dcsimg

Lifespan, longevity, and ageing

المقدمة من AnAge articles
Maximum longevity: 12 years (wild) Observations: Unverified estimates suggest these animals may live up to 23 years (http://www.fishbase.org/).
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Joao Pedro de Magalhaes
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de Magalhaes, J. P.
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AnAge articles

Trophic Strategy ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata is carnivorous. Juveniles feed predominantly on bottom dwelling aquatic crustaceans such as amphipods, mysids and crangonid shrimps. Larger adults feed on larger crustaceans such as swimming crabs and fish, such as sandeels, small gadoids and dragonets.

Animal Foods: fish; aquatic crustaceans

Primary Diet: carnivore (Piscivore , Eats non-insect arthropods)

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
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Jason Dooney, The College of New Jersey
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Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Life Cycle ( الإنجليزية )

المقدمة من Animal Diversity Web

Young Raja clavata hatch at 11 to 13 cm in length and 8 to 9 g in weight with a male:female sex ratio of 1:1. Juveniles remain in shallow water. As they increase in size, juveniles grow large button like structures called bucklers. When R. clavata reaches sexual maturity, usually around 8.8 years, it begins seasonally migrating. During summer, adults move in shore during the summer and migrate into deeper waters during the winter.

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
موقع الشريك
Animal Diversity Web

Conservation Status ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata is classified as near threatened on to the ICUN's Red List of Threatened Species. Declines in R. clavata landings are thought to be due to over exploitation and poor species identification. It is caught primarily as by-catch in both otter and beam trawl fisheries, and is targeted with gillnets, long-lines, and recreational anglers. Landings of rays and skates are often lumped together under one generic category (i.e., Raja spp.), but market samplings suggest that R. clavata is the most dominant species landed in the North Sea and Skagerrak. Over exploitation in this species is especially relevant as it has slow growth, long life span, and long period before individuals become reproductively mature make. Research suggests that R. clavata may be extinct from the North Sea due to over fishing. In an effort to help R. clavata recover (as well as other batids), total allowable catch (TAC) for all skates and rays was reduced by 47% from 1999 to 2005. Some fisheries in the United Kingdom have also implemented a minimum landing size. Unfortunately, localized management efforts are not expected to have an impact on regional population conservation.

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: near threatened

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
موقع الشريك
Animal Diversity Web

Behavior ( الإنجليزية )

المقدمة من Animal Diversity Web

Like most other elasmobranchs, Raja clavata uses a diverse array of senses that allow it to perceive orientation as well as locate potential predators and prey. In order locate predators and prey, it depends on its acute sense of smell and a lateral line system that allows it to detect changes in pressure in the local environment. The lateral line system is made up of individual mechano-receptors called neuromasts which run the length of the body. These neuromasts are hair filled structures surrounded with a jelly like substance. These are connected to lateral line canals which interact with the environment. Pressure changes in the water displace the hair and fires impulses to the organism’s brain alerting it to the direction and strength of the disturbance. Raja clavata also depends on electrosensory perception organs known as Ampullae of Lorenzini. The ampullae, which consists of a series of gelatin filled pits that create an electrically sensitive network along the ventral surface of the animal, help sense minor electrical pulses created by the muscle contractions of organisms in the local environment. The ampullary organs interact with local electric fields through an opening in the skin on the snout of the animal that leads to a jelly filled canal. The electrical fields produced by animals dissipate rather quickly so R. clavata can only perceive the weak current at short range. Within that short range the thornback ray can accurately determine the location of the prey, even if buried in the sand. Some studies have indicated that R. clavata as well as other sharks and rays can use their Ampullae of Lorenzini to detect the strength and direction of the Earth’s magnetic field as a sort of rudimentary positioning system. Studies have shown that the thornback ray does not rely sight for locating prey.

Raja clavata acquires information from its three otilith organs (the utriculus, the sacculus, and the lagena) to maintain spatial equilibrium in its marine habitat. These organs are located in the inner ear vestibular system where the semicircular canals converge next to the cochlea. The three organs respond to tilts in both the longitudinal and transverse axes. The utriculus and sacculus produce similar responses and thus create mutually reinforcing signals. The macula lagena is made up of gravity receptors which provide a marker for R. clavata upright position. The macula utriculus produces an increase in discharge to signal side-up and nose-up displacements while the macula sacculus signals side-up and nose-down displacements. All three maculae act as an "out-of-position receptor" when the individual is subjected to a constant speed spatial deviation.

Thornback rays have a very acute sense of smell that they use for locating prey. They have two openings called nares that are located ventrally on the organism’s snout. Water enters the nares and passes through a structure known as the olfactory sac. The olfactory sac is lined with a series of folded tissue called the olfactory lamellae which provide increased surface area for molecule-receptor interactions. Molecules dissolved in the water bind to neuroreceptors in the olfactory lamellae to provide chemosensory information to the brain.

Perception Channels: visual ; acoustic ; vibrations ; chemical ; electric ; magnetic

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الاقتباس الببليوغرافي
Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
موقع الشريك
Animal Diversity Web

Benefits ( الإنجليزية )

المقدمة من Animal Diversity Web

There are no known adverse effects of Raja clavata on humans.

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Benefits ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata is more highly valued for its meat than other skates and rays. Thus, R. clavata plays an important economic role in fisheries, particularly those off the coast of Portugal.

Positive Impacts: food

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
موقع الشريك
Animal Diversity Web

Associations ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata feeds on a variety of benthic animals. Juveniles feed on small crustaceans such as amphipods, mysids and crangonid shrimps, and adults feed on fish and larger crustaceans such as swimming crabs and fish such as sandeels, small gadoids and dragonets. This species is also known to feed on conspecifics as well. The eggs of R. clavata act as prey for some fish and mollusks such as necklace shells. Juveniles are preyed upon by common skate and adult R. clavata. Due to its high placement in the costal sea floor food web, R. clavata has a significant effect on trophic levels of the benthos. Parasites of this species have not been documented.

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
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Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Distribution ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata is wide spread in coastal waters from Iceland to Norway. Its geographic ranges extends into the North Sea, the Mediterranean Sea, the western Black Sea, Madeira Island, the Atlantic coasts of Africa, as well the waters off of South Africa and the south-western Indian Ocean. Adult R. clavata remains inshore during summer and move off shore into deeper waters during winter.

Biogeographic Regions: palearctic (Native ); ethiopian (Native ); mediterranean sea (Native )

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
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زيارة المصدر
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Habitat ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata lives at the bottom of the continental shelf and upper slope of the coastal sea floor. It inhabits a wide range of sea floor habitats including mud, sand, shingle, gravel and rocky areas. Raja clavata has been found in coastal waters as deep as 300 m but is most abundant in waters between 10 and 60 meters. Juveniles are often found further inshore in more shallow waters than their adult counterparts.

Range depth: 10 to 300 m.

Average depth: 60 m.

Habitat Regions: saltwater or marine

Aquatic Biomes: coastal

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Life Expectancy ( الإنجليزية )

المقدمة من Animal Diversity Web

In the wild, Raja clavata lives between 12 and 15 years. There is no information available regarding the average lifespan of captive individuals.

Typical lifespan
Status: wild:
12 to 15 years.

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The Regents of the University of Michigan and its licensors
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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Morphology ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata shares its general body shape with all rays in that it is a flattened, disc shape with broad pectoral fins connected to the head and body with triangular pelvic fins at the base of the body. Raja clavata has a long, narrow tail that is roughly equivalent to the length of the body. As the common name denotes (thornback ray), the upper portion of the body and tail are covered with thorn-like projections. When R. clavata reaches sexual maturity the bases of the thorns thicken to resemble small button like projections called buckler. The snout and small portions of the body are covered in thorns in sub adults and the underside may be thorny in large females. Adults typically have between 21 and 25 large thorns running from the nape to its first pectoral fin. It is sexually dimorphic, as females have a maximum length of 118 cm and males have a maximum length of 98 cm. The dorsal surface of R. clavata exhibits a wide range of coloration from light brown to grey and is sprinkled with light and dark spots of varying size. Its underside is solid white except for the snout which is grey. The largest specimen ever recorded weighed 18 kg.

Range mass: 18 (high) kg.

Range length: 98 to 120 cm.

Other Physical Features: ectothermic ; heterothermic ; bilateral symmetry

Sexual Dimorphism: female larger

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Associations ( الإنجليزية )

المقدمة من Animal Diversity Web

Raja clavata eggs are preyed upon by some fish and mollusks, such as necklace shells. Juveniles are commonly preyed upon by larger fish, specifically common skates and adult R. clavata. The coloration of R. clavata likely helps reduce risk of predation.

Known Predators:

  • necklace shells, Polinices
  • common skate, Dipturus batis
  • thornback rays, Raja clavata

Anti-predator Adaptations: cryptic

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
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Animal Diversity Web

Reproduction ( الإنجليزية )

المقدمة من Animal Diversity Web

Little is known of the mating behavior of many skates, including Raja clavata. Some speculate that this is because the animals mate in the cover of night. Studies of sting rays may provide insight into the mating behaviors of elasmobranchs. During mating, females swim in shore and spend the morning either buried in the sand or lay atop each other in large groups. Those females buried are attempting to avoid males that already mated or are not yet ready for reproduction whilst those in the aggregate groups are prepared for mating. Meanwhile, males swim up and down the beach searching for potential mates. In the afternoon, they switch roles and the males bury themselves in the sand while the females become mobile to search for food in the grassy beds. This cycle can continue for multiple weeks.

Thornback rays are oviparous with females migrating further in shore to lay their eggs. Egg-laying season for Raja clavata occurs between March and September. Mature females deposit one egg at a time in sandy or muddy substrate close to shore. Eggs range in length from 5 to 9 cm long and 3.8 to 6.4 cm wide. Mature females can lay between 140 and 160 eggs in a single year. Eggs are rectangular and oblong in shape and are marked by rigid horns on each corner. During development, embryos feed solely on their yolk. Embryonic development typically takes between 4 and 6 months and is largely dependent on ambient water temperature.

Breeding season: Thornback rays breed from March to September.

Range number of offspring: 140 to 160.

Range gestation period: 4 to 6 months.

Average age at sexual or reproductive maturity (female): 8.8 years.

Average age at sexual or reproductive maturity (male): 7.1 years.

Key Reproductive Features: iteroparous ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; fertilization (Internal ); oviparous

There is no information available regarding parental care in Raja clavata; however, studies on closely related elasmobranches indicate a high level of parental investment.

