Nepenthes benstonei /nɪˈpɛnθiːz bɛnˈstoʊniaɪ/ is a tropical pitcher plant endemic to Peninsular Malaysia, where it grows at elevations of 150–1350 m above sea level.[3][10] The specific epithet benstonei honours botanist Benjamin Clemens Stone, who was one of the first to collect the species.[1]
In their 1997 monograph, "A skeletal revision of Nepenthes (Nepenthaceae)", Matthew Jebb and Martin Cheek tentatively referred specimens collected from Bukit Bakar, near Macang, Kelantan, to N. sanguinea.[7] These were Stone & Chin 15238, deposited at Universiti Kebangsaan Malaysia near Kuala Lumpur (KLU), and Shah & Shukor 3168, also held at KLU as well as the Forest Research Institute of Malaysia in Kepong (KEP). They noted that the plants exhibited some unusual morphological features, such as larger leaves and decurrent, almost petiolate leaf bases, suggesting that they might represent an as-yet undescribed taxon.[7]
Field studies confirmed that the taxon represented a separate species, and it was formally described as N. benstonei in 1999 by Charles Clarke.[1][11]
The holotype of N. benstonei, Clarke s.n., was collected by Charles Clarke on 24 July 1998, on Bukit Bakar in Kelantan at an altitude of between 450 and 550 m. It is deposited at the Forest Research Institute of Malaysia in Kepong (KEP). Isotypes are held at Herbarium Bogoriense (BO), the Royal Botanic Gardens, Kew (K), the National Herbarium of the Netherlands in Leiden (L), the Forest Department in Sandakan (SAN), and the Singapore Botanic Gardens (SING).[3][12]
Although only described towards the very end of the twentieth century, N. benstonei was probably first collected in July 1911 by Henry Nicholas Ridley on Mount Tahan in Pahang. The Ridley 16097 series comprises three herbarium sheets: one deposited at the Singapore herbarium and two at Kew. The former consists of a climbing stem fragment with two upper pitchers and two female inflorescences. The two sheets at Kew are barcoded K000651565 (climbing stem with immature female inflorescence but no pitchers) and K000651564 (climbing stem with upper pitchers), and differ significantly in morphology.[3] Ridley referred the Ridley 16097 material to N. singalana.[3]
For his seminal monograph of 1928, "The Nepenthaceae of the Netherlands Indies", B. H. Danser examined the sheet of Ridley 16097 held in Singapore, lumping it with the variable N. alata from the Philippines.[2] Danser briefly mentioned this specimen in his discussion of N. alata:[2]
The specimen recorded by me from the Malay Peninsula deviates more [from N. alata], especially by the long, narrow inflorescence and 2-flowered pedicels, but also in the Philippines and Sumatra forms with 2-flowered pedicels have been found (Ramos 14650, Lörzing 11603).
Danser also treated N. eustachya from Sumatra in synonymy with N. alata, giving rise to a taxon with a puzzling geographical distribution: widespread across Sumatra and the Philippines, apparently very rare in the Malay Peninsula (Ridley 16097 being the sole record), and completely unknown from Borneo and the other major islands of the Malay Archipelago.[2][12]
In 1990, Ruth Kiew identified Ridley 16097 as belonging to N. gracillima. Kiew explained the apparent near-absence of N. alata from Peninsular Malaysia as follows:[6]
Why has this species not been recollected since 1911 in spite of several botanical expeditions to G. Tahan since then? The answer is quite simple, Ridley's specimen (16097) actually belongs to N. gracillima. Although N. alata Blanco is superficially similar to N. gracillima in its narrow leaf blade which has an attenuate base, the pitchers of these two species are distinct. Those of N. alata are broader (about 4 cm wide) and are distinctly bulbous towards the base compared with the very slender, non-bulbous pitchers of N. gracillima that are 1.5 to 3 cm wide. Nor does the G. Tahan specimen have truly petiolate leaves - its leaf base is narrow and has rolled up during drying. Ridley's specimen from G. Tahan determined by Danser as N. alata bears pitchers typical of N. gracillima and examination of the Tahan population in the field leaves no doubt that it belongs to this species.
