Beronaphaenops

Body elongate, head and pronotum narrow, hind body oval, clearly convex on the dorsum (Fig 1). Integument thin, translucent (as being depigmented), moderately shiny; short, sparse and uniform pubescence presents on both the dorsal and ventral surfaces, tibiae and tarsi densely pubescent. Microsculpture distinctly impressed on the head and elytra, consisting of isodiametric and transverse meshes on head and isodiametric meshes on the elytra, almost indistinct on the pronotum. Colour from reddish-brown on the head and pronotum to yellowish on the elytra and appendages; ventral part of the body more or less lighter than the respective parts of dorsum. Appendages long and slender. Head elongate, evidently longer than wide, strongly constricted at the neck; frontal grooves complete, extending onward onto the clypeus, deep along the apical three-fourth of the head where outwardly sinuate, somewhat shallow at the level of posterior supraorbital seta, and again deeper downhill along the collar constriction; eyes perfectly wanting, each one replaced by short, concave, slanting suture; praeocular traits present; genae convex; two pair of supraorbital setae on lines convergent posteriad, the posterior setae being within the frontal grooves. Clypeus sub-hexagonal (Fig. 2), surface covered besides some normal hairs, with four large basic setae, the outer setae being rather longer than the inner ones, one or two additional microsetae situated at marginal apical and lateral positions, as long as the inner basic setae; frontoclypeal sulcus more or less distinct. Labrum transverse (Fig. 2), apically slightly dilated, with six apical setigerous pores, inner and medial pairs of setae situated closer to each other than medial and outer pairs; anterior margin wavy, with a three or five distinct concavities, the middle one much deeper than the outer ones. Mandibles long but fairly stout (Figs 3–4), bidentate, with sharply hooked apices; retinacle of the left mandible bidentate, retinacle of the right mandible tridentate. Labium (Fig. 5) completely fused as no trace of labial suture clear between the mentum and submentum; mentum with pair of setae, median tooth large, long and porrect (at distal position reaching or slightly exceeding the level of epilobes), rather slanting ventrally, deeply bifid at the tip as denticles somewhat divergent onward; submentum with a transverse row of six setae, inner and outer pair of setae shorter than the medial one; ligula slightly broadened apically (Fig. 6), its anterior margin wavy, more pronouncedly in the middle, with eight setae of nearly equal length, two middlemost at ventral position and six lateral at dorsal position, paraglossae short, fragile, transparent, slightly curved and covered with microtrichia dorsolaterally, hardly surpassing the anterior margin of ligula; labial palpus slender (Fig. 5), with penultimate segment quadrisetose, gradually dilated towards the apex, about one-third as long as ultimate segment. Maxillae long and fine (Fig. 7), each with arcuate and sharply pointed lacinia, inner margin of lacinia with 5-6 stout setae at distal position and numerous smaller hairs at medial and proximal position; maxillary palpus entirely glabrous and asetose, slender, ultimate segment fusiform, as long as penultimate one; stipes with two setae at lateroventral position. Antennae filiform, very long and slender, posteriad not exceeding the middle of elytra. Pronotum cordate (Fig. 8), short in relation with the elytra, wider than long, constricted towards the base, with maximum width at its second-fifth. Anterior margin concave, longer than the posterior one, fore angles prominent, round; posterior margin concave. Sides convex, round anteriad, straight, convergent posteriorly, hardly sinuate before the hind angles; hind angles pointed, projecting outside and turned upwards; lateral margins grooved and their extremities reflexed upward throughout. Anterolateral setae present, posterolateral ones absent. Disc smooth, slightly convex, midline well impressed, not reaching both the basal and apical margins; apical transverse impression less distinct, laterally reaching the anterior margin; basal transverse impression sharply impressed, laterally merging into the basal foveae. Scutellum small, triangular. Elytra (Figs 1, 9–10) subovate, with shoulders obtuse though still marked, basal border absent. Sides narrower at the basis, slanting at shoulders, becoming wider posteriorly, with maximal width at the third-fourth, round at apex; lateral grooves moderately reflexed upward, and becoming flat at the apical sixth. Disc convex; striae superficial and more or less obliterated at the sides and apically, striae 2 and 3 forming an apical anastomosis, stria 8 