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Dooney, J. 2011. "Raja clavata" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Raja_clavata.html
مؤلف
Jason Dooney, The College of New Jersey
محرر
Matthew Wund, The College of New Jersey
محرر
John Berini, Special Projects
النص الأصلي
زيارة المصدر
موقع الشريك
Animal Diversity Web

Biology ( الإنجليزية )

المقدمة من Arkive
All skates are oviparous (8) (egg-laying), with large female thornbacks laying up to 140 to 160 egg capsules per year (6). The species moves into shallower, inshore waters in spring when ready to spawn, laying eggs between March and September, which take a further four to six months to hatch (6). Young feed on small crustaceans and other bottom-dwelling creatures, predating more on fish and larger crustaceans such as crabs as they develop (5) (6). Maturity is reached at around eight years of age (5), and the maximum lifespan is at least 15 years (6).
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Conservation ( الإنجليزية )

المقدمة من Arkive
Fishing of the thornback skate is controlled through the authorities limiting the Total Allowable Catch (TAC), although the quota is set for all rajids combined, covering EU waters in the North Sea and Norwegian Sea. This measure helps ensure continuity of economic activity in the fisheries concerned, while giving depleted stocks a reasonable chance of recovery. The TAC for skates and rays has been reduced by approximately 47 % between 1999 and 2005, providing greater protection to the species by reducing the quota allowed to be caught. However, records show that the actual numbers caught each year have in fact been below what is permissible through the TAC (6). Thus, these skate are not currently considered to be particularly over-fished and are not seen as a high conservation priority.
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Description ( الإنجليزية )

المقدمة من Arkive
The thornback skate is probably one of the commonest skates encountered by divers (3), being among the most abundant rajids in the north-eastern Atlantic and Mediterranean (1). As with all skates, the body is flattened and disc-shaped, with the pectoral fins broadly expanded and joined to the head and body (4). The tail is distinctly demarcated from the disc-like body, relatively narrow, and about as long as body length (3) (4). The upper surface of the disc and tail are covered with numerous thorns, which become thickened with button-like bases (known as bucklers) once the skate is sexually mature, hence the species' common name (3) (5). Only the snout and margins of the disc are prickly in young, and the underside is only prickly in large, mature females, which also possess more developed 'bucklers' on their back and tail (2) (3). The colour varies from light brown to grey on the upper surface, variegated with dark and light spots and blotches that camouflage the skate in the dappled light of the sea bed, while the underside is creamy-white (2) (3).
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Habitat ( الإنجليزية )

المقدمة من Arkive
A bottom-dwelling species occupying continental shelf and upper slope waters to depths of 300 m, but mainly from 10 to 60 m (6). However, off South Africa this species occurs to 1000 m (4). Usually found on sediment type sea beds such as mud, sand or gravel, occasionally adjacent to rocky reefs (3) (7).
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Range ( الإنجليزية )

المقدمة من Arkive
Found in the eastern Atlantic from Iceland, Norway, North Sea and the western Baltic southward to Morocco and Namibia, including the Mediterranean and the Black Sea (2). Also reported from South Africa eastward into the south-western Indian Ocean (4).
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Status ( الإنجليزية )

المقدمة من Arkive
Classified as Lower Risk/near threatened (LR/nt) on the IUCN Red List 2004 (1).
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Threats ( الإنجليزية )

المقدمة من Arkive
The thornback skate is an important component of mixed trawl fisheries, has been targeted by recreational anglers (1), and is also caught incidentally as bycatch in beam trawl fisheries (9). The thorniness of this species adds to their chance of being entangled in nets (9). There is some evidence of a decline in catch rates in north-west European waters, which could imply a decline in population numbers, but more data is required to confirm this (1). The impact of the fishing industry on rays and skates is the subject of increasing concern due to their slow growth rate, late maturity and low fecundity, making them particularly vulnerable to population collapse due to fishing activity (10).
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Brief Summary ( البلجيكية الهولندية )

المقدمة من Ecomare
Stekelroggen danken hun naam aan de vele stekels op hun rug en staart. Toch zijn ze ongevaarlijk, de stekels zijn niet giftig. Om aan eten te komen, verstoppen ze zich onder een laagje zand van de zeebodem. Samen met de stippen op hun rug zorgt dat er voor dat ze goed gecamoufleerd zijn. Als er wormen, kreeftachtigen, zeeslakken of kleine bodemvissen voorbij zwemmen, vangen ze die vanuit een hinderlaag. Stekelroggen hebben rijen met sterke, botte tanden. Wanneer er een tand versleten is komt de volgende in de rij naar voren. Volwassen stekelroggen hebben wel 36 tot 44 rijen tanden.
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Brief Summary ( الإنجليزية )

المقدمة من Ecomare
The thornback ray owes its name to the numerous thorns on its back and tail. Nevertheless, the animal is not dangerous because the thorns are not poisonous. In order to catch its food, it hides under a layer of sand on the sea floor. Together with the spots on its back, it is well camouflaged. Any worms, crustaceans, sea snails or small benthic fish that happen to swim past are caught from its ambush. Thornbacks have rows of strong blunt teeth. When a tooth is worn down, the next one in line appears. Adult thornback rays have 36 to 44 rows of teeth.
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Trophic Strategy ( الإنجليزية )

المقدمة من Fishbase
Nocturnal predator. Feeds on all kinds of bottom animals (Ref. 3167). Polychaetes and crustaceans are important prey items of smaller individuals, bony fishes become more important prey items of larger individuals (Ref. 28070).
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Life Cycle ( الإنجليزية )

المقدمة من Fishbase
Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).
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Diagnostic Description ( الإنجليزية )

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Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).
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Morphology ( الإنجليزية )

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Dorsal spines (total): 0; Analspines: 0; Analsoft rays: 0
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Biology ( الإنجليزية )

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Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (
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Importance ( الإنجليزية )

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fisheries: commercial; gamefish: yes
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Rae vouklet ( البريتانية )

المقدمة من wikipedia BR

Ar rae vouklet (liester: raeed bouklet), pe rae lagadek (liester: raeed lagadek), Raja clavata eus hec'h anv skiantel, a zo ur pesk plat, kar d'ar rinkin, hag a vev er meurvor Atlantel hag er mor Kreizdouar[1].

Doareoù pennañ

Etre 2 ha 4 kg eo he fouez ha war-dro 90 cm he hirder. Gris-gell eo he c'hein ha sklaer he c'hof. Goloet eo he c'horf gant drein e stumm krogoù a zo kaoz eus hec'h anv. Kuzhat a ra e strad ar mor, evel an darn vrasañ eus ar raeed.

Dave


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  1. Aquamaps.org
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Rae vouklet: Brief Summary ( البريتانية )

المقدمة من wikipedia BR

Ar rae vouklet (liester: raeed bouklet), pe rae lagadek (liester: raeed lagadek), Raja clavata eus hec'h anv skiantel, a zo ur pesk plat, kar d'ar rinkin, hag a vev er meurvor Atlantel hag er mor Kreizdouar.

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Clavellada ( الكتالونية )

المقدمة من wikipedia CA
 src=
Clavellada

La clavellada, l'escrita, la bastina (per extensió errònia[4]), la bastina clavejada, el clavell, la rajada, la rajada punxosa o la rajada vera (Raja clavata) és una espècie de peix de la família dels raids i de l'ordre dels raïformes.

Morfologia

Reproducció

És ovípar,[5] els ous tenen com a banyes a la closca[8] i les femelles en ponen entre 52 i 170 a l'any.[5]

Alimentació

Menja tota mena d'animals bentònics, preferiblement crustacis.[9]

Depredadors

És depredat per Chelidonichthys gurnardus.[10][11]

Hàbitat

És un peix marí, de clima subtropical (70°N-29°S, 25°W-42°E) i demersal que viu entre 20–577 m de fondària.[5][12]

Distribució geogràfica

Es troba a l'Oceà Atlàntic oriental (des d'Islàndia, Noruega, la Mar del Nord i l'oest del Mar Bàltic fins al Marroc i Namíbia), la Mediterrània i la Mar Negra.[5][13][14][15][16][17][18][19][20][21][22][23][24][25][26][27][28][29][30][31][32][33][34][35][36][37][38][39][40][41][42][43][44][45][46][47][48][49][50][51][52][53][54][55][56][57][58][59][60][61][62][63][64][65][66]

Ús comercial

Es comercialitza fresc i congelat.[67]

Observacions

És inofensiu per als humans.[5]

