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Kleberg's Bluestem

Dichanthium annulatum (Forssk.) Stapf

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This is a very excellent fodder grass which is sought out by stock from a mixture of grasses and eaten in preference to all others.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Gramineae (Poaceae) in Flora of Taiwan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Comments

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Dichanthium annulatum is morphologically and cytologically very variable. For details of its variation see Mehra & Celarier in Proc. Okla. Acad. Sci. 38: 22-25.1958; Mehra in Fyton 17:157-166. 1961; and Mehra in Caryologia 17:545-556. 1964. The species is an excellent fodder grass eagerly sought by stock.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
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Flora of Pakistan Vol. 0: 280 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Description

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Culm tufted, about 1 mm in diameter, hard, nodes densely covered with silky hairs. Blade 8-10 cm long by 3-4 mm wide, surface covered with tubercle based silky hairs; ligule chartaceous, about 1 mm long, upper margin fimbriate. Inflorescence of digitate racemes, racemes 2-8, up to 5 cm long, pale purple in color. Spikelets paired, the upper ones pedicelled; the lower ones sessile, about 3 mm long. Lower glume oblong, chartaceous, as long as the spikelet, 5-nerved, lower argins inrolled, upper part 2-keeled, with long tubercled hairs along keels, lower back ciliate; upper glume chartaceous, lanceolate, margins inrolled, fimbriate, backside 1-keeled, 3-nerved; lemma linear, 1-nerved, tipped with a long flexuous awn arising from the apex, about 6 times the length of the lemma; palea oblong, hyaline, about 2.5 mm long, nerveless; anther about 1.8 mm long; lodicules 2, truncate, conspicuously several-nerved, about 0.3 mm long. Widely distributed in India, Burma, Tropical and North Africa, now introduced as a fodder in many countries.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Gramineae (Poaceae) in Flora of Taiwan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Poaceae in Flora of Taiwan @ eFloras.org
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Chang-Sheng Kuoh
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eFloras.org
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Description

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Perennial. Culms tufted, erect or sometimes straggling, 30–100 cm tall, nodes bearded with spreading hairs. Leaf sheaths terete, shorter than internodes; leaf blades flat, 8–30 × 0.2–0.4 cm, glabrous or adaxial surface stiffly pilose, margins smooth or scaberulous, apex acuminate; ligule 1–2 mm, lacerate. Inflorescence terminal; peduncle glabrous; racemes 2–8, subdigitate, suberect, 4–5 cm, with 0–6 pairs of homogamous spikelets. Sessile spikelet 3–5 mm; lower glume elliptic-oblong or oblong, firmly papery, 5–9-veined, subglabrous or pubescent to villous on lower back, upper flanks often with long spreading hairs, keels shortly ciliate, not or barely winged, apex obtuse; upper glume ciliate along keel and margins, apex acute or obtuse; awn 1.6–2.4 cm. Caryopsis obovate. Pedicelled spikelet many-veined, pubescent to villous with spreading tubercle-based hairs. Fl. and fr. Jun–Nov. 2n = 20, 40.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 22: 604, 605, 608 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Description

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Perennial; culms 25-100 cm high, geniculately ascending. Leaf-blades 3-30 cm long, 2-7 mm wide. Inflorescence composed of (1-) 2-15 subdigitate shortly peduncled racemes, the peduncles glabrous; racemes 3-7 cm long, the spikelets subimbricate with 0-6 smaller homogamous pairs at the base; internodes and pedicels solid. Sessile spikelet narrowly oblong, 2-6 mm long; lower glume firmly cartilaginous, slightly concave, pubescent to villous below the middle with long bulbous-based hairs on the margins above, obtuse to subacute; awn 8-25 mm long.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0: 280 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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eFloras.org
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Distribution

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Tropical & N. Africa, Nepal, India, Burma.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Annotated Checklist of the Flowering Plants of Nepal Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Annotated Checklist of the Flowering Plants of Nepal @ eFloras.org
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K.K. Shrestha, J.R. Press and D.A. Sutton
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Distribution

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Guangdong, Guangxi, Guizhou, Hainan, Hubei, Sichuan, Taiwan, Yunnan [India, Indonesia, Malaysia, Myanmar, Nepal, Pakistan, Philippines; Africa, Pacific Islands; introduced in America and Australia].
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cc-by-nc-sa-3.0
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 22: 604, 605, 608 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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eFloras.org
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Distribution

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Distribution: Pakistan (Sind, Baluchistan, Punjab, N.W.FY. & Kashmir); Kenya, Tanzania and Senegal, through the Middle East to Indonesia; introduced to southern Africa, Tropical America and Australia.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0: 280 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of Pakistan @ eFloras.org
editor
S. I. Ali & M. Qaiser
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eFloras.org
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Elevation Range

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150-1100 m
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Annotated Checklist of the Flowering Plants of Nepal Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Annotated Checklist of the Flowering Plants of Nepal @ eFloras.org
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K.K. Shrestha, J.R. Press and D.A. Sutton
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eFloras.org
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Flower/Fruit