Jebb and Cheek supported this interpretation in their 1997 monograph, "A skeletal revision of Nepenthes (Nepenthaceae)",[7] and in their 2001 treatment for Flora Malesiana, "Nepenthaceae".[8] They also restored N. eustachya as a separate species, making N. alata a Philippine endemic once again — a circumscription that has been accepted by subsequent authors.[12][13]
However, in his 2001 book, Nepenthes of Sumatra and Peninsular Malaysia, Charles Clarke disagreed with Kiew's identification of Ridley 16097 as N. gracillima. He noted that the inflorescences of both N. gracillima and the closely related N. ramispina are very short, rarely exceeding 10 and 20 cm, respectively. Both female inflorescences of Ridley 16097 have a long peduncle and rachis, each exceeding 20 cm in length. In addition, the flowers of these species are almost always borne on pedicels, unlike those of Ridley 16097, which mostly have two-flowered partial peduncles.[14] Furthermore, the shape of the lamina is unlike that of N. gracillima or N. ramispina; despite being narrow and lanceolate, it is proportionately considerably longer and has a much narrower, almost sub-petiolate base. Kiew partly attributed the narrower leaf bases of Ridley 16097 to a preservation artefact, but Clarke stated that this explanation could not fully account for the differences. He also noted that the specimen exhibits a decurrent leaf attachment. Taking all of these morphological features into account, Clarke felt that Ridley 16097 most likely represented a specimen of N. benstonei.[12][15] At the time, he believed that Kiew had grouped both N. benstonei and N. ramispina with N. gracillima.[12] But in a 2012 revision of the Nepenthes of Mount Tahan, which included a reappraisal of the taxonomically confused N. alba and N. gracillima, Clarke and Ch'ien Lee concluded that Kiew's concept of N. gracillima had encompassed N. alba, N. benstonei, and N. gracillima.[3] Clarke and Lee also showed that Ridley 16097 represents a mixed collection and that only two of the three sheets belong to N. benstonei; the Kew specimen barcoded K000651564 is representative of N. alba.[3]
In addition to the two sheets of Ridley 16097, populations of N. benstonei from Mount Tahan are represented by the specimen Holttum 20643, held at the herbarium of the Singapore Botanic Gardens (SING).[3] A number of specimens collected from Terengganu also belong to N. benstonei. These are Shah et al. 3274, which is deposited at the Forest Research Institute of Malaysia in Kepong, and Shah et al. 3283, held at the Universiti Kebangsaan Malaysia near Kuala Lumpur.[12][16]
Specimens from peninsular Thailand originally assigned to N. benstonei in Cheek and Jebb's 2001 monograph, "Nepenthaceae",[8] have since been identified as belonging to a new species, N. thai.[9]
Nepenthes benstonei is a climbing plant. The stem, which may be branched, can attain a length of 10 m[10] and is up to 0.6 cm in diameter. Internodes are cylindrical and up to 15 cm long.[12]
Leaves are coriaceous and sessile to sub-petiolate. The lamina is usually broadly linear-lanceolate in shape, but may also be slightly spathulate. Its base is a broad, amplexicaul sheath with decurrent margins. The lamina can reach 60 cm in length and 9 cm in width. It has a rounded to acute apex. The margins of the lamina usually meet the tendril unequally on both sides, being up to 3 mm apart. Three to five longitudinal veins are present on either side of the midrib. Pinnate veins are almost indistinct. Tendrils are up to 60 cm long.[12]
Rosette and lower pitchers reach 20 cm in height[16] and 5 cm in width. They are ovoid in the lower part and cylindrical above, with a pronounced hip in the middle. A pair of fringed wings (≤4 mm wide) runs the whole length of the pitcher cup. The pitcher mouth is round to ovate and oblique throughout. The peristome is up to 6 mm wide and bears very small but distinct teeth along its inner margin. The pitcher lid is ovate and lacks appendages. It bears a short but distinct keel and often has a very broad insertion. A simple or bifurcate spur (≤12 mm long) is inserted near the base of the lid.[12]
Upper pitchers are similar in most respects to their lower counterparts. They are up to 20 cm high[16] and 3 cm wide. They are infundibular in the lowermost part, narrowly ovoid in the next part, and cylindrical above. The peristome lacks teeth in upper pitchers. The lid is narrower and has a less obtuse apex. The spur is simple and much smaller, reaching only 5 mm in length.[12]
Nepenthes benstonei has a racemose inflorescence. The peduncle is up to 20 cm long and the rachis up to 30 cm long. The first flower is generally borne on a pedicel, sometimes with a simple, lanceolate bracteole (≤1.5 cm long). Subsequent flowers are produced on pedicels or two-flowered partial peduncles, which lack bracteoles. Sepals are ovate and around 4 mm long. Male inflorescences usually bear around twice as many flowers as female ones. N. benstonei is one of the few Nepenthes species known to produce multiple inflorescences concurrently on a single stem. Two to three are usually produced, originating from sequential nodes at the top of the stem. This unusual reproductive habit has also been observed, although much more rarely, in N. alba, N. ampullaria, N. attenboroughii, N. rigidifolia, N. sanguinea, and N. thai.[9][12][13] It is seen even more frequently in N. philippinensis.[13]