Referències

  1. Linnaeus C. 1758. Systema Naturae, Ed. X. (Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.) Holmiae. Systema Nat. ed. 10 v. 1. i-ii + 1-824.
  2. BioLib (anglès)
  3. Linnaeus, C., 1758. Systema Naturae, Ed. X. (Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.) Holmiae. Systema Nat. ed. 10 v. 1: i-ii + 1-824.
  4. https://dialnet.unirioja.es/servlet/articulo?codigo=1191807 Comentaris sobre algunes causes d'errors o d'inexactituds en la nomenclatura catalana dels peixos marins, Miquel Duran i Ordiñana
  5. 5,0 5,1 5,2 5,3 5,4 5,5 FishBase (anglès)
  6. Dorel, D., 1986. Poissons de l'Atlantique nord-est relations taille-poids. Institut Francais de Recherche pour l'Exploitation de la Mer. Nantes, França. 165 p.
  7. Muus, B.J. i J.G. Nielsen, 1999. Sea fish. Scandinavian Fishing Year Book, Hedehusene, Dinamarca. 340 p.
  8. Breder, C.M. i D.E. Rosen, 1966. Modes of reproduction in fishes. T.F.H. Publications, Neptune City, Nova Jersey, Estats Units. 941 p.
  9. Stehmann, M. i D.L. Bürkel, 1984. Rajidae. p. 163-196. A: P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen i E. Tortonese (eds.) Fishes of the north-eastern Atlantic and Mediterranean. UNESCO, París. vol. 1.
  10. De Gee, A. i A.H. Kikkert, 1993. Analysis of the grey gurnard (Eutrigla gurnardus) samples collected during the 1991 International Stomach Sampling Project. ICES C.M. 1993/G:14. 25 p.
  11. FishBase (anglès)
  12. Mytilineou, C., C.-Y. Politou, C. Papaconstantinou, S. Kavadas, G. D'Onghia i L. Sion, 2005. Deep-water fish fauna in the Eastern Ionian Sea. Belg. J. Zool., 135(2): 229-233.
  13. Ajayi, T.O., 1982. Food and feeding habits of Raja species (Batoidei) in Carmarthen Bay, Bristol Channel. J. Mar. Biol. Assoc. U.K. 62:215-223.
  14. Alegre, M., J. Lleonart i J. Veny, 1992. Espècies pesqueres d'interès comercial. Nomenclatura oficial catalana. Departament de Cultura, Generalitat Catalunya, Barcelona, Països Catalans.
  15. Bauchot, M.-L., 1987. Raies et autres batoides. p. 845-886. A: W. Fischer, M.L. Bauchot i M. Schneider (eds.). Fiches FAO d'identificationpour les besoins de la pêche. (rev. 1). Mèditerranée et mer Noire. Zone de pêche 37. Vol. II. Commission des Communautés Européennes i FAO, Roma.
  16. Bianchi, G., 1986. Fichas FAO de identifacao de espécies para propósitos comerciais. Guia de campo para as espécies comerciais marinhas e de águas salobras de Angola. Preparado com o apoio da NORAD e da FAO (FIRM) Programa Regular, FAO, Roma. 184 p.
  17. Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer i H.J. Boyer, 1993. FAO species identification field guide for fishery purposes. The living marine resources of Namibia. FAO, Roma, Itàlia. 250 p.
  18. Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer i H.J. Boyer, 1999. Field guide to the living marine resources of Namibia. FAO species identification guide for fishery purposes. Roma, FAO. 265 p.
  19. Bilecenoglu, M., E. Taskavak S. Mater i M. Kaya, 2002. Checklist of the marine fishes of Turkey. Zootaxa (113):1-194.
  20. Borges, T.C., S. Olim i K. Erzini, 2003. Weight-length relationship for fish species discarded in commercial fisheries of the Algarve (southern Portugal). J. Appl. Ichthyol. 19(6):394-396.
  21. Brander, K. i D. Palmer, 1985. Growth rate of Raja clavata in the north east Irish Sea. J. Cons. CIEM 42:125-128.
  22. Brito, A., 1991. Catálogo de los peces de las Islas Canarias. Francisco Lemus, La Laguna, Illes Canàries, Espanya. 230 p.
  23. Compagno, L.J.V., D.A. Ebert i M.J. Smale, 1989. Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., Londres. 158 p.
  24. Costa, F., 1991. Atlante dei pesci dei mari italiani. Gruppo Ugo Mursia Editore S.p.A. Milà, Itàlia. 438 p.
  25. Coull, K.A., A.S. Jermyn, A.W. Newton, G.I. Henderson i W.B. Hall, 1989. Length/weight relationships for 88 species of fish encountered in the North Atlantic. Scottish Fish. Res. Rep. (43):80 p.
  26. Demirhan, S.A., S. Semih Engin, K. Seyhan i E. Akamca, 2005. Some biological aspects of thornback ray (Raja clavata L., 1758) in the southeastern Black Sea. Turk. J. Fish. Aquat. Sci. 5: 75-83 (2005).
  27. Ebeling, E., 1988. A brief survey of the feeding preferences of Raja clavata in Red Wharf Bay in the Irish Sea. p. 1-5 plus tables. ICES, C.M. 1988/G:58, Demersal Fish Committee.
  28. Ellis, J.R., A Cruz-Martinez, B.D. Rackham i S.I. Rodgers. 2004. The distribution of Chondrichthyan fishes around the British Isles and implications for conservation J. Northw. Atl. Fish. Sci., Vol. 35: 195-213.
  29. Filiz, H. i G. Bilge, 2004. Length-weight relationships of 24 fish species from the North Aegean Sea, Turkey. J. Appl. Ichthyol. 20:431-432.
  30. Fischer, W., M.-L. Bauchot i M. Schneider (eds.). 1987. Fiches FAO d'identification des espèces pour les besoins de la pêche. (Révision 1). Méditerranée et mer Noire. Zone de Pêche 37. FAO, Roma. 1529 p.
  31. Fricke, R., 1999. Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez): an annotated checklist, with descriptions of new species. Koeltz Scientific Books, Koenigstein, Theses Zoologicae, Vol. 31: 759 p.
  32. Fricke, R., M. Bilecenoglu i H.M. Sari, 2007. Annotated checklist of fish and lamprey species (Gnathostoma and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beitr. Naturk. Sea A (706):1-172.
  33. Gallagher, M.J., C. P. Nolan i F. Jeal, 2005. Age, growth and maturity of the commercial ray species from the Irish Sea. J. Northw. Atl. Fish. Sci., 35: 47-66.
  34. George, M., 2004. Haie, Rochen und Chimären in der Nord- und Ostsee. Elasmoskop 8(1):10-16.
  35. Gibson, R.N. i I.A. Ezzi, 1987. Feeding relationships of a demersal fish assemblage on the west coast of Scotland. J. Fish Biol. 31:55-69.
  36. Holden, M.J. i R.N. Tucker, 1974. The food of Raja clavata Linnaeus 1758, Raja montagui Fowler, 1910, Raja naevus Müller and Henle, 1841 and Raja brachyura Lafont, 1873 in British waters. J. Cons. Int. Explor. Mer. 35(2):189-193.
  37. Jonsson, G., 1992. Islenskir fiskar. Fiolvi, Reykjavik, 568 pp.
  38. Krstulovic Sifner, S., N. Vrgoc, V.: Isajlovic, I. Dadic, M. Peharda i C. Piccinetti, 2009. Long-term changes in distribution and demographic composition of thornback ray, Raja clavata, in the northern and central Adriatic Sea. J. Appl. Ichthyol. 25 (Suppl. 1) (2009):40-46.
  39. Labropoulou, M. i C. Papaconstantinou, 2000. Community structure of deep-sea demersal fish in the North Aegean Sea (northeastern Mediterranean). Hydrobiologia 440:281-296.
  40. Louisy, P., 2001. Guide d'identification des poissons marins. Europe et Méditerranée. París: Eds. Eugène Ulmer.
  41. Luther, W. i K. Fiedler, 2002. Guida della fauna marina costiera del Mediterraneo. Atlante illustrato a colori. Franco Muzzio & C. (editore), Roma. 244p.
  42. Lythgoe, J. i G. Lythgoe, 1976. Vissen van de Europese kutswateren en de Middlellandse Zee. 335 p.
  43. Machado, P.B., L.S. Gordo i I. Figueiredo. 2004. Skate and ray species composition in mainland Portugal from the commercial landings. Aquat. Living Resour. 17:231-234.
  44. Mendes, B., P. Fonseca i A. Campos, 2004. Weight-length relationships for 46 fish species of the Portugese west coast. J. Appl. Ichthyol. 20:355-361.
  45. Merella, P., A. Quetglas, F. Alemany i A. Carbonell, 1997. Length-weight relationship of fishes and cephalopods from the Balearic Islands (western Mediterranean). Naga ICLARM Q. 20(3/4):66-68.
  46. Morato, T., E. Solà, M.P. Grós i G. Menezes, 2003. Diets of thornback ray (Raja clavata) and tope shark (Galeorhinus galeus) in the bottom longline fishery of the Azores, Northeastern Atlantic. Fish Bull. 101:590-602.
  47. Nijssen, H. i S.J. de Groot, 1974. Catalogue of fish species of the Netherlands. Beaufortia 21(285):173-207.
  48. Notarbartolo-di-Sciara, G. i I. Bianchi, 1998. Guida degli squali e delle razze del Mediterraneo. 388 p.
  49. Nunes, A., 1953. Peixes da Madeira. Edição da Junta Geral do distrito Autón.
  50. Plejic, T., 2007. Fische der Adria. Das romanische Element in der kroatischen Ichthyofauna. Mit Glossar der Adriafische Kroatisch-Englisch-Deutsch-Italienisch. Diploma Thesis. University of Leipzig Institute of Applied Linguistics and Translational Studies.
  51. Quignard, J.-P. i J.A. Tomasini, 2000. Mediterranean fish biodiversity. Biol. Mar. Mediterr. 7(3):1-66.
  52. Salekhova, L.P., N.S. Kostenko, T.A. Bogachick i O.N. Minibaeva, 1988. Composition of ichthyofauna in the region of the Karadag State Reserve (Black Sea). J. Ichthyol. 28(2):16-23.
  53. Sanches, J.G., 1991. Catálogo dos principais peixes marinhos da República de Guiné-Bissau. Publicações avulsas do I.N.I.P. Núm. 16. 429 p.
  54. Santos, R.S., F.M. Porteiro i J.P. Barreiros, 1997. Marine fishes of the Azores: annotated checklist and bibliography. Bulletin of the University of Azores. Supplement 1. 244 p.
  55. Schneider, W., 1990. FAO species identification sheets for fishery purposes. Field guide to the commercial marine resources of the Gulf of Guinea. Prepared and published with the support of the FAO Regional Office for Africa. FAO, Roma. 268 p.
  56. Serena, F., R. Baino i P. Righini, 1988. Geographical and depth distribution of rays in northern Tyrrhenian sea. Rapp. Comm. Int. Mer. Méditerr. 31(2):277.
  57. Soljan, T., 1975. I pesci dell'Adriatico Arnoldo Mondadori Editore, Verona, Itàlia.
  58. Stehmann, M., 1979. Rajidae. p. 58-69. A: J.C. Hureau i Th. Monod (eds.) Checklist of the fishes of the north-eastern Atlantic and of the Mediterranean (CLOFNAM), UNESCO, París, Vol. 1.
  59. Stehmann, M., 1990. Rajidae. p. 29-50. A: J.C. Quero, J.C. Hureau, C. Karrer, A. Post i L. Saldanha (eds.) Check-list of the fishes of the eastern tropical Atlantic. Junta Nacional de Investigaçao Cientifica e Tecnológica, Lisboa, Portugal. Vol. 1.
  60. Stehmann, M., 1995. A record of Raja clavata, the eastern atlantic Thornback Skate, from the southern Madagascar ridge at Walters Shoal (Elasmobranchii, Rajidae). J. Ichthyol. 35(5):63-73.
  61. Sánchez, F., F. de la Gándara i R. Gancedo, 1995. Atlas de los peces demersales de Galicia y el Cantábrico. Otoño 1991-1993. Publicaciones Especiales, Instituto Español de Oceanografía (Madrid), (20). 100 p.
  62. Went, A.E.J., 1957. List of Irish fishes. Department of Lands, Fisheries Division, Dublín. 31 p.
  63. Wheeler, A., 1992. A list of the common and scientific names of fishes of the British Isles. J. Fish Biol. 41(1):1-37.
  64. Whittamore, J.M. i I.D. McCarthy, 2005. The population biology of the thornback ray, Raja clavata in Caernarfon Bay, North Wales. J. Mar. Biol. Ass. U.K. 85:1089-1094.
  65. Winkler, H.M., K. Skora, R. Repecka, M. Ploks, A. Neelov, L. Urho, A. Gushin i H. Jespersen, 2000. Checklist and status of fish species in the Baltic Sea. ICES CM 2000/Mini:11, 15 p.
  66. Wirtz, P., R. Fricke i M.J. Biscoito, 2008. The coastal fishes of Madeira Island-new records and an annotated check-list. Zootaxa 1715: 1-26.
  67. Frimodt, C., 1995. Multilingual illustrated guide to the world's commercial coldwater fish. Fishing News Books, Osney Mead, Oxford, Anglaterra. 215 p.