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Fl. & Fr. Per.: March-November.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of Pakistan Vol. 0: 280 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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eFloras.org
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Habitat

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Mountain slopes, disturbed ground; 100–2200 m.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 22: 604, 605, 608 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
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eFloras.org
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Synonym

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Andropogon annulatus Forsk., Fl. Aegypt. Arab. 173. 1775.
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Gramineae (Poaceae) in Flora of Taiwan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Poaceae in Flora of Taiwan @ eFloras.org
editor
Chang-Sheng Kuoh
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eFloras.org
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Synonym

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Andropogon annulatus Forsskål, Fl. Aegypt.-Arab. 173. 1775; Bothriochloa tuberculata W. Z. Fang; Dichanthium annulatum var. bullisetosum B. S. Sun & S. Wang.
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 22: 604, 605, 608 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of China @ eFloras.org
editor
Wu Zhengyi, Peter H. Raven & Hong Deyuan
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eFloras.org
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Comprehensive Description

provided by North American Flora
Andropogon nodosus (Willem.) Nash
Dichanihiufn nodosum Willem. Ann. Bot. Usteri 18: 11. 1796.
Andropogon arisiatus Poir. in Ivam. Encyc. Suppl. 1 : 585. 1810.
Andropogon molUcomus Kunth, R§v. Gram. 365. 1830.
Diplasanthum lanosum Desv. Opusc. 67. 1831.
Andropogon caricosus molUcomus Hack, in DC. Monog. Phan. 6 : 569. 1889.
Stems up to 1.5 m. tall, rather stout, usually branched above, decumbent and rooting below, softly pubescent at the apex below the inflorescence; leaf-sheaths compressed, keeled, shorter than the internodes; blades up to 3 dm. long, 4-6 mm. wide, usually rough, glabrous, or pubescent at the base; racemes in 2's-4's, sometimes solitary, 6-10 cm. long, the internodes and pedicels about one third as long as the sessile spikelets, ciliate on one margin; perfect sessile spikelet 4^5 mm. long, somewhat convolute below, the first scale chartaceous-herbaceous, broadly obovate-elliptic, about 12-nerved, the nerves vanishing below the somewhat 3-toothed apex, pubescent on the smooth shining back, the second scale much narrower and almost enclosed by the first, the foiurth scale bearing an awn 2-2.5 cm. long; pedicellate spikelet equaling the sessile one and similar to it, but flat and more pubescent, the first scale 15-19nerved, ciliate on the keels.
Type locality : Mauritius.
Distribution : Introduced into Florida, Antigua, Guadeloupe, and Barbados ; a native of the Old World tropics.
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George Valentine Nash. 1912. (POALES); POACEAE (pars). North American flora. vol 17(2). New York Botanical Garden, New York, NY
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Physical Description

provided by USDA PLANTS text
Perennials, Terrestrial, not aquatic, Rhizomes present, Stolons or runners present, Stems nodes swollen or brittle, Stems geniculate, decumbent, or lax, sometimes rooting at nodes, Stems caespitose, tufted, or clustered, Stems terete, round in cross section, or polygonal, Stem internodes hollow, Stems with inflorescence less than 1 m tall, Stems, culms, or scapes exceeding basal leaves, Leaves mostly cauline, Leaves conspicuously 2-ranked, distichous, Leaves sheathing at base, Leaf sheath mostly open, or loose, Leaf sheath smooth, glabrous, Leaf sheath and blade differentiated, Leaf blades linear, Leaf blades 2-10 mm wide, Leaf blades mostly flat, Leaf blades mostly glabrous, Ligule present, Ligule an unfringed eciliate membrane, Inflorescence terminal, Inflorescence solitary, with 1 spike, fascicle, glomerule, head, or cluster per stem or culm, Inflorescence with 2 or more spikes, fascicles, glomerules, heads, or clusters per culm, Inflorescence a panicle with narrowly racemose or spicate branches, Inflorescence a panicle with digitately arranged spicate branches, Inflorescence with 2-10 branches, Flowers bisexual, Flowers unisexual, Spikelets pedicellate, Spikelets sessile or subsessile, Spikelets dorsally compressed or terete, Spikelet less than 3 mm wide, Spikelets with 2 florets, Spikelet with 1 fertile floret and 1-2 sterile florets, Spikelets paired at rachis nodes, Spikelets in paired units, 1 sessile, 1 pedicellate, Spikelets bisexual, Spikelets unisexual, Spikelets disarticulating below the glumes, Spikelets falling with parts of disarticulating rachis or pedicel, Rachilla or pedicel glabrous, Glumes present, empty bracts, Glumes 2 clearly present, Glumes equal or subequal, Glumes equal to or longer than adjacent lemma, Glume surface hairy, villous or pilose, Glumes 3 nerved, Glume margins or apex erose-ciliate, Lemmas thin, chartaceous, hyaline, cartilaginous, or membranous, Lemma 1 nerved, Lemma glabrous, Lemma apex acute or acuminate, Lemma distinctly awned, more than 2-3