Smicridea (Smicridea) manzanara Flint & Denning 1989

Smicridea (Smicridea) manzanara

Although related to penai and pucara, this species is the most easily recognized of the three. The very long lateral plates of the phallus and, especially, the long slender dorsolateral lobes of the phallus are diagnostic. The differences in the females of the three species are rather slight. The deep, rounded clasper receptacle, with a large, reticulate surface area anteriad in combination with the distinctive anterior knobs of the internal plate offer the best means of identification.

ADULT.—Length of forewing, 10–11.5 mm. Color pale brown; appendages slightly paler, antennae annulate; forewing pale brown marked with darker brown flecks, generally producing an indistinct pattern. Eye of male in frontal aspect distinctly less in diameter than half of interocular width. Anterolateral process of fifth sternum distinctly larger than segment; with 2 pairs of internal sacs, each slightly shorter than segment within which it lies.

Male Genitalia: Ninth segment with anterior margin nearly vertical. Tenth tergum elongate with apex divided apicomesally; tergite with tip rounded in dorsal, upturned and pointed in lateral aspect. Clasper with basal segment slightly inflated apicad, paralled-sided basally; apical segment short, bluntly pointed. Phallus tubular, base at 110° angle to stem, width of basal opening less than 3 times the width of narrowest part of stem, apex inflated; lateral plate at least 3 times as long as broad, ventral margin confluent with venter of stem, in ventral aspect with mesobasal shoulder distinct, shallowly cupped, with dorsal and ventral margins deeply concave, dorsolateral lobe long and slender, nearly attaining apex of lateral plate.

Female Genitalia: Eighth sternite elongate, inner margin sinuate, posteromesal angle produced and truncate, anterolateral angle produced, obliquely truncate. Ninth tergum produced ventrad and anteriad, very broad laterally. Clasper receptacle with inner surface strongly reticulate, produced mesally and posteroventrally, deep and pocket-like, with a large clasper groove anteriad; in lateral aspect with a deep ventral overhang on the surface of which is an oblique dark mark. Vaginal sclerites lightly sclerotized; posterior sclerite, trans4 erse, with a darkened posteromesal lobe, anterior sclerite, trans4 erse, darkened posteromesally. Internal plate darkened, with lateral arms produced posteriad and bearing distinct dark knobs from anterior margin.

MATERIAL EXAMINED.—Holotype, male: CHILE: PCIA. MALLECO: Rí Manzanares [∼10 km W Purén], 2 Jan 1966, Flint & Cekalo4 ic. Type NMNH.

Paratypes: CHILE: PCIA. ARAUCO: Butamalal [Nahuelbuta Mountains, 37°51′S; 73°12′W], 23–31 Jan 1954, L.E. Peña G., 1 (CNC); Caramávida, 25–31 ec 1953, L.E. Peña G., 1 (CNC); same, but 17–19 Oct 1969, Flint & Barria, 4 , 10 (NMNH). PCIA. CACHAPOAL: Pichi-Alhué, 25–27 Nov 1967, L.E. Peña G., 1, 1 (NMNH). PCIA. CAQUENES, near coastal stream, 17.5 km S Curanipe, 50 m, 25 Jan 1979, Davis et al., 3 , 1 (NMNH); Alto Tregualemu, ∼20 km SE Chovellén, 500 m, 26–27 Jan 1979, Davis et al., 9 (, 4 (NMNH); same, but 1–3 Dec 1981, D.R. Davis, 1 (NMNH); same, but 27–28 Jan 1981, L.E. Peña G., 1 (NMNH); Paso Garcia, ∼23 km W Cauquenes, 300 m, 29–30 Nov 1981, D.R. Davis, 1 (NMNH); Cayurranquil, W Caquenes, 400 m, 23–31 Jan 1981, L.E. Peña G., 1 (NMNH). PCIA. CONCEPCIÓN: Fundo Pinares [about 10 km E Concepción on south side of Río Bío-Bío], 30 Dec 1965, Flint & Cekalovic, 4 (NMNH); same, but 18 Dec 1964, T. Cekalovic K., 5 (NMNH); same, but 18 Feb 1967, 2 (NMNH); Lirquén, 17 Oct 1965, T. Cekalovic K., 1 (NMNH); Quebrada Honda, N Lirquén, 31 ec 1965, Flint & Cekalovic, 7 , 4 (NMNH); same, but 5 Feb 1966, T. Cekalovic K., 5 (NMNH); same, but 14–15 Oct 1969, Flint & Barria, 26 , 5 (NMNH). PCIA. CORDILLERA: El Manzano, 26 Oct 1951, L. E. Peña G., 1 (CNC); El Alfalfal, 29 Feb 1968, Flint & Peña, 2 (NMNH). PCIA. CURICO: Estero La Jaula, Los Queñes, 4–18 Jan 1964, L.E. Peña G., 19 , 23 (NMNH). PCIA. MALLECO: same data as holotype, 16 , 3 (NMNH); Parque Nacional Contulmo, 2 Jan 1966, Flint & Cekalovic, 1 (NMNH); same, but 20 Feb 1967, T. Cekalovic K., 4 (NMNH); same, but 19 Oct 1969, Flint & Barria, 1 (NMNH); Angol, Los Alpes, 650 m, 17 Mar 1979, Nielsen et al., 2, 2 (ZMC); Río Blanco, 5–27 Mar 1950, L.E. Peña G., 6 , 1 (CNC); same, but 21–24 Feb 1954, l (CNC). AREA METROPOLITANA: Peñalolen, 1 Oct 1950, L.E. Peña G., 3 (CNC); Quebrada Macul, 2 Mar 1950, L.E. Peña G., 2 (CNC); Macul, 11 Mar 1949, L.E. Peña G., 1 (CNC). PCIA. PETORCA: LOS Hornos, Guaguen, 2–3 Oct 1986, L.E. Peña G., 2 , 2 (NMNH). PCIA. QUILLOTA: Estero Marga-Marga, near Perales, 9 Mar 1968, Flint & Peña G., 4 (NMNH). PCIA. TALCA: Tonlemo, 15 Feb 1950, L.E. Peña G., 27 , 2 (CNC); same, but 14–15 Dec 1984, 1 (NMNH); Constitución, 16 Dec 1976, Gurney & Barria, 1 (NMNH); Forel Carrizalillo [near Rí Maule, ∼25 km E coast], 250 m, 30 Jan–5 Feb 1981, L.E. Peña G., 1 (NMNH). PCIA. VALPARAÍSO: Cerro Vizcachas, Dec 1982, R. Madariaga, 1 (NMNH).