Bibliografia

  • Anònim, 1997. Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Bèlgica.
  • Anònim, 2000. Base de dades de la col·lecció de peixos del J.L.B. Smith Institute of Ichthyology, Grahamstown, Sud-àfrica. J.L.B. Smith Institute of Ichthyology, Grahamstown, Sud-àfrica.
  • Anònim, 2001. Base de dades de la col·lecció de peixos del National Museum of Natural History (Smithsonian Institution). Smithsonian Institution - Division of Fishes.
  • Anònim, 2002. Base de dades de la col·lecció de peixos del American Museum of Natural History. American Museum of Natural History, Central Park West, NY 10024-5192, Estats Units.
  • Capapé, C. i M. Desoutter, 1981. Nouvelle description de Raja (Raja) clavata Linnaeus, 1758 et note sur la validité de Raja (Raja) capensis Müller et Henle, 1841. Cybium 5(4):23-39.
  • Chiarelli, A.B. i E. Capanna, 1973. Cytotaxonomy and vertebrate evolution. Academic Press Inc., Londres.
  • Compagno, L.J.V., 1999. Checklist of living elasmobranchs. p. 471-498. A W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland, Estats Units.
  • Crespo, J., J. Gajate i R. Ponce, 2001. Clasificación científica e identificación de nombres vernáculos existentes en la base de datos de seguimiento informático de recursos naturales oceánicos. Instituto Español de Oceanografía (Madrid)
  • Dulvy, N.K. i J.D. Reynolds, 1997. Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315.
  • Eschmeyer, William N., ed. 1998. Catalog of Fishes. Special Publication of the Center for Biodiversity Research and Information, núm. 1, vol. 1-3. California Academy of Sciences. San Francisco, Califòrnia, Estats Units. ISBN 0-940228-47-5.
  • Holden, M.T., 1972. The growth rates of Raja brachyura, Raja clavata and Raja montagui as determined from tagging data. J. Cons. Int. Explor. Mer. 34(2):161-168.
  • Hughes, G.M., 1972. Distribution of oxygen tension in the blood and water along the secondary lamella of icefish gills. J. Exp. Biol. 56:481-492.
  • Kotlyar, A.N., 1984. Dictionary of names of marine fishes on the six languages. All Union Research Institute of Marine Fisheries and Oceanography, Moscou. 288 p.
  • McEachran, J.D. i K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia (2):271-290.
  • Möller, H. i K. Anders, 1986. Diseases and parasites of marine fishes. Verlag Möller, Kiel. 365 p.
  • Nygren, A., B. Nilsson i M. Jahnke, 1971. Cytological study in Hypotremata and Pleurotremata (Pisces). Hereditas 67:275-282.
  • Ricker, W.E., 1973. Russian-English dictionary for students of fisheries and aquatic biology. Fisheries Research Board of Canada, Ottawa.
  • Sanches, J.G., 1989. Nomenclatura Portuguesa de organismos aquáticos (proposta para normalizaçao estatística). Publicaçoes avulsas do I.N.I.P. Núm. 14. 322 p.
  • Stingo, V., 1979. New developments in vertebrate cytotaxonomy. II. The chromosomes of the cartilaginous fishes Genetica 50(3):227-239.
  • Vasil'ev, V.P., 1980. Chromosome numbers in fish-like vertebrates and fish. J. Ichthyol. 20(3): 1-38.
  • Wu, H.L., K.-T. Shao i C.F. Lai (eds.), 1999. Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan.


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Clavellada: Brief Summary ( الكتالونية )

المقدمة من wikipedia CA
 src= Clavellada

La clavellada, l'escrita, la bastina (per extensió errònia), la bastina clavejada, el clavell, la rajada, la rajada punxosa o la rajada vera (Raja clavata) és una espècie de peix de la família dels raids i de l'ordre dels raïformes.

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Morgath Styds ( الويلزية )

المقدمة من wikipedia CY

Pysgodyn sy'n byw yn y môr ac sy'n perthyn i deulu'r Rajidae ydy'r Morgath Styds sy'n enw benywaidd; lluosog: morgathod styds (Lladin: Raja clavata; Saesneg: Thornback ray).

Mae ei diriogaeth yn cynnwys Ewrop, Môr y Gogledd, y Môr Baltig a Chefnfor yr Iwerydd ac mae i'w ganfod ym Môr y Gogledd ac arfordir Cymru.

Ar restr yr Undeb Rhyngwladol dros Gadwraeth Natur (UICN), caiff y rhywogaeth hon ei rhoi yn y dosbarth 'Yn agos at fod dan fygythiad' o ran niferoedd, bygythiad a chadwraeth.[1]

Gweler hefyd

Cyfeiriadau

  1. Gwefan www.marinespecies.org; adalwyd 4 Mai 2014
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Morgath Styds: Brief Summary ( الويلزية )

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Pysgodyn sy'n byw yn y môr ac sy'n perthyn i deulu'r Rajidae ydy'r Morgath Styds sy'n enw benywaidd; lluosog: morgathod styds (Lladin: Raja clavata; Saesneg: Thornback ray).

Mae ei diriogaeth yn cynnwys Ewrop, Môr y Gogledd, y Môr Baltig a Chefnfor yr Iwerydd ac mae i'w ganfod ym Môr y Gogledd ac arfordir Cymru.

Ar restr yr Undeb Rhyngwladol dros Gadwraeth Natur (UICN), caiff y rhywogaeth hon ei rhoi yn y dosbarth 'Yn agos at fod dan fygythiad' o ran niferoedd, bygythiad a chadwraeth.

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Rejnok ostnatý ( التشيكية )

المقدمة من wikipedia CZ
ikona
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Rejnok ostnatý (Raja clavata) je paryba z řádu rejnoků pravých. Je nejčastějším druhem rejnokovitých v Evropě.

Popis

 src=
Rejnok ostnatý (Raja clavata)

Hřbetní strana je hnědošedá v různých odstínech, s tmavými skvrnami a světlými tečkami. Břišní strana těla je světlá. Hrudní ploutve se táhnou od rypce po obou stranách těla až k ocasu. Samičky i samci mají na hřbetě trny.

Žije na dně moří s lehce písčitým až bahnitým dnem od 20 metrů hloubky. Lze se s nimi setkat na mělčinách a v ústí řek.

Období rozmnožování je obvykle v jarních měsících kdy rejnoci jsou nejaktivnější. K oplodnění dochází v těle samice, samec se pevně přidrží samice. Samice za rok vyprodukuje až 150 vajec umístěných v pouzdru. Samice rejnoka dorůstají do 120 cm, ale samci jen 80 cm.

Rejnoci se živí kraby, krevetami, malými rybami, škeblemi a jedí dokonce i různé druhy korýšů.

Odkazy

Reference

  1. Červený seznam IUCN 2018.1. 5. července 2018. Dostupné online. [cit. 2018-08-10]
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Rejnok ostnatý: Brief Summary ( التشيكية )

المقدمة من wikipedia CZ
ikona Tento článek není dostatečně ozdrojován a může tedy obsahovat informace, které je třeba ověřit.
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Rejnok ostnatý (Raja clavata) je paryba z řádu rejnoků pravých. Je nejčastějším druhem rejnokovitých v Evropě.

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Sømrokke ( الدانماركية )

المقدمة من wikipedia DA

Sømrokken (latin: Raja clavata) er en bruskfisk i ordenen rokker, der er udbredt fra Nordnorge til Middelhavet. Den er almindelig i Vesterhavet, Skagerrak og det dybe Kattegat. Hunnen måler op til 120 cm, mens den mindre han er 70 cm. Spredt på oversiden samt i rækker på ryggens midte og ud ad halen findes store torne, der har en sømhovedlignende roddel, fra hvilken selve tornen krummer sig bagud. Sømrokken lever mest på blød bund i 20-100 meters dybde, hvor den skjuler sig, så kun øjnene stikker op over sandet.

Beskrivelse

Sømrokken lever af krabber, håising, tobis og andre bunddyr. Den yngler sommeren igennem. De glatte mørke ægkapsler er seks cm lange og fire cm bredde og ses ofte opskyllet på stranden. Hannens bugfinner er som hos andre rokker omdannet til parringsorganer, der næsten når til midt på halen. De kønsmodne hanner har ligesom andre ægte rokker (Rajidae) såkaldte karter, der er krumme hudtænder. De findes i felter på de vingeformede brystfinners overside nær hjørnerne.

Sømrokken styrer med halen, og "flyver" langsomt gennem vandet med bugfinnerne, der bølger op og ned. Der sidder, som hos andre rokker, to gange fem gælleåbninger på undersiden lidt bag munden.

Dens kød smager som hummer ligesom for de andre arter af ægte rokker.

Kilder/Henvisninger

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Sømrokke: Brief Summary ( الدانماركية )

المقدمة من wikipedia DA

Sømrokken (latin: Raja clavata) er en bruskfisk i ordenen rokker, der er udbredt fra Nordnorge til Middelhavet. Den er almindelig i Vesterhavet, Skagerrak og det dybe Kattegat. Hunnen måler op til 120 cm, mens den mindre han er 70 cm. Spredt på oversiden samt i rækker på ryggens midte og ud ad halen findes store torne, der har en sømhovedlignende roddel, fra hvilken selve tornen krummer sig bagud. Sømrokken lever mest på blød bund i 20-100 meters dybde, hvor den skjuler sig, så kun øjnene stikker op over sandet.

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Nagelrochen ( الألمانية )

المقدمة من wikipedia DE
 src=
Ein Nagelrochen

Der Nagelrochen (Raja clavata) ist ein Knorpelfisch aus der Familie Echte Rochen (Rajidae). Er lebt vor allem an den europäischen und afrikanischen Küsten des Atlantischen Ozeans, in der Nordsee, dem Mittelmeer und dem südwestlichen Schwarzen Meer.

Verbreitung

Der Nagelrochen ist die häufigste Rochenart der europäischen Meere. Er lebt an den Küsten des Ostatlantiks von Norwegen bis Namibia, in der Nordsee, im Skagerrak und im Kattegat. Außerdem im Mittelmeer, im südwestlichen Schwarzen Meer und möglicherweise auch noch im äußersten Südwesten des Indischen Ozeans an der Küste Südafrikas, Mosambiks und an der Südküste Madagaskars.

In deutschen Gewässern ist der Nagelrochen selten geworden. Während im Jahr 1910 in Ostfriesland noch 18.000 Glatt- und Nagelrochen gefischt wurden, wurden wegen Überfischung und Beifang ab 1980 fast keine Nagelrochen mehr im deutschen Wattenmeer gesichtet. Erst seit 2010 steigt ihre Zahl wieder.[1]

Merkmale

Der Körper der Fische ist rhombisch, die Schnauze kurz, die flügelartigen Brustflossen spitz. Die Oberseite ist bestachelt und rau, braun und mit hellen und dunklen Flecken gemustert, die Unterseite weiß. Der Schwanz trägt weit hinten zwei kleine Rückenflossen und ist mit hellen und dunklen Querbändern versehen. Charakteristisch ist die Mittelreihe schlanker, nagelähnlicher Dornen auf Rücken und Schwanzoberseite. Ältere Tiere bekommen noch zusätzliche Dornen auf Ober- und Unterseite der Flügel. Die Männchen erreichen eine Größe von 70 Zentimetern, Weibchen werden bis zu 120 Zentimetern lang.