The stem and lamina have a sparse indumentum of simple white hairs. Short, branched reddish-brown hairs line the margins of the lamina. The outer surfaces of the pitchers bear a sparse covering of short, branched red hairs. The same hairs are more densely present on the margins of the lid and upper part of the pitcher directly below the peristome. Immature inflorescences have an indumentum of short white and red hairs throughout.[12]
The stem and leaves of N. benstonei bear a thick, waxy cuticle that often gives a whitish-blue sheen to the lamina and pitchers. Inflorescences are distinctly waxy throughout.[12]
No infraspecific taxa of N. benstonei have been described.[12]
Nepenthes benstonei is endemic to Peninsular Malaysia. It is known with certainty only from the summits of low hills in Kelantan and northern Terengganu,[12][16] and from Mount Tahan in Taman Negara, Pahang.[3] The species has a relatively wide altitudinal range of 150[10] to 1350 m above sea level.[3]
Nepenthes benstonei grows terrestrially among open, secondary vegetation, where it is exposed to direct sunlight. It is very abundant near the summit of Bukit Bakar, where it grows on cuttings beside a paved road leading to a Telekom Malaysia station at the summit.[12][16] There, its altitudinal distribution appears to be restricted to 450–600 m.[12]
The species is also present on Mount Tahan, which at 2187 m is the highest mountain in Peninsular Malaysia. Its altitudinal range on Mount Tahan is known to extend from 800 to 1350 m. It is common on the mountain's lower slopes and can be seen along the western summit route from Sungai Relau, particularly on the tops of steep ridges at around 800–1200 m.[nb 1] It has been recorded growing along the western trail itself and from other disturbed sites, including areas affected by landslides. Plants have also been observed in dense forest, but these bear comparatively few pitchers. Though there were no confirmed reports of N. benstonei from Mount Tahan prior to 2012, the species's presence there is attested by much older herbarium material.[3]
Although the extent of its range is uncertain, N. benstonei appears to have a secure future in the wild as the type locality lies within a protected area and the species's unremarkable appearance means over-collection does not pose a serious threat.[12]
In his description of N. benstonei, Charles Clarke noted two characteristics that he considered unique among Nepenthes. These were the production of multiple inflorescences and the presence of a thick, waxy cuticle on the leaves.[1] Subsequent field studies have shown that the former is not unique to N. benstonei, but also occasionally occurs in other Nepenthes. Likewise, a number of other species, such as N. hirsuta from Borneo, are known to produce a waxy cuticle, although it is less developed than in N. benstonei.[12] Otherwise, N. benstonei lacks remarkable characteristics and is distinguished from related species on the basis of its stem, leaves, peristome, lid, indumentum, and glands of the digestive zone.[1]
Nepenthes benstonei appears to be related to N. sanguinea, which is also native to Peninsular Malaysia. It can be distinguished on the basis of its significantly larger leaves, which are often sub-petiolate and differ in shape. Nepenthes benstonei also has longer tendrils and a denser indumentum. The presence of teeth on the peristome of lower pitchers and of a thick, waxy cuticle on the leaves also serve to distinguish these taxa. In addition, herbarium specimens of N. benstonei tend to dry to a lighter colour than those of N. sanguinea.[12]
The pitchers of N. benstonei also resemble those of N. smilesii from Indochina. Clarke suggests that N. benstonei may represent an evolutionary link between the Nepenthes taxa of Indochina and Peninsular Malaysia.[16] Nepenthes benstonei also superficially resembles N. macrovulgaris from Borneo. It differs in producing multiple inflorescences, which are longer than those of N. macrovulgaris and bear one- or two-flowered partial peduncles, as opposed to exclusively two-flowered in the latter. The waxy coating of its leaves also separates these species.[12] Nepenthes benstonei has also been compared to N. albomarginata, although the presence of a white band below the peristome, which gives the latter its name, makes identification easy.[12] Upper pitchers of N. benstonei could be confused with those of N. mirabilis, although all other parts of the plant have little in common.[12]
In 2001, Charles Clarke performed a cladistic analysis of the Nepenthes species of Sumatra and Peninsular Malaysia using 70 morphological characteristics of each taxon. The resultant cladogram placed N. benstonei in an unresolved polytomy at the base of the Montanae/Nobiles clade, together with N. rhombicaulis.[12]
Only one natural hybrid involving N. benstonei is known.[13] A single example of N. benstonei × N. mirabilis was discovered by Andrew Hurrell at the foot of Bukit Bakar, where the two species occur sympatrically.[12][16]
Nepenthes benstonei /nɪˈpɛnθiːz bɛnˈstoʊniaɪ/ is a tropical pitcher plant endemic to Peninsular Malaysia, where it grows at elevations of 150–1350 m above sea level. The specific epithet benstonei honours botanist Benjamin Clemens Stone, who was one of the first to collect the species.
