L.E. Peña G., 15 (NMNH); Río Rahue, 20 Oct 1969, Flint & Barria, 1 , 11 (NMNH); Río Rahue, outlet Lago Rupanco, 31 Oct 1957, J. Illies, 1 (NMNH); La Picada, W Volcán Osorno, 600 m, 12–22 Jan 1980, L.E. Peña G., 3 , 5 (NMNH).

PCIA. PALENA: Río Futaleufú, 37 km SW Futaleufú, 27 Jan 1987, C.M. & O.S. Flint, Jr., 5 , 4 (NMNH); Río Ventisquero, 16 km S Puerto Cardenas, 23–24 Jan 1987, C.M. & O.S. Flint, Jr., 2 (NMNH).

PCIA. PETORCA: Cuesta El Melón, 14–16 Nov 1985, L.E. Peña G., 44 (NMNH); E La Ligua, 26 Sep 1980, L.E. Peña G., 1 , 2 (NMNH); Los Hornos, Guaguen, 2–3 Oct 1986, L.E. Peña G., 1 , 3 (NMNH).

PCIA. QUILLOTA: Estero Marga-Marga, near Perales, 9 Mar 1968, Flint & Peña, 20 , 6 (NMNH).

PCIA. TALCA: El Radal, Jan 1950, L.E. Peña G., 1 (CNC); same, but 23–30 Nov 1957, 10 (CNC); Tonlemo, 15 Feb 1950, L.E. Peña G., 22 , 13 (CNC); same, but 14–21 Dec 1984, 21 , 13 (NMNH); Alto Vilches, 11 Dec 1976, L.E. Peña G., 1 (NMNH); Molina, 7–8 Mar 1968, Flint & Peña, 1 , 1 (NMNH); Los Cipreses, 13–15 Jan 1968, L.E. Peña G., 3 , 1 (NMNH); Constitución, 16 Dec 1976, Gurney & Barria, 5 , 1 (NMNH); Forel Carrizalillo [near Río Maule, ∼25 km E coast], 250 m, 30 Jan–5 Feb 1980, L.E. Peña G., 13 , 7 (NMNH); Río Pinotalca [8 km N Empedrado], 100 m, 29–30 Jan 1981, L.E. Peña G., 1 , 2 (NMNH).

PCIA. VALDIVIA: Enco, 26 Feb 1955, L.E. Peña G., 2 , 1 (CNC); Rincon de Piedra, ∼30 km SE Valdivia, 50 m, 24–25 Feb 1979, Davis et al., 2 (NMNH); same, but 180 m, 24 Sep 1981, Nielsen et al., 3 (ZMC); same, but 14 Dec 1981, 23 (ZMC); S Valdivia, 23 Oct 1969, Flint & Barria, 3 (NMNH); 30 km S Valdivia, 13 Jan 1951, Ross & Michelbacher, 8 , 6 (CAS); Río Bueno, 16 Feb 1978, L.E. Peña G., 1 (NMNH); 8 mi [∼13 km] E Río Bueno, 15 Jan 1951, Ross & Michelbacher, 1 , 1 (CAS); Río Bueno (Margen N), W Río Bueno City, 14–17 Feb 1978, L.E. Peña G., 26 , 11 (NMNH); W Río Bueno, 3–4 Mar 1987, L.E. Peña G., 2 , 3 (NMNH); Choshuenco, 20 Feb 1978, L.E. Peña G., 1 (NMNH); Puerto Fuy, 16–19 Feb 1978, L.E. Peña G., 31 , 28 (NMNH); Lago Calafquén, 10–11 Mar 1984, L.E. Peña G., 5 , 5 (NMNH); Huilo-Huilo, 9–10 Mar 1984, L.E. Peña G., 4 , 4 (NMNH); Rinihue, 7–8 Mar 1984, L.E. Peña G., 1 (NMNH); [Lago] Pellaifa [39°36'S; 71°58′W, ∼600 m], mountain brook, 22 Jan 1958, J. Illies, 1 , 4 (NMNH); Llifen, Lago Ranco, 4–5 Mar 1967, T. Cekalovic K., 4 , 9 (NMNH); Chollinco, Lago Ranco, 5 Mar 1967, T. Cekalovic K., 5 , 2 (NMNH); Río Calcurrupe [40°13′S; 72°17′W], 4 Mar 1967, T. Cekalovic K., 2 , 1 (NMNH).

PCIA. VALPARAÍSO: Cerro Vizcachas, Dec 1982, Madariaga & Escobar, 2 , 5 (NMNH); Marga-Marga, Colliguay, 14–15 Mar 1964, L.E. Peña G., 1 , 5 (NMNH); Quilpué, 13–14 Mar 1964, L.E. Peña G., 2 , 1 (NMNH).