fragmentary, not deepening posteriorly; scutellar stria absent; apical recurrent stria well-defined, subparallel to the suture, joining stria 5; intervals flat, each one with single longitudinal row of hairs in the middle; stria 3 with single (? anterior) discal setigerous puncture before the middle, at the level of the fourth pore of humeral umbilicate series; apical triangle of pores complete, the preapical pore as developed as the discal one, on the apical anastomosis of striae 2 and 3, more distant from the apex than from the suture, larger and rather removed from other two apical pores, the internal apical pore somewhat closer to the apex than to the suture, the external apical pore lies inward, at the recurrent stria field, not near to the recurrent stria, closer to the apex than to the suture. Umbilicate series of elytra consisting of 8 pores, not aggregate, very peculiar in disposition, six pores situated inwards, at discal, not marginal position; humeral umbilicate series consisting of four setigerous pores, the first one distant from the marginal gutter, slightly before the level of the second pore, which laying at the marginal gutter, the third pore slightly remote from the gutter, the fourth pore widely distant from both the other pores of humeral group and the gutter, the distance between third and fourth being larger than that between the first and third; two pairs of pores each of the middle and apical groups of umbilicate series not closely set together, the distance between the pores of each group being smaller than the distance between the sixth and seventh, the distance between the pores of the middle umbilicate series almost twice smaller than the distance between the pores of the apical umbilicate series; fifth pore more remote inward than sixth one, seventh pore rather remote inward, the eight pore near to the marginal gutter. Wings absent. Thorax and lateral sides of abdominal sternites smooth, middle parts of the sternites pubescent, sternites 3-5 each with pair of setigerous punctures; anal sternites with a pair of setae in the males, with two pair of setae in the females. Legs long and slender; protibia straight, densely pubescent, without longitudinal groove on the external face; tarsi thin, densely pubescent, without modified appendages underneath; protarsi with segments 1 and 2 strongly dilated and inwardly denticulate (more in the first segment, less in the second one), furnished beneath with sexual adhesive appendages in the males (Fig. 11), same segments in the females without such characterization (Fig. 12), segment 1 of the protarsi shorter than segments 2-4 together and clearly shorter than segment 5 in the males, while segments 1 of protarsi hardly shorter than segment 5 in females; both the mesotarsi and metatarsi with segment 1 almost as long as segments 2-4 together and rather longer than segment 5 in the two sexes. Aedeagus (Figs 13–15) with straight apex and poorly differentiated basal bulb in lateral aspect, thinner to the apex, moderately wide at basal and medial parts; sagittal carina moderately large, round; ventral margin straight apically, slightly concave at the middle; dorsal margin little convex with preputial bulge before the basal part; apical orifice very large, reaching the basal part of aedeagus; basal orifice small, flat, weakly differentiated; aedeagus slightly asymmetrical in dorsal aspect, with left side concave at the middle, apex widely round. Inner sac (endophallus) with an “anisotopic”, well differentiated, large copulatory piece surrounded of a field of sclerous scales, situated in the medial and distal parts of the aedeagus; copulatory piece double, its ventral branch partially visible at left lateral aspect, same longer in dorsal aspect than in the lateral one, V-shaped, with dorsal branch well visible from above, more chitinized proximally, ventral branch less evident as sunken in compact chitin. Parameres elongate and thick, each one with four apical setae, the distal three large and long, the proximal one smaller, left paramere with additional minute seta in medial position. Ovipositor consisting of large valvifer and two-segmented stylus (Fig. 16). Valvifer nearly twice larger than the stylus, with more chitinized, glabrous internal margin and less chitinized, pubescent (17-20 setae visible) external margin in sternal position. Basal stylomere of stylus larger than the apical one, with three marginal setae at central sternal position and two bigger ones distally, one on the sternal face, another on the tergal face; apical stylomere subtriangular, with two rather large ensiform setae (as long as or longer than the half length of stylomere) on tergal position, and sensorial fovea bearing two minute nematiform setae on sternal position.