Nepenthes benstonei C.Clarke è una pianta carnivora della famiglia Nepenthaceae[1], endemica della Malaysia Peninsulare, dove cresce a 150–1350 m.
Nepenthes benstonei C.Clarke è una pianta carnivora della famiglia Nepenthaceae, endemica della Malaysia Peninsulare, dove cresce a 150–1350 m.
Nepenthes benstonei /nɪˈpɛnθiːz bɛnˈstoʊni.aɪ/ là một loài nắp ấm đặc hữu của bán đảo Malaysia, nơi nó phát triển ở độ cao 150–1350 m trên mực nước biển.[3][10] Tên loài benstonei dùng để ghi danh nhà thực vật học Benjamin Clemens Stone, một trong những người đầu tiên để thu thập loài này.[1]
Nepenthes benstonei /nɪˈpɛnθiːz bɛnˈstoʊni.aɪ/ là một loài nắp ấm đặc hữu của bán đảo Malaysia, nơi nó phát triển ở độ cao 150–1350 m trên mực nước biển. Tên loài benstonei dùng để ghi danh nhà thực vật học Benjamin Clemens Stone, một trong những người đầu tiên để thu thập loài này.
本斯通猪笼草（学名：Nepenthes benstonei）是西马来西亚特有的热带食虫植物，生长于海拔150米至1350米的地区。[3][10]其种加词“benstonei”来源于植物学家本杰明·克莱门斯·斯通，他采集到了本斯通猪笼草的第一份标本。[1]
马修·杰布和马丁·奇克在其1997年发表的专著《猪笼草属（猪笼草科）的框架性修订》中，提到了采集自吉兰丹州马樟（Machang）附近的巴卡山（Bukit Bakar）的血红猪笼草（N. sanguinea）标本。其中，编号为“Stone & Chin 1523”，存放于吉隆坡附近的马来西亚国民大学，编号为“Shah & Shukor 3168”的标本存放于甲洞的马来西亚森林研究院（Forest Research Institute of Malaysia）。他们注意到这个类群表现出的一些不寻常的形态特征，如叶片大型，基部具叶柄且下延，这使他们认为其可能是一个尚未描述的类群。[7]
实地考察证实，这个类群的确是一个独立的物种。1999年，查尔斯·克拉克对本斯通猪笼草进行了描述。[1][11]
1998年7月24日，查尔斯·克拉克在吉兰丹州的巴卡尔山海拔450米至550米的地区采集了本斯通猪笼草的模式标本，其编号为“Clarke s.n.”。此标本存放于甲洞的马来西亚森林研究院。同模标本存放于茂物植物园、英国皇家植物园、荷兰国家植物标本馆、山打根林业局和新加坡植物园。[3][12]
尽管本斯通猪笼草在二十世纪末才被描述，但早在1911年7月，亨利·尼古拉斯·里德利就首次在彭亨的大汉山采集到了本斯通猪笼草的标本。这个编号为“Ridley 16097”号标本分为三部分：其中一部分位于新加坡植物园，另外两部分位于邱园。前者包含一段带2个上位笼及2个雌性花序的攀援茎。位于邱园的后者的编码分别为“K000651565”（包含一段带未程序雌性花序且无捕虫笼的攀援茎）及“K000651564”（包含一段带上位笼的攀援茎），两者形态明显不同。[3]但里德利认为“Ridley 16097”号标本为欣佳浪山猪笼草（N. singalana）。[3]
