Cotoneaster verruculosus Diels

Specimens examined at A: Sikkim: Patria, Chiyabanjan-Phalut, 3100–3800 m, 2 Jun 1960, H. Hara, H. Kanai, G.Murata, M. Togashi & T. Tuyama 2012; Bhutan & Sikkim Border, 2 Sep 1913, R. E. Cooper 786; E. Himalaya: Phallut, 3048 m, 15 Apr 1918, G. H. Cave n.s.

Shrubs deciduous or semi-evergreen, prostrate to erect, 0.6–1.2(–2) m tall. Branchlets initially dark reddish brown to grayish brown, dark gray when old, terete, initially densely yellow strigose, glabrate, conspicuously verruculose when old. Petiole 2–5 mm, pubescent; stipules persistent, lanceolate, 2–6 mm, pubescent; leaf blade suborbicular or broadly ovate to broadly obovate, 0.7–1.4 cm × (4–)6–12 mm, both surfaces appressed pilose, glabrescent, abaxially subglabrous, base broadly cuneate or rounded, apex emarginate or partly mucronulate. Inflorescences usually 1-flowered. Flowers 7–8 mm in diam., nearly sessile. Hypanthium glabrous abaxially. Sepals ovate-triangular, 1.5–2.5 mm and ca. as broad, apex ± obtuse. Petals erect, pink, suborbicular, 4–4.5 mm in diam. Stamens 20, shorter than petals. Ovary pubescent apically; styles 2, free, not exceeding stamens. Fruit red, globose, 7–8 mm in diam.; pyrenes often 2. Fl. May–Jun, fr. Sep–Oct.

Shrubs deciduous or semievergreen, erect, 0.6–2 m tall. Branchlets initially dark reddish brown to grayish brown, dark gray when old, terete, initially densely yellow strigose, glabrate, conspicuously verruculose when old. Petiole 2–5 mm, pubescent; stipules persistent, lanceolate, 2–6 mm, pubescent; leaf blade suborbicular or broadly ovate to broadly obovate, 0.7–1.4 cm × 6–12 mm, both surfaces appressed pilose, glabrescent, abaxially subglabrous, base broadly cuneate or rounded, apex emarginate or partly mucronulate. Inflorescences usually 1-flowered. Flowers 7–8 mm in diam., nearly sessile. Hypanthium glabrous abaxially. Sepals ovate-triangular, 1.5–2.5 mm and ca. as broad, apex ± obtuse. Petals erect, pink, suborbicular, 4–4.5 mm in diam. Stamens 20, shorter than petals. Ovary pubescent apically; styles 2, free, not exceeding stamens. Fruit red, globose, 7–8 mm in diam., pyrenes often 2. Fl. May–Jun, fr. Sep–Oct.

Himalaya (Nepal to Sikkim), Burma, W. China.

Himalaya (Nepal to Sikkim); Bhutan, W. China (Sichuan, Xizang/Tibet, Yunnan), N. India, Myanmar, Nepal, Sikkim.

Sichuan, Xizang, Yunnan [Bhutan, N India, Myanmar, Nepal, Sikkim].

3000-3400 m

Mixed forests on dry slopes, grasslands, fields, roadsides; [2800–]3000–3400[–3600] m.

Mixed forests on dry slopes, grasslands, fields, roadsides; 2800--3600 m.

Cotoneaster distichus Lange var. verruculosus (Diels) T. T. Yü, Bull. Brit. Mus. (Nat. Hist.), Bot. 1: 128. 1954.

Cotoneaster sandakphuensis G. Klotz, Bull. Bot. Surv. India 5: 213. 1964; Type: "Sikkim-Himalaya, prope Sandakphu, infruticetis via, Rimbick", 9 Oct 1941, K. P. Biswas 5721, fr.

Cotoneaster distichus Lange var. verruculosus (Diels) T. T. Yü; C. improvisus G. Klotz.