Lebensweise

Nagelrochen halten sich auf sandigen und schlammigen Weichböden in Tiefen von 20 bis 300 Metern auf, im östlichen Ionischen Meer in Tiefen von 300 bis 577 Metern. Sie ernähren sich von bodenbewohnenden Wirbellosen, vor allem von Krebstieren, aber auch von kleinen Fischen.

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Skelettpräparat eines Nagelrochens

Wie alle Rajidae sind sie ovipar, legen also Eier. Die Paarungszeit ist im Frühjahr. Etwa täglich legen die Weibchen einige Wochen danach ein Ei ab, insgesamt 50 bis 170 in einem Jahr. Die Eier sind von einer viereckigen, mit Haltefäden an jeder Ecke versehen Hornkapsel umhüllt, die fünf bis neun Zentimeter lang und 3,4 bis 6,8 Zentimeter breit sind. Nach vier bis fünf Monaten schlüpfen die Jungtiere bei einer Körperlänge von etwa zwölf Zentimeter.

Literatur

  • Bent J. Muus, Jørgen G. Nielsen: Die Meeresfische Europas in Nordsee, Ostsee und Atlantik. Kosmos, Stuttgart 1999, ISBN 3-440-07804-3.
  • Hans A. Baensch, Robert A. Patzner: Mergus Meerwasser-Atlas Band 6 Non-Perciformes (Nicht-Barschartige), Mergus-Verlag, Melle, ISBN 3-88244-116-X.

Einzelnachweise

  1. Seltener Gast im Wattenmeer: Nagelrochen gesichtet. In: NDR 1 Niedersachsen. 5. Juli 2020, abgerufen am 6. Juli 2020.
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Nagelrochen: Brief Summary ( الألمانية )

المقدمة من wikipedia DE
 src= Ein Nagelrochen

Der Nagelrochen (Raja clavata) ist ein Knorpelfisch aus der Familie Echte Rochen (Rajidae). Er lebt vor allem an den europäischen und afrikanischen Küsten des Atlantischen Ozeans, in der Nordsee, dem Mittelmeer und dem südwestlichen Schwarzen Meer.

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Raja clavata ( الإسكتلنديون )

المقدمة من wikipedia emerging languages

The Raja clavata, kent as the thorny-back or jennie skate, is a speshies o ray fish in the faimily Rajidae.[2]

References

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Raja clavata: Brief Summary ( الإسكتلنديون )

المقدمة من wikipedia emerging languages

The Raja clavata, kent as the thorny-back or jennie skate, is a speshies o ray fish in the faimily Rajidae.

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Түлкүкөөрчөк ( القيرغستانية )

المقدمة من wikipedia emerging languages
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Түлкүкөөрчөк.

Түлкүкөөрчөк (лат. лат. Raja clavata) — деңизде жашоочу көөкөрчөк балыктардын бир түрү.

Колдонулган адабияттар

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Thornback ray ( الإنجليزية )

المقدمة من wikipedia EN

The thornback ray (Raja clavata), or thornback skate, is a species of ray fish in the family Rajidae.[2]

Distribution

The Thornback ray is found in the Atlantic coastal waters of Europe and western Africa. It is also present from south Africa to the southwestern Indian Ocean [2] and in the Mediterranean and Black Seas.[3] It is native possibly as far south as Namibia and South Africa.[2]

Its natural habitats are open large seas and shallow seas. It is sometimes seen trapped in large estuarine pools at low tide.[2]

The thornback ray is probably one of the most common rays encountered by divers.

Habitat

The thornback ray is usually found on sedimentary seabeds such as mud, sand or gravel at depths between 10-60m. Juvenile fish feed on small crustaceans, particularly amphipods and bottom-living shrimps; adults feed on crabs, shrimps and small fish.

Description

Like all rays, the thornback ray has a flattened body with broad, wing-like pectoral fins. The body is kite-shaped with a long, thorny tail. The back is covered in numerous thorny spines, as is the underside in older females.[2][4]

Adult fish can grow to 1 metre (3.3 ft) in length, although most are less than 85 centimetres (33 in). This ray can weigh from 4.5 to 8.75 lb (2 to 4 kg).[5]

Their colours vary from light brown to grey with darker blotches and numerous small darker spots and yellow patches. Sometimes the yellow patches are surrounded by small dark spots. The underside is creamy-white with a greyish margin. When threatened they can appear black.[6][7]

In sexually mature fish, some of the spines are thickened with button-like bases (known as bucklers). These are particularly well developed on the tails and backs of sexually mature females.[7]

Personificated face of a Raja clavata .

Taxonomy

Raja clavata, the thornback ray (or thornback skate, roker), was named by Carl Linnaeus in the 10th edition of Systema Naturae in 1758, in the genus Raja of the Order Rajiformes in the Family Rajidae.[2]

It is one of about 13 species of skate (family Rajidae) that are known from the North Sea and adjacent Atlantic waters.

Common names

Raja clavata at the Aquarium de Vannes.

Common names include:[8]

  • Danish — Sømrokke.
  • Dutch — Stekelrog.
  • English — Roker / Thornback ray / Dorn.
  • Estonian — Astelrai.
  • Faeroese — Naglaskøta.
  • Finnish — Okarausku.
  • French — Raie bouclée.
  • German — Nagelrochen.
  • Icelandic — Dröfnuskata.
  • Latvian — Dzelkņraja.
  • Norwegian — Piggskate.
  • Polish — Raja nabijana a. ciernista.
  • Portuguese — Raia lenga / Raia brocheada
  • Russian — Колючий скат.
  • Spanish — Raya de clavos.
  • Swedish — Knaggrocka.

Morphometrics

Dorsal spines : 0; Anal spines: 0; Anal soft rays: 0. ; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length; pectoral fins with clear angles on lateral side; triangular pelvic fins . Dorsally prickly; large females also prickly throughout their ventral surface; young and large males prickly along the borders of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults . Max length : 105 cm male/unsexed; 139.0 cm (female); common length : 85.0 cm ; max. weight: 18.0 kg.

Top side view of Thornback ray on a white background.

Life cycle

Oviparous. Polyandrous species. Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms . Up to 170 egg cases can be laid by a single female in a year, average fecundity around 48-74 eggs. In northwestern Europe, egg cases are laid during spring, and in the Mediterranean during winter and spring. Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo. Capsules are 5.0-9.0 cm long without the horns and 3.4-6.8 cm wide. Egg cases are anchored with an adhesive film.

Embryos feed solely on yolk. Egg cases hatch after about 4–5 months and pups are about 11–13 cm.

Mating season is from February to September, peaking in June. Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males. Mating does not occur in the Baltic Sea.[2] This is one of the fish used by the marine leech Pontobdella muricata as a host.[9]

British Isles population
Skeletal mount

A search about the growth and maturation of Raja clavata in the Solway Firth (part of the border between Cumbria, England and Dumfries and Galloway, Scotland) shows that the males and females appear to mature at 42 and 45 cm in disc width respectively. The Solway population is heavily exploited by an unrestricted commercial fishery and a considerable proportion (48.6%) of the retained catch is immature. It is suggested that fishing pressure has brought about a reduction in the size at which female fish mature.[10]

See also

References

  1. ^ Ellis, J. (2016). "Raja clavata". IUCN Red List of Threatened Species. 2016: e.T39399A103110667. doi:10.2305/IUCN.UK.2016-3.RLTS.T39399A103110667.en. Retrieved 15 November 2021.
  2. ^ a b c d e f g Fishbase.org: Raja clavata
  3. ^ Guide of Mediterranean Skates and Rays (Raja clavata). Oct. 2022. Mendez L., Bacquet A. and F. Briand. http://www.ciesm.org/Guide/skatesandrays/raja-clavata
  4. ^ Filaman.ifm: Morphology of Raja clavata
  5. ^ Kindersley, Dorling (2005) [2001]. Animal. New York City: DK Publishing. ISBN 0-7894-7764-5.
  6. ^ Shark-references.com: Lists/literature by V.A. Vanov
  7. ^ a b Oceanario.pt: Raja clavata
  8. ^ http://www.ices.dk/explore-us/projects/EU-RFP/EU Repository/ICES ICES Fish Map: species factsheet for Raja clavata
  9. ^ "Pontobdella muricata Linnaeus, 1758". SeaLifeBase. Retrieved 27 December 2019.
  10. ^ Nottage, A. S. (1983). "Growth and maturation of roker Raja clavata L. in the Solway Firth". Journal of Fish Biology. 23: 43–48. doi:10.1111/j.1095-8649.1983.tb02880.x.

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Thornback ray: Brief Summary ( الإنجليزية )

المقدمة من wikipedia EN

The thornback ray (Raja clavata), or thornback skate, is a species of ray fish in the family Rajidae.

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Raja clavata ( الإسبانية، القشتالية )

المقدمة من wikipedia ES

La raya común o raya de clavos (Raja clavata) es una especie de peces de la familia de los Rajidae en el orden de los Rajiformes. Los machos pueden llegar alcanzar los 105 cm de longitud total y las hembras 120. Se alimenta de animales bentónicos, preferiblemente crustáceos. Entre sus depredadores se encuentra Chelidonichthys gurnardus.

Es ovíparo y las hembras ponen huevos envueltos en una cápsula córnea.[1][2]

Es un pez marino, de aguas templadas (70 ° N-29 ° S, 25 ° W-42 ° E), y demersal que vive entre los 20 y los 600 m de profundidad. Se encuentra en el océano Atlántico oriental (desde Islandia, Noruega, mar del Norte y mar Báltico, hasta Marruecos y Namibia), el Mediterráneo y el mar Negro.

Referencias

  1. Dulvy, N.K. i J.D. Reynolds, 1997. Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315.
  2. Breder, C.M. i D.E. Rosen, 1966. Modes of reproduction in fishes. T.F.H. Publications, Neptune City, Nueva Jersey, Estados Unidos. 941 p.

Bibliografía

  • Fritzsch, B. i P. Moller, 1995. A history of electroreception. p. 39-55. A: P. Moller (ed.) Electric fishes: history and behavior. Fish and Fisheries Series 17. Chapman & Hall, Londres.
  • McEachran, J.D. i K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia (2):271-290.
  • Wu, H.L., K.-T. Shao i C.F. Lai (eds.), 1999. Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwán.

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Raja clavata: Brief Summary ( الإسبانية، القشتالية )

المقدمة من wikipedia ES

La raya común o raya de clavos (Raja clavata) es una especie de peces de la familia de los Rajidae en el orden de los Rajiformes. Los machos pueden llegar alcanzar los 105 cm de longitud total y las hembras 120. Se alimenta de animales bentónicos, preferiblemente crustáceos. Entre sus depredadores se encuentra Chelidonichthys gurnardus.