B·H·丹瑟在其1928年的开创性著作《荷属东印度群岛的猪笼草科植物》中，将采自新加坡的“Ridley 16097”号标本归入了分布于菲律宾的翼状猪笼草（N. alata）。B·H·丹瑟在其专著对翼状猪笼草的讨论中关于该标本是这样写道的：[注 1][2]
“ 来源于西马来西亚的该标本相对（翼状猪笼草）而言，其花序特别的长而窄，且花梗带2朵花，虽然也发现菲律宾及苏门答腊的变型带2朵花的。（Ramos 14650，Lörzing 11603） ”B·H·丹瑟将也将产于苏门答腊岛的真穗猪笼草（N. eustachya）归入了翼状猪笼草，这使得这一类群的地理分布变得非常的奇怪：其广布于苏门答腊及菲律宾，且奇怪的出现于西马来西亚（“Ridley 16097”号标本为唯一记录），而其间的婆罗洲及其他主要岛屿都均未发现。[2][12]
1990年，鲁思·裘（Ruth Kiew）将“Ridley 16097”号标本归入瘦小猪笼草（N. gracillima），并解释了为什么翼状猪笼草明显不存在于西马来西亚：[注 2][6]
马修·杰布和马丁·奇克在其1997年发表的专著《猪笼草属（猪笼草科）的框架性修订》及《猪笼草科》中支持该观点。[7]他们也将真穗猪笼草独立为一个物种，使翼状猪笼草再次被划归为菲律宾特有的物种——之一处理得到了随后其他学者的认同。[12][13]
但在查尔斯·克拉克2001年的著作《苏门答腊岛与西马来西亚的猪笼草》中并不赞同鲁思·裘将“Ridley 16097”号标本归入瘦小猪笼草的处理。他指出，瘦小猪笼草及其密切相关的岔刺猪笼草（N. ramispina）的花序非常短，很少超过10厘米。而“Ridley 16097”号标本的花序都具有较长的总花梗和花序轴，都可超过20厘米。且瘦小猪笼草与岔刺猪笼草的花梗只带一朵花，而“Ridley 16097”号标本的花梗大部分带两朵花。[14]此外，其叶片的形状也与瘦小猪笼草和岔刺猪笼草不同。鲁思·裘认为“Ridley 16097”号标本的叶片呈现出狭窄状部分是由于制作标本时造成的，但查尔斯·克拉克认为这不能完全解释其差异。他还指出这个种群具有下延的槽状叶柄。考虑到这些形态特征，查尔斯·克拉克感觉“Ridley 16097”号标本更可能是本斯通猪笼草。[12][15]查尔斯·克拉克当时相信鲁思·裘已将本斯通猪笼草和岔刺猪笼草都归入了瘦小猪笼草中。[12]但在2012年，查尔斯·克拉克与李乾对大汉山的猪笼草的分类学修订中重新评估了白猪笼草（N. alba）及瘦小猪笼草的分类，他们认为鲁思·裘对瘦小猪笼草的划分已包含了白猪笼草、本斯通猪笼草及瘦小猪笼草。[3]他们还表明“Ridley 16097”号标本是一个混合的标本，三份中仅两份为本斯通猪笼草，而邱园编号为“K000651564”的标本实际为白猪笼草。[3]
除了“Ridley 16097”号标本中属于本斯通猪笼草的两份标本外，新加坡植物园还存放着采集自大汉山，编号为“Holttum 20643”的本斯通猪笼草标本。[3]采集自丁加奴的许多标本也属于本斯通猪笼草。其中编号为“Shah et al. 3274”的标本存放于马来西亚森林研究院；编号为“Shah et al. 3283”的标本存放于吉隆坡附近的马来西亚国民大学。[12][16]
马修·杰布和马丁·奇克2001年发表的专著《猪笼草科》中[8]，将一个产自泰国的猪笼草归入本斯通猪笼草，之后其被认定一个新的物种——泰国猪笼草（N. thai）。[9]
本斯通猪笼草为藤本植物，可分叉。茎可长达10米[10]，直径可达0.6厘米，呈圆柱形。节间距可达15厘米。[12]
尚未发现本斯通猪笼草的种下类群。[12]
本斯通猪笼草的叶片革质，无柄至具小叶柄。叶片通常为宽线形至披针形，也可能略呈匙形。叶基宽大，抱茎，边缘下延。叶片可长达60厘米，宽至9厘米。叶尖圆至急尖。笼蔓与叶片衔接处，叶片两侧边缘通常不等侧，可有3毫米的差距。中脉的两侧各有3至5条纵脉。羽状脉大多不明显。笼蔓可长达60厘米。[12]
本斯通猪笼草的下位笼可高达20厘米[16]，宽至5厘米。其下半部为卵形，上半部为圆柱形，中部具明显的笼肩。腹面具一对笼翼，不宽于4毫米。笼口为圆形至卵形，倾斜。唇可宽达6毫米，具有小型的唇齿。笼盖为卵形，无附属物。具有一个短但明显的笼盖骨，常非常的宽大。笼盖基部的后方具有一根不分叉或分叉的笼蔓尾，不长于12毫米。[12]
