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Purple Ramping Fumitory

Fumaria purpurea Pugsley

Biology

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This plant is an annual, and flowers between July and October. The fruits are produced on downward curved pedicels. As the purple ramping-fumitory resembles others in the family, it has been somewhat overlooked by botanists, and not a lot is known about the plant.
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Conservation

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Purple ramping-fumitory is listed in the UK Biodiversity Action Plans (UKBAPs), and included in English Nature's Species Recovery Programme (SRP). The priority action for this species is to establish the true status of the plant, and to protect it at its present sites. Once this is achieved, work can begin on increasing the range and populations. With any endangered plant, it is important to ensure that seeds are collected and stored as an insurance against possible extinction, and to assist in any re-introduction programme. The Millennium Seed Bank, maintained by the Royal Botanic Gardens, Kew, at Wakehurst Place, has stored seeds from many UK plants. Purple ramping-fumitory is one of many arable plants that could benefit from agricultural/environment schemes. Efforts are underway to encourage farmers and landowners to adopt these schemes in order to preserve these scarce plants.
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Description

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Purple ramping-fumitory is a climbing plant which can also be found sprawling across the ground. It bears up to 24 purple flowers, 12 mm in length, in clusters on the end of its stems. The leaves form three or five lobes and are arranged alternately up the stems. The fumitory family's English and scientific names derive from the Latin fumus terrae, meaning 'earth smoke'. This is believed to stem from an early botanist who described the appearance of fumitory 'as if the ground were all of a smoak'.
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Habitat

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Purple ramping-fumitory is a plant of hedge-banks, arable land, waste ground and occasionally sea cliffs. It favours sites subject to recent disturbance, or drought affected areas.
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Range

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This species is endemic to Britain, Ireland and the Channel Islands. It is widely distributed but the main concentrations are in western and northern Britain. Its present strongholds are Cornwall and Lancashire.
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Status

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Classified as Lower Risk: Nationally Scarce.
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Threats

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This species was not discovered until 1902 and, as it has not been fully studied, it is little understood why it is declining. However, as with so many of our arable weeds, the likely cause is changes in farming practices.
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Fumaria purpurea

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Fumaria purpurea, known as purple ramping-fumitory,[1] is an annual flowering herbaceous plant in the poppy family which is endemic to the British Isles.

Description

A sprawling or climbing plant with brittle stems which exude white sap when broken, up to about 2 m tall, typically found growing up through hedges. The whole plant is hairless and smooth, with pale green stems and leaves, and flowers that range from white through to dark purple, but generally pink. The leaves are flat but often curled and deeply divided into numerous irregular-shaped lobes, based on multiples of three (a ternate pattern). The inflorescence is a raceme with 15-24 individual flowers on short recurved stalks, each hermaphroditic flower being 10–13 mm long, with large oval sepals up to 6.5 mm in length, and 4 petals arranged into a characteristic tube shape. The fruits are roughly spherical, 2.5 mm in diameter, with a distinct ring or neck at the base when fresh.[2]

Distinctive features for separating this species from other fumitories are the purplish flowers that are strongly bent back along the stalk, the large sepals and neck on the fruits.[3][4]

Distribution and habitat

Distribution of purple ramping-fumitory, as shown in the BSBI handbook[3]

Fumaria purpurea has only been recorded in Britain, Ireland, the Isle of Man and the Channel Isles (an area sometimes referred to by naturalists as the British Isles), which makes it endemic to this area. It has a striking and unusual distribution pattern, being found in Britain mainly along a vertical line from Devon to Orkney. It grows in hedges and on disturbed ground, including gardens, arable fields and construction sites.[5][3] It was first collected in 1726 by Johann Jacob Dillenius, "ad sepes prope Shrewsbury" (in hedges near Shrewsbury, where it still occurs), according to a specimen at Oxford University (OXF)[6] which was identified much later by Pugsley.[7] In Britain and Ireland it is classed as "least concern" by the JNCC and the National Parks and Wildlife Service, respectively, while it is also listed as "vulnerable" in England and "critically endangered" in Wales.[8][9]

Taxonomy

Typical growth form of purple ramping-fumitory in the gardens of Attingham Park

Purple ramping-fumitory was named in 1902 by H.W. Pugsley, who also described two varieties: var. longisepala, with sepals up to 6.5 mm long, and var. brevisepala, with shorter sepals only 5 mm long, which could be confused with F. muralis var. boraei.[10] The two varieties are usually ignored now. There is some debate about its origin and its relationship to other species in the genus. Some authorities consider it to be closely related to F. muralis because of chemical and cytological similarities, while others place it alongside F. capreolata, owing to their morphological similarity.[3]

Fumaria purpurea has the chromosome number 2n = 80.

References

  1. ^ BSBI List 2007 (xls). Botanical Society of Britain and Ireland. Archived from the original (xls) on 2015-06-26. Retrieved 2014-10-17.
  2. ^ Stace, C.A. (2019). New Flora of the British Isles. Suffolk. ISBN 978-1-5272-2630-2.
  3. ^ a b c d Murphy, R.J. (2009). Fumitories of Britain and Ireland. London: Botanical Society of the British Isles. ISBN 978-0-901158-40-6.
  4. ^ Rich, T.C.G. "Plant Crib: Fumaria" (PDF).
  5. ^ Lockton, A.J. "BSBI species accounts: Fumaria purpurea". Retrieved 30 August 2010.
  6. ^ New York Botanic Gardens. "Index Herbariorum".
  7. ^ Lockton, Alex; Whild, Sarah (2015). Flora and vegetation of Shropshire. Montford Bridge: Shropshire Botanical Society.
  8. ^ Cheffings, C.; Farrell, L. (2005). The Vascular Plant Red Data List for Great Britain. Peterborough: Joint Nature Conservation Committee.
  9. ^ Wyse Jackson, Michael (2016). Ireland Red List No. 10: Vascular Plants (PDF). Dublin: National Parks and Wildlife Service.
  10. ^ Pugsley, H.W. (1902). "The British capreolate fumitories". Journal of Botany. 40: 129–136, 173–181.
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Fumaria purpurea: Brief Summary

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Fumaria purpurea, known as purple ramping-fumitory, is an annual flowering herbaceous plant in the poppy family which is endemic to the British Isles.

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