Es ovíparo y las hembras ponen huevos envueltos en una cápsula córnea.​​

Es un pez marino, de aguas templadas (70 ° N-29 ° S, 25 ° W-42 ° E), y demersal que vive entre los 20 y los 600 m de profundidad. Se encuentra en el océano Atlántico oriental (desde Islandia, Noruega, mar del Norte y mar Báltico, hasta Marruecos y Namibia), el Mediterráneo y el mar Negro.

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Arraia gastaka ( الباسكية )

المقدمة من wikipedia EU

Arraia gastaka (Raja clavata) ur gaziko arrain kartilaginosoa da, Rajidae familiakoa[1].

Ekialdeko Ozeano Atlantikoan bizi da.

Erreferentziak

  1. Bigelow, H.B. & W.C. Schroeder (1953) Fishes of the Western North Atlantic, Part 2 Sears Foundation for Marine Research, Yale University 352–362. or..



Biologia Artikulu hau biologiari buruzko zirriborroa da. Wikipedia lagun dezakezu edukia osatuz.
(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Arraia gastaka: Brief Summary ( الباسكية )

المقدمة من wikipedia EU

Arraia gastaka (Raja clavata) ur gaziko arrain kartilaginosoa da, Rajidae familiakoa.

Ekialdeko Ozeano Atlantikoan bizi da.

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Okarausku ( الفنلندية )

المقدمة من wikipedia FI

Okarausku (Raja clavata) on itäisen Atlantin rauskukalalaji. Sitä tavataan etupäässä Euroopan ja Afrikan rannikoilla.

Okarausku viihtyy tavallisesti melko syvällä ja hiekkapohjalla. Ravinto koostuu pääasiassa nilviäisistä ja kaloista. Kyseessä on Skandinavian ja Yhdistyneen kuningaskunnan yleisin rauskulaji. Ihmiselle okarausku on vaaraton.[2]

Pisimmillään okarausku voi olla noin 120 cm ja painavimmillaan jopa 18 kg.[3] Laji on ovipaarinen eli sen munat kehittyvät munakoteloissa emon ulkopuolella.

Okarausku on tätä nykyä silmälläpidettävä laji.[1] Kantaa ei siis voi pitää elinvoimaisena muttei vaarantuneenakaan. Laji on viime vuosikymmeninä selvästi vähentynyt etenkin pohjoisemman Euroopan vesillä.[4]

Lähteet

  1. a b Ellis, J.: Raja clavata IUCN Red List of Threatened Species. Version 2013.1. 2005. International Union for Conservation of Nature, IUCN, Iucnredlist.org. Viitattu 03.08.2013. (englanniksi)
  2. EFSA: Thornback Ray (Raja clavata)
  3. Raja clavata (peilipalvelin) FishBase. Froese, R. & Pauly, D. (toim.). (englanniksi)
  4. DISSERTATIONS University of Groningen

Aiheesta muualla

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Okarausku: Brief Summary ( الفنلندية )

المقدمة من wikipedia FI

Okarausku (Raja clavata) on itäisen Atlantin rauskukalalaji. Sitä tavataan etupäässä Euroopan ja Afrikan rannikoilla.

Okarausku viihtyy tavallisesti melko syvällä ja hiekkapohjalla. Ravinto koostuu pääasiassa nilviäisistä ja kaloista. Kyseessä on Skandinavian ja Yhdistyneen kuningaskunnan yleisin rauskulaji. Ihmiselle okarausku on vaaraton.

Pisimmillään okarausku voi olla noin 120 cm ja painavimmillaan jopa 18 kg. Laji on ovipaarinen eli sen munat kehittyvät munakoteloissa emon ulkopuolella.

Okarausku on tätä nykyä silmälläpidettävä laji. Kantaa ei siis voi pitää elinvoimaisena muttei vaarantuneenakaan. Laji on viime vuosikymmeninä selvästi vähentynyt etenkin pohjoisemman Euroopan vesillä.

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Raja clavata ( الفرنسية )

المقدمة من wikipedia FR

La raie bouclée (Raja clavata) est une espèce de raies de l'ordre des Rajiformes que l'on rencontre de la Norvège à l'Afrique du Sud, en Méditerranée, autour des îles Britanniques, et même jusqu'en Islande. Sur le plan commercial, c'est l'espèce de rajidé la plus importante des pêches françaises. C'est également la plus recherchée au point de vue gustatif.

Dénominations

Noms normalisés

Noms FAO :

  • Français : raie bouclée.
  • Anglais : thornback ray.
  • Espagnol : raya de claros.

Autre dénomination de vente admise : raie.

Noms français régionaux

  • Manche : clouée (Boulogne), raie grise (Boulogne, Port-en-Bessin, Cherbourg, Granville, La Rochelle), raie grise bouclée (Cherbourg), raie frisée, raitène (Cotentin[1]), grisard (Granville), rea griz (Bretagne).
  • Atlantique : raie verte (Douarnenez), raie grise (Concarneau), ikara,

zerra bouké (côte Basque).

  • Méditerranée : clavaillada (Port-Vendres), clavelada, clavelado, clabelada (Languedoc, Provence), razza spiniza (Bastia).

Répartition géographique

Rare au sud-est de l'Islande, au nord-ouest de la Norvège et autour des îles Féroé. Commune en mer du Nord et dans l'Atlantique nord-est, de l'Écosse à la Mauritanie. Méditerranée, mer Noire, Atlantique sud-est et océan Indien sud-ouest.

Répartition bathymétrique

Elle vit de la côte à 500 m de profondeur, les concentrations les plus élevées se trouvant entre -10 et -60 m.

Caractères distinctifs

La raie bouclée est la seule raie européenne à avoir la queue colorée de zones transversales alternativement sombres et claires, ce qui lui donne un aspect annelé. Elle se caractérise aussi par la présence fréquente sur le dos et sur le ventre de boucles, sortes de grosses épines recourbées à base circulaire. Chez les individus en provenance de certains secteurs de pêche, comme le golfe de Gascogne, ces boucles peuvent être totalement absentes. Il faut également souligner que l'on ne doit pas tenir compte de la coloration dorsale qui peut présenter de très fortes variations d'un spécimen à l'autre.

  • Taille maximale : 115 cm.
  • Taille commune : 40–105 cm.

Photos

Notes et références

  1. Raie bouclée, n° 2658, base DORIS.

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Raja clavata: Brief Summary ( الفرنسية )

المقدمة من wikipedia FR

La raie bouclée (Raja clavata) est une espèce de raies de l'ordre des Rajiformes que l'on rencontre de la Norvège à l'Afrique du Sud, en Méditerranée, autour des îles Britanniques, et même jusqu'en Islande. Sur le plan commercial, c'est l'espèce de rajidé la plus importante des pêches françaises. C'est également la plus recherchée au point de vue gustatif.

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Roc garbh ( الأيرلندية )

المقدمة من wikipedia GA

Is iasc é an roc garbh. Is baill d'fhine na Rajidae iad.


Ainmhí
Is síol ainmhí é an t-alt seo. Cuir leis, chun cuidiú leis an Vicipéid.
Má tá alt níos forbartha le fáil i dteanga eile, is féidir leat aistriúchán Gaeilge a dhéanamh.


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Raja clavata ( الإيطالية )

المقدمة من wikipedia IT

La razza chiodata[1] (Raja clavata Linnaeus, 1758), conosciuta commercialmente con il nome generico di razza[2] , è una specie di pesce cartilagineo appartenente alla famiglia Rajidae. L'habitat naturale è rappresentato sia dal mare aperto che da acque costiere, dell'Europa, della costa atlantica dell'Africa, probabilmente fino alla Namibia e al Sudafrica in un range batimetrico che varia da 0 a 100 m circa. Talvolta viene anche vista in trappola in grandi pozze naturali che si formano per la bassa marea nei pressi di grandi estuari.

Descrizione

 src=
Mandibola di Raja clavata
 src=
Coda di Raja clavata

La Raja clavata è probabilmente una delle specie di razza che gli appassionati di subacquea incontrano con maggiore frequenza nelle acque dei mari italiani e del Mediterraneo.

Come tutte le razze ha un corpo appiattito con ampie pinne pettorali simili a delle ali. Il corpo è trapezoidale, a forma di aquilone con una coda lunga e spinosa, interamente ricoperta da dentelli dermici ricurvi e pungenti, come anche il resto dell'intero corpo, il che gli conferisce una maggiore idrodinamicità. Gli adulti possono crescere fino a 1 m di lunghezza anche se la maggior parte sono inferiori a 85 cm. Questa specie di razza pesa da 2 a 3,98 kg.

Negli esemplari sessualmente maturi alcune delle spine sono inspessite e più resistenti con una base più larga (note come "scudi"). Queste sono particolarmente ben sviluppate sulla coda come è visibile nell'immagine a lato e sul retro delle femmine sessualmente mature.

Il colore varia dal marrone chiaro al grigio, con numerose piccole macchie più scure e macchie gialle. A volte le macchie gialle sono circondate da piccole macchie scure. La parte inferiore è bianco-crema, con un margine grigio.

Habitat

La razza chiodata vive di solito su fondali fangosi, sabbiosi o ghiaiosi ad una profondità media tra i 10-60 m. I giovani esemplari si nutrono di piccoli crostacei, in particolare anfipodi e gamberetti. Gli adulti si nutrono invece di granchi, gamberi e piccoli pesci.

Note

  1. ^ vedi: (G.U. della Repubblica Italiana 2ª Serie speciale - n. 22 del 18/03/2013) e (G.U. della Repubblica Italiana 2ª Serie speciale - n. 24 del 26/03/2018) e (Regolamento (CE) N. 1638/2001) e (Regolamento (CE) N. 1581/2004) e (Regolamento (CE) N. 216/2009) e (Regolamento (UE) 2016/458) e (Regolamento (UE) 2017/127) e (Regolamento (UE) 2018/120) e (Regolamento (UE) 2019/124) e (Decisione di Esecuzione (UE) 2016/1251)
  2. ^ Decreto Ministeriale n°19105 del 22 settembre 2017 - Denominazioni in lingua italiana delle specie ittiche di interesse commerciale, su politicheagricole.it.

Bibliografia

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Raja clavata: Brief Summary ( الإيطالية )

المقدمة من wikipedia IT

La razza chiodata (Raja clavata Linnaeus, 1758), conosciuta commercialmente con il nome generico di razza , è una specie di pesce cartilagineo appartenente alla famiglia Rajidae. L'habitat naturale è rappresentato sia dal mare aperto che da acque costiere, dell'Europa, della costa atlantica dell'Africa, probabilmente fino alla Namibia e al Sudafrica in un range batimetrico che varia da 0 a 100 m circa. Talvolta viene anche vista in trappola in grandi pozze naturali che si formano per la bassa marea nei pressi di grandi estuari.

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Stekelrog ( البلجيكية الهولندية )

المقدمة من wikipedia NL

De stekelrog of gewone rog (Raja clavata) is een rog uit de familie Rajidae.