本斯通猪笼草的上位笼较小，大部分结构类似于下位笼。可高达20厘米，宽至3厘米。下部为漏斗形，中部为窄卵形，上部为圆柱形。唇无唇齿。笼盖较窄，末端为钝尖。笼蔓尾不分叉，大大缩小，仅可长达5毫米。[12]
本斯通猪笼草的花序为总状花序。总花梗可达20厘米，花序轴可达30厘米。最顶端的花梗带一朵花，偶尔具披针形的苞片（长达可达1.5厘米）。基部花梗一朵或两朵花，无苞片。萼片为卵形，长约4毫米。雄性花序花朵数量约为雌性的两倍。本斯通猪笼草是猪笼草属中少数可以在一条茎上产生多个花序的猪笼草。茎顶部节点通常可发出2至3条花序。这种不同寻常的生殖习性也出现其他一些猪笼草属物种上，但更为罕见，在包括苹果猪笼草（N. ampullaria）、硬叶猪笼草（N. rigidifolia）、血红猪笼草（N. sanguinea）和泰国猪笼草。[9][12][13]菲律宾猪笼草（N. philippinensis）则较为常见。[13]
本斯通猪笼草的茎部和叶片披被着稀疏的白色毛被。在叶片的边缘具有红褐色的分叉短毛被。捕虫笼的外表面覆盖着稀疏的红色分叉短毛被。同样的毛被还更密集的出现于笼盖的边缘和唇下方的笼身上。发育中的花序具有白色或红色的短毛被。[12]
本斯通猪笼草的茎和叶片具很厚的蜡质表皮，其使得叶片和捕虫笼具有蓝白色的光泽。花序整体也密布蜡质。[12]
本斯通猪笼草是西马来西亚特有的热带食虫植物。仅能确定其分布于吉兰丹州低地山丘的顶部和丁加奴北部[12][16]，及大汉山国家公园的大汉山[3]。其分布于海拔150米[10]至1350米[3]处。
但其分布范围可能更为广泛，部分标本采集于彭亨的大汉山国家公园。[12][16]
本斯通猪笼草陆生于开阔，有阳光直射的次生植被中。在巴卡山顶峰附近有大量的分布，其生长于去往山顶马来西亚电讯公司基站的道路两旁。在此其多存在于海拔450至600米的地区内。[12][16]
其也分布于西马来西亚的大汉山海拔800米至1350米处。其多见于山脉低海拔的斜坡上，也可见于惹拉河（Sungai Relau）去往西部山峰的沿途，特别是海拔800米至1200米的陡峭山脊处。[注 3]再往西，其生长就会受到包括滑坡等地质灾害的影响。本斯通猪笼草的植株多位于茂密的森林中，但很少出现捕虫笼。虽然在2012年前并没有确认本斯通猪笼草存在于大汉山，但许多以往的标本证明了其的存在。[3]
虽然其分布范围还不确定，但野外的本斯通猪笼草在未来一段时间内还是较为安全的。其模式产地处于保护区内，并且该物种的外观并不惹眼，因此遭受到过度采集威胁的可能性较小。[12]
在查尔斯·克拉克关于本斯通猪笼草的描述中，指出了两个区别于其他猪笼草的特点。即是其可产生多个花序，及叶片具有很厚的蜡质表皮。[1]但后来的实地考察发现，前者并不是本斯通猪笼草特有的，其也偶尔出现于其他猪笼草上。同样，其他一些猪笼草也具有蜡质表皮，如婆罗洲的刚毛猪笼草（N. hirsuta），但其不如本斯通猪笼草的发达。[12]所以本斯通猪笼草并没有标志性的特征，要将本斯通猪笼草和与之具有亲缘关系的物种区分开来，需要从茎、叶片、唇、笼盖、毛被和消化腺体区等各个方面综合比较。[1]
本斯通猪笼草表现出与西马来西亚特有的血红猪笼草之间存在着较为密切的亲缘关系。它们的区别在于本斯通猪笼草的叶片更大些且形状不同，并通常具有小叶柄。笼蔓也较长，并覆盖着密集的毛被。下位笼具唇齿，且叶片具有较厚的蜡质角质层。这些特征都有助于将其与血红猪笼草相区别。此外，本斯通猪笼草干燥标本的颜色要比血红猪笼草的更浅。[12]
本斯通猪笼草的捕虫笼形态也类似于存在于中南半岛的斯迈尔斯猪笼草（N. smilesii）。查尔斯·克拉克认为本斯通猪笼草在进化树中可能处于中南半岛类群和西马来西亚类群之间。[16]
本斯通猪笼草在表面上也类似于婆罗洲特有的大型平庸猪笼草（N. macrovulgaris）。它们的区别在于本斯通猪笼草可产生多个花序，且都长于大型平庸猪笼草的花序。花梗带1至2朵花，而后者均带2朵花。此外，大型平庸猪笼草的叶片不具有蜡质表皮。[12]
本斯通猪笼草也类似于白环猪笼草（N. albomarginata），但其不具有后者唇下特有的白色带状附属物，所以将其区别开来并不困难。[12]
本斯通猪笼草的上位笼可能会与奇异猪笼草（N. mirabilis）的上位笼相混淆，但植株的其他部分并没有什么共同之处。[12]