Kenmerken

Deze rog is bruingrijs met donkere vlekken en stipjes, de buikzijde is lichter. Hij heeft stekels op de rug en de staart, en ook de buik is stekelig. Een stekelrog wordt ongeveer 85 cm lang.

Leefwijze

Deze dieren liggen meestal op de zeebodem te wachten op een passerende prooi. Ze zijn nagenoeg onzichtbaar dankzij hun camouflage.

Verspreiding en leefgebied

Deze kraakbeenvis komt voor in ondiep water met modder- en zandbodem, onder andere in de Atlantische Oceaan, de Middellandse Zee en Zwarte Zee.

Relatie met de mens

Stekelroggen zijn in veel openbare aquaria te bezichtigen. Soms worden ze zoals in Nausicaä in Boulogne-sur-Mer in open "aaibakken" gehouden, omdat het dier totaal ongevaarlijk is, in tegenstelling tot bijvoorbeeld de giftige pijlstaartrog.

Status aan de Nederlandse en Belgische kusten

Stekelroggen komen voor op een diepte tussen de 2 en 60 m in de Noordzee. Zij zetten hun eieren af in ondiep water. De zwarte eikapsels van de stekelrog kan men langs de Belgische en Nederlandse kust aantreffen. Omdat ze betrekkelijk weinig eieren produceren, zijn ze erg gevoelig voor overbevissing.[2] De soort staat als gevoelig op de internationale Rode Lijst van de IUCN[1] en als kwetsbaar op de Nederlandse Rode Lijst.

Externe links

Bronnen, noten en/of referenties
  1. a b (en) Stekelrog op de IUCN Red List of Threatened Species.
  2. Nijssen, H. & S.J. de Groot, 1987. De vissen van Nederland, KNNV, Utrecht.
  • David Burnie (2001) - Animals, Dorling Kindersley Limited, London. ISBN 90-18-01564-4 (naar het Nederlands vertaald door Jaap Bouwman en Henk J. Nieuwenkamp).
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Stekelrog: Brief Summary ( البلجيكية الهولندية )

المقدمة من wikipedia NL

De stekelrog of gewone rog (Raja clavata) is een rog uit de familie Rajidae.

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Raja nabijana ( البولندية )

المقدمة من wikipedia POL
Commons Multimedia w Wikimedia Commons

Raja nabijana[3], raja ciernista[4], płaszczka nabijana[5], płaszczka kolczasta[5] (Raja clavata) – gatunek ryby chrzęstnoszkieletowej z rodziny rajowatych (Rajidae).

Zasięg występowania

Wschodni Atlantyk od północnej Norwegii i południowej Islandii do południowej Afryki oraz Morze Śródziemne i Morze Czarne.

Występuje na piaszczystym lub mulistym dnie, zwykle na głębokości od 20 do 60 m, ale spotykana jest także na głębokościach większych do 500 m.

Cechy morfologiczne

Osiąga maksymalnie do 70 cm (samce) i do 125 cm (samice) długości. Ciało spłaszczone grzbietobrzusznie o kształcie rombowatej tarczy, płetwa piersiowa z ostrymi krawędziami. Pysk krótki, rozwartokątny. Na skórze występują kolce, tak na stronie wierzchniej jak również dolnej. Dorosłe osobniki posiadają na stronie grzbietowej liczne, duże kolce osadzone w skórze razem z szerokimi, gładkimi tarczkami. U samic i młodych osobników ciągnie się wzdłuż grzbietu i trzonu ogonowego długi szereg kolców. U samców szereg kolców wstępuje tylko na trzonie ogonowym. Samce dodatkowo posiadają ruchome kolce na płetwach piersiowych. Uzębienie składa się z 36–45 zębów, u samców spiczaste a samic spłaszczonych. Dwie małe płetwy grzbietowe o takiej samej wielkości znajdują się na końcu trzonu ogonowego. Płetwa ogonowa mała. Brak płetwy odbytowej. Posiada narządy elektryczne.

Strona grzbietowa jasnoszara lub jasnobrązowa, u młodych osobników wyraźnie widoczne brązowo obwiedzione jasne plamy oraz małe czarne punkty. U starszych osobników plamy zanikają. Strona brzuszna biała z ciemnymi krawędziami.

Odżywianie

Żywi się głównie krabami, ale także wieloszczetami, mięczakami, jeżowcami i małymi rybami.

Rozmnażanie

Ryba jajorodna. Dojrzałość płciową samice osiągają w 9 roku życia, a samce w 7 roku.

Przypisy

  1. Raja clavata, w: Integrated Taxonomic Information System (ang.).
  2. Raja clavata. Czerwona księga gatunków zagrożonych (IUCN Red List of Threatened Species) (ang.).
  3. Stanisław Rutkowicz: Encyklopedia ryb morskich. Gdańsk: Wydawnictwo Morskie, 1982. ISBN 83-215-2103-7.
  4. Eugeniusz Grabda, Tomasz Heese: Polskie nazewnictwo popularne krągłouste i ryby - Cyclostomata et Pisces. Koszalin: Wyższa Szkoła Inżynierska w Koszalinie, 1991.
  5. a b G. Nikolski: Ichtiologia szczegółowa. Tłum. Franciszek Staff. Warszawa: Państwowe Wydawnictwo Rolnicze i Leśne, 1970.

Bibliografia

  • Fritz Terofal, Claus Militz: Ryby morskie. Warszawa: Świat Książki, 1996. ISBN 83-7129-306-2.
  • Raja clavata. (ang.) w: Froese, R. & D. Pauly. FishBase. World Wide Web electronic publication. www.fishbase.org [dostęp 1 sierpnia 2009]
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Raja nabijana: Brief Summary ( البولندية )

المقدمة من wikipedia POL

Raja nabijana, raja ciernista, płaszczka nabijana, płaszczka kolczasta (Raja clavata) – gatunek ryby chrzęstnoszkieletowej z rodziny rajowatych (Rajidae).

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Raja clavata ( البرتغالية )

المقدمة من wikipedia PT

Raja clavata é uma espécie de peixe da família Rajidae conhecida pelo nome comum de raia-lenga.

Descrição

Pode ser encontrada nos seguintes países: Albânia, Alemanha, Argélia, Angola, Bélgica, Benim, Bósnia e Herzegovina, Bulgária, Camarões, República do Congo, República Democrática do Congo, Croácia, Chipre, Costa do Marfim, Dinamarca, Egito, Guiné Equatorial, Ilhas Feroé, França, Gabão, Gana, Gibraltar, Grécia, Guiné, Guiné-Bissau, Islândia, Irlanda, Israel, Itália, Líbano, Libéria, Líbia, Madagáscar, Malta, Mauritânia, Maurícia, Monaco, Marrocos, Holanda, Nigéria, Noruega, Polónia, Portugal, Roménia, Rússia, São Tomé e Príncipe, Senegal, Serra Leoa, Eslovénia, Espanha, Suécia, Síria, Togo, Tunísia, Turquia, Reino Unido, possivelmente Namíbia e possivelmente África do Sul.

Os seus habitats naturais são: mar aberto e mar costeiro.

Referências

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Raja clavata: Brief Summary ( البرتغالية )

المقدمة من wikipedia PT

Raja clavata é uma espécie de peixe da família Rajidae conhecida pelo nome comum de raia-lenga.

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Vatos ( الرومانية، المولدوفية )

المقدمة من wikipedia RO
Identificare externă pentru
Raja clavata NCBI 7781 ITIS 160901 WoRMS 105883 Vedeți și clasificarea de la Wikispecii

Vatosul (Raja clavata) este un pește marin cartilaginos din ordinul rechinilor batoizi (plați), familia Rajidae, genul Raja. [2] Mai este denumit și „vulpe de mare”.

Habitat

Zonele în care poate fi întâlnit sunt cele subtropicale (70°N - 29°S, 25°V – 42°E), estul și nordul Oceanului Atlantic, Marea Nordului, Marea Baltică, Marea Mediterană și Marea Neagră, sud-vestul Oceanului Indian.

Este un pește criofil, care duce o viață solitară, îngropat în nisip la adâncimea de 60-70 m, cu apă mai rece și mai sărată, alături de calcan și morun.

Descriere

Vatosul are corpul turtit lateral și acoperit cu plăci osoase, înzestrate cu câte un spin. Lungimea masculului ajunge până la 70 cm, femela până la 1,5 m, greutatea 5-8 kg. Pe fața ventrală a corpului se află 5 deschideri branhiale. Pe spate are numeroși țepi care pe abdomen sunt mai rari, de asemeni și pe muchia cozii.
Ochii sunt mari și așezați pe partea dorsală, apropriați între ei. Gura se află pe fața ventrală.
Culoarea este schimbătoare după mediu, de obicei variază între cenușiu, brun roșcată și galben-cenușiu cu pete neregulate. [3]
Este ovipar, în lunile martie-mai se apropie de țărm până la 25-45 m, pentru a depune 6-10 ouă. După depunere, se retrage la adâncimi mai mari până în septembrie-noiembrie, când se apropie de țărm odată cu curenții reci.
Se hrănește cu pești, crustacei și moluște. [4]

Valoare comercială

Carnea vatosului este foarte apreciată. Se consumă înotătoarele pectorale (comercializate ca „aripi de vulpe”), considerate delicatese, fiind asemănătoare la gust cu carnea de crab. Ficatul reprezintă 6,5% din greutatea totală a corpului și conține 60-67% grăsimi, fiind foarte bogat în vitamina A. Pielea vatosului, prelucrată și colorată în verde este folosită în pielărie sub numele de „piele de rechin”.

Referințe

  1. ^ Ellis, J. (2005). Raja clavata. Lista roșie a speciilor periclitate IUCN. Versiunea 2012.1. International Union for Conservation of Nature. Accesat în 08 August 2015. Verificați datele pentru: |access-date= (ajutor)
  2. ^ marinespecies.org: Raja species - WoRMS - World Register of Marine Species
  3. ^ Dan Manoleli, Teodor T. Nalbant: Viața în Marea Neagră, Editura științifică, București, 1976, pag.211-214
  4. ^ ecomareaneagra.wordpress.com: Peștii din Marea Neagră

Vezi și

Legături externe

Commons
Wikimedia Commons conține materiale multimedia legate de Vatos
Wikţionar
Caută „vatos” în Wikționar, dicționarul liber.
Wikţionar
Caută „vulpe-de-mare” în Wikționar, dicționarul liber.
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Vatos: Brief Summary ( الرومانية، المولدوفية )

المقدمة من wikipedia RO

Vatosul (Raja clavata) este un pește marin cartilaginos din ordinul rechinilor batoizi (plați), familia Rajidae, genul Raja. Mai este denumit și „vulpe de mare”.

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Trnjevka ( الإسبانية، القشتالية )

المقدمة من wikipedia SL

Trnjevka (znanstveno ime Raja clavata) je morska riba iz družine pravih raž.

Opis

Trnjevka je svoje ime dobila po trnastih izrastkih, ki jih ima po telesu. Trnjevka je, kot vsi skati, sploščena riba hrustančnica, ki ima obliko romba. Prsne plavuti so od glave do repa široko prirasle na telo. Rep trnjevke je dolg in bičast, ribi pa ne pomaga pri plavanju[1]. Oči ima trnjevka na hrbtu, usta pa na spodnji strani telesa. Po hrbtu je trnjevka rumeno rjava ali sivo zelena (odvisno od podlage na kateri živi) in je posuta s pegami, po trebuhu pa je okrasto bele barve s sivkastim pridihom. Odrasle ribe lahko dosežejo 1 m v dolžino, običajno pa dosežejo le okoli 85 cm. Tehtajo od 2 do 4 kg[2].

Razširjenost in uporabnost

Trnjevke živijo na blatnem ali peščenem dnu, v katerega se lahko tudi delno zakopljejo, ko čakajo na plen. To je riba srednjih globin, ki se zadržuje v globinah med 10 in 80 metrov. Mlade ribe se hranijo pretežno z mahnimi rakci in rakovicami, odrasle pa k temu dodajo še manjše ribe.

Razširjene so v obalnih vodah vzhodnega Atlantika, njihov atlantski habitat pa seže vse do Namibije in Južne Afrike. Razširjena je tudi v Sredozemlju in Jadranu. Samice ležejo velika jajca v velikih rjavih lupinah.

Trnjevke lovijo od pomladi do poznega poletja, najpogosteje na parangal, neredko pa se znajdejo tudi v mrežah vlečnicah.

Meso trnjevk je mehko, belo in rahlo kiselkastega okusa, sodi pa med najboljše med vsemi hrustančnicami. Najpogosteje jo pripravljajo v brodetu, pa tudi kuhano. Kulinarično sodi med drugo ali tretjerazredne ribe.

Viri in reference

  1. "Morpholgy" in [1]
  2. Burnie, David; Wilson, Don E. (2001). Animal. New York City: DK Publishing. ISBN 0-7894-7764-5.

http://www.habitas.org.uk/marinelife/species.asp?item=ZF1360

  • Zei, M. (1983). Morski ribji trg. Ljubljana: Mladinska knjiga.
Wikivrste vsebujejo še več podatkov o temi: Raja clavata
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Trnjevka: Brief Summary ( الإسبانية، القشتالية )

المقدمة من wikipedia SL

Trnjevka (znanstveno ime Raja clavata) je morska riba iz družine pravih raž.

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Knaggrocka ( السويدية )

المقدمة من wikipedia SV

Knaggrocka (Raja clavata) är den vanligaste rockan i Europas hav. Den hittas bland annat på Västkustens grundare bottnar i Sverige.

Denna broskfisk känns igen på sin gulaktiga färg med ljusa fläckar omgivna av mörka ringar. Nosen är rät eller trubbvinklig (till skillnad från klorocka), och på ovansidan har den stora taggar. Undersidan är sträv.

Liksom andra broskfiskar har den munnen på undersidan (där även gälarna sitter) och ett skelett uppbyggt av brosk.

Noter

  1. ^ Ellis, J. & Walker, P. 2000. Raja clavata. Från: IUCN 2006. 2006 IUCN Red List of Threatened Species Läst 2007-01-20.
  2. ^ Naturhistoriska riksmuseets databas över fisknamn Arkiverad 29 september 2007 hämtat från the Wayback Machine.. Läst 20070120.
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Knaggrocka: Brief Summary ( السويدية )

المقدمة من wikipedia SV

Knaggrocka (Raja clavata) är den vanligaste rockan i Europas hav. Den hittas bland annat på Västkustens grundare bottnar i Sverige.

Denna broskfisk känns igen på sin gulaktiga färg med ljusa fläckar omgivna av mörka ringar. Nosen är rät eller trubbvinklig (till skillnad från klorocka), och på ovansidan har den stora taggar. Undersidan är sträv.

Liksom andra broskfiskar har den munnen på undersidan (där även gälarna sitter) och ett skelett uppbyggt av brosk.

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Морська лисиця ( الأوكرانية )

المقدمة من wikipedia UK

Будова

Розмір самців приблизно 70 — 85 см, самиць — 125 см, вага до 4 кг. Тіло морської лисиці має форму сплюснутого ромбу, який утворений зрощеними головою, тулубом та грудними плавцями. Верхня частина тіла та хвіст вкриті крупними шипами, кінці яких загнуті назад, між якими знаходяться дрібні шипи. Невелика кількість шипів є також на нижній частині тіла. На кінці хвоста розташовані два невеликих спинних плавця. Забарвлення досить мінливе, спина жовтувато-сіра або бура з темними та світлими плямами, нижня частина — біла.

Спосіб життя

Морська лисиця — донна риба. Малорухлива риба, тримається на дні, де заривається у пісок або мул. Зустрічається переважно на невеликих глибинах, але може опускатися до глибини 300 м (у Чорному морі — до 100 м). Плаває хвилеподібно, рухаючи бічними плавцями. Оскільки рот риби знаходиться на нижній частині тіла, полюючи здобич, що рухається, скат пливе таким чином, щоб опинитися над нею, потім швидко падає вниз та хапає її. Живиться донними ракоподібними (крабами, десятиногими раками), рибою, рідше молюсками. Зимує на глибині.

Розмноження

Розмножується навесні. Для нересту підходить до берегів, але уникає занадто мілких ділянок з теплою водою та опріснених вод. Самиця відкладає на дно яйця, що мають 6 — 9 см у довжину та 4 — 7 см у ширину. Плодючість від кількох десятків, до кількох сотень яєць, які відкладаються протягом всього літнього періоду. Яйця являють собою випуклі капсули, вкриті роговою оболонкою, що мають у кутках рогові відростки з пучками ниток. Цими нитками яйця заплутуються у водорості та тримаються біля дна. Зародок розвивається протягом 4,5 — 5,5 місяців. Рогова оболонка яйця захищає його від хижаків, але вона здатна пропускати морську воду, забезпечуючи таким чином дихання зародку. Мальок, що з'являється, має довжину 12 — 13 см.

Значення

Промислове значення в Україні невелике. У деяких північних країнах (Швеція, Данія) м'ясо скатів цінується за свої смакові якості.

Литература

  • Словник зоологичної номенклатури. Назви хребетних тварин. Mammalia. –Reptilia. –Amphibia. –Pisces. (Проєкт). Уклали М. Шарлемань та К. Татарко (Природничий відділ). Київ: ДВУ, 1927, 124+1 с.

Посилання

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Raja clavata ( الفيتنامية )

المقدمة من wikipedia VI

Raja clavata là một loài thuộc họ Rajidae. Nó được tìm thấy ở vùng nước ven biển of châu Âu and the Atlantic bờ biển của Africa, có thể xa về phía nam như Namibia và cả Nam Phi. Các môi trường sống tự nhiên của chúng lànhững vùng biển mở và biển nông.

Hình ảnh

Tham khảo

http://www.habitas.org.uk/marinelife/species.asp?item=ZF1360

 src= Wikispecies có thông tin sinh học về Raja clavata


Hình tượng sơ khai Bài viết về Cá sụn này vẫn còn sơ khai. Bạn có thể giúp Wikipedia bằng cách mở rộng nội dung để bài được hoàn chỉnh hơn.
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Raja clavata: Brief Summary ( الفيتنامية )

المقدمة من wikipedia VI

Raja clavata là một loài thuộc họ Rajidae. Nó được tìm thấy ở vùng nước ven biển of châu Âu and the Atlantic bờ biển của Africa, có thể xa về phía nam như Namibia và cả Nam Phi. Các môi trường sống tự nhiên của chúng lànhững vùng biển mở và biển nông.

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Морская лисица ( الروسية )

المقدمة من wikipedia русскую Википедию
Латинское название Raja clavata
Linnaeus, 1758

wikispecies:
Систематика
на Викивидах

commons:
Изображения
на Викискладе

ITIS 160901 NCBI 7781
Эта статья — о скате. Об акуле см. Обыкновенная морская лисица.

Морская лисица, или колючий скат[1], или шиповатый скат[2] (лат. Raja clavata) — наиболее распространённый европейский вид скатов. Обитает у восточного побережья Атлантического океана от Исландии, Норвегии и Северного моря до Южной Африки. Присутствует в Средиземном, Чёрного морях, был обнаружен в Азовском море[2].

Внешний вид

Тело морских лисиц ромбообразное, голова короткая, а крылоподобные грудные плавники на концах заострённые. Верхняя часть шершавая и усеянная шипами, покрытая узором из светлых и тёмных пятен. Нижняя сторона белая. На задней части хвоста, покрытого светлыми и тёмными поперечными полосками, два маленьких спинных плавника. Характерен ряд стройных иглообразных шипов на спине и верхней части хвоста. У зрелых особей имеются дополнительные шипы на верхней и нижней стороне крыльев. Самцы достигают величины от 70 см, а величина самок доходит до 120 см.

Поведение

Морские лисицы обитают на песчаном и илистом дне моря на глубине от 20 до 300 м. На востоке Ионического моря их встречали на глубине до 577 м. Они питаются обитающими на дне беспозвоночными, прежде всего ракообразными, а также небольшой рыбой.

Как и для всех скатов, для морских лисиц характерно яйцерождение. Спаривание проходит весной. После этого самки на протяжении нескольких недель ежедневно откладывают по яйцу, всего от 50 до 170 за год. По истечении четырёх-пяти месяцев из них появляются на свет детёныши, величина которых составляет при рождении около 12 см.

Примечания

  1. Решетников Ю. С., Котляр А. Н., Расс Т. С., Шатуновский М. И. Пятиязычный словарь названий животных. Рыбы. Латинский, русский, английский, немецкий, французский. / под общей редакцией акад. В. Е. Соколова. — М.: Рус. яз., 1989. — С. 42. — 12 500 экз.ISBN 5-200-00237-0.
  2. 1 2 Васильева Е. Д. Рыбы Чёрного моря. Определитель морских, солоноватоводных, эвригалинных и проходных видов с цветными иллюстрациями, собранными С. В. Богородским. — М.: ВНИРО, 2007. — С. 20-21. — 238 с. — 200 экз.ISBN 978-5-85382-347-1.
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Морская лисица: Brief Summary ( الروسية )

المقدمة من wikipedia русскую Википедию
Эта статья — о скате. Об акуле см. Обыкновенная морская лисица.

Морская лисица, или колючий скат, или шиповатый скат (лат. Raja clavata) — наиболее распространённый европейский вид скатов. Обитает у восточного побережья Атлантического океана от Исландии, Норвегии и Северного моря до Южной Африки. Присутствует в Средиземном, Чёрного морях, был обнаружен в Азовском море.

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