2001年，查尔斯·克拉克对产自苏门答腊及西马来西亚的猪笼草进行了分支系统学分析，利共用了70个形态特征。但本斯通猪笼草与菱茎猪笼草（N. rhombicaulis）一起还未确定到底是归入山地分支（Montanae）还是高贵分支（Nobiles）分支。[12]
仅发现了一个关于本斯通猪笼草的自然杂交种。[13]安德鲁·赫里尔（Andrew Hurrell）于巴卡山山脚本斯通猪笼草与奇异猪笼草分布地相互重叠的地区发现了它们的自然杂交种。[12][16]
寬葉豬籠草
源小猪笼草
拟翼状猪笼草
翼状猪笼草
白猪笼草
白环猪笼草
阿札潘山猪笼草
苹果猪笼草
安达曼猪笼草
昂嘎桑猪笼草
附盖猪笼草
阿金特猪笼草
马兜铃猪笼草
阿滕伯勒猪笼草
贝卡利猪笼草
贝里猪笼草
本斯通猪笼草
二齿猪笼草
波哥猪笼草
邦苏猪笼草
博世猪笼草
豹斑猪笼草
伯克猪笼草
风铃猪笼草
塞西尔猪笼草
象岛猪笼草
陈氏猪笼草
熙德猪笼草
圆盾猪笼草
柯普兰猪笼草
丹瑟猪笼草
N. adnata
N. abgracilis
N. abalata
N. alata
N. alba
N. albomarginata
N. alzapan
N. ampullaria
N. andamana
N. angasanensis
N. appendiculata
N. argentii
N. aristolochioides
N. attenboroughii
N. beccariana
N. bellii
N. benstonei
N. bicalcarata
N. bokorensis
N. bongso
N. boschiana
N. burbidgeae
N. burkei
N. campanulata
N. ceciliae
N. chang
N. chaniana
N. cid
N. clipeata
N. copelandii
N. danseri
迪安猪笼草
密花猪笼草
上位猪笼草
滴液猪笼草
疑惑猪笼草
爱德华猪笼草
鞍型猪笼草
附生猪笼草
真穗猪笼草
绝灭猪笼草
艾玛猪笼草
法萨猪笼草
杏黄猪笼草
暗色猪笼草
甘通山猪笼草
无毛猪笼草
有腺猪笼草
小花猪笼草
小猪笼草
瘦小猪笼草
裸瓶猪笼草
钩唇猪笼草
汉密吉伊坦山猪笼草
赫姆斯利猪笼草
刚毛猪笼草
粗毛猪笼草
霍尔登猪笼草
胡瑞尔猪笼草
无刺猪笼草
卓越猪笼草
泉氏猪笼草
N. deaniana
N. densiflora
N. diatas
N. distillatoria
N. dubia
N. edwardsiana
N. ephippiata
N. epiphytica
N. eustachya
N. extincta
N. eymae
N. faizaliana
N. flava
N. fusca
N. gantungensis
N. glabrata
N. glandulifera
N. graciliflora
N. gracilis
N. gracillima
N. gymnamphora
N. hamata
N. hamiguitanensis
N. hemsleyana
N. hirsuta
N. hispida
N. holdeni
N. hurrelliana
N. inermis
N. insignis
N. izumiae
贾桂琳猪笼草
马桶猪笼草
容洪猪笼草
贡布猪笼草
克尔猪笼草
印度猪笼草
奇坦兰山猪笼草
克罗斯猪笼草
空堪达猪笼草
仓田猪笼草
蓝姆猪笼草
熔岩猪笼草
莱昂纳多猪笼草
莱特岛猪笼草
小舌猪笼草
长叶猪笼草
劳氏猪笼草
麦克法兰猪笼草
大叶猪笼草
大型平庸猪笼草
马达加斯加猪笼草
曼塔灵阿汉山猪笼草
马普鲁山猪笼草
马索亚拉半岛猪笼草
大猪笼草
美林猪笼草
小瓮猪笼草
迈克猪笼草
棉兰老岛猪笼草
惊奇猪笼草
奇异猪笼草
N. jacquelineae
N. jamban
N. junghuhnii
N. kampotiana
N. kerrii
N. khasiana
N. kitanglad
N. klossii
N. kongkandana
N. kurata
N. lamii
N. lavicola
N. leonardoi
N. leyte
N. lingulata
N. longifolia
N. lowii
N. macfarlanei
N. macrophylla
N. macrovulgaris
N. madagascariensis
N. mantalingajanensis
N. mapuluensis
N. masoalensis
N. maxima
N. merrilliana
N. micramphora
N. mikei
N. mindanaoensis
N. mira
N. mirabilis
柔毛猪笼草
山地猪笼草
姆鲁山猪笼草
毛律山猪笼草
龙猪笼草
内格罗斯岛猪笼草
新几内亚猪笼草
黑猪笼草
诺斯猪笼草
卵形猪笼草
巴拉望岛猪笼草
圆锥猪笼草
巴布亚猪笼草
盾葉毛豬籠草
伯威尔猪笼草
有柄猪笼草
菲律宾猪笼草
细毛猪笼草
皮托庞猪笼草
宽唇猪笼草
美丽猪笼草
莱佛士猪笼草
馬來王豬籠草
岔刺猪笼草
拉莫斯猪笼草
两眼猪笼草
菱茎猪笼草
硬叶猪笼草
罗伯坎特利猪笼草
罗恩猪笼草
N. mollis
N. monticola
N. muluensis
N. murudensis
N. naga
N. negros
N. neoguineensis
N. nigra
N. northiana
N. ovata
N. palawanensis
N. paniculata
N. papuana
N. peltata
N. pervillei
N. petiolata
N. philippinensis
N. pilosa
N. pitopangii
N. platychila
N. pulchra
N. rafflesiana
N. rajah
N. ramispina
N. ramos
N. reinwardtiana
N. rhombicaulis
N. rigidifolia
N. robcantleyi
N. rowanae
萨马岛猪笼草
血红猪笼草
萨兰加尼猪笼草
辛布亚岛猪笼草
欣佳浪山猪笼草
斯迈尔斯猪笼草
匙叶猪笼草
显目猪笼草
窄叶猪笼草
苏门答腊猪笼草
素叻猪笼草
苏里高猪笼草
塔蓝山猪笼草
坚韧猪笼草
毛盖猪笼草
细猪笼草
泰国猪笼草
高棉猪笼草
多巴猪笼草
托莫里猪笼草
特勒布猪笼草
宝特瓶猪笼草
波叶猪笼草
超基猪笼草
维奇猪笼草
葫芦猪笼草
维耶亚猪笼草
长毛猪笼草
绿猪笼草
佛氏猪笼草
N. samar
N. sanguinea
N. saranganiensis
N. sibuyanensis
N. singalana
N. smilesii
N. spathulata
N. spectabilis
N. stenophylla
N. sumatrana
N. suratensis
N. surigaoensis
N. talangensis
N. tenax
N. tentaculata
N. tenuis
N. thai
N. thorelii
N. tobaica
N. tomoriana
N. treubiana
N. truncata
N. undulatifolia
N. ultra
N. veitchii
N. ventricosa
N. vieillardii
N. villosa
N. viridis
N. vogelii
阿里猪笼草
石龙门猪笼草
坎特利猪笼草
雪线猪笼草
红脉猪笼草
N. × alisaputrana
N. × bauensis
N. × cantleyi
N. × cincta
N. × ferrugineomarginata
哈里猪笼草
虎克猪笼草
基纳巴卢山猪笼草
古晋猪笼草
美翼猪笼草
N. × harryana
N. × hookeriana
N. × kinabaluensis
N. × kuchingensis
N. × merrilliata
妙翼猪笼草
潘丘卢保山猪笼草
梨形猪笼草
沙捞越猪笼草
沙礼花-哈萨猪笼草
N. × mirabilata
N. × pangulubauensis
N. × pyriformis
N. × sarawakiensis
N. × sharifah-hapsahii
毛果猪笼草
宝翼猪笼草
特鲁斯马迪山猪笼草
曾氏猪笼草
红瓶猪笼草
N. × trichocarpa
N. × truncalata
N. × trusmadiensis
N. × tsangoya
N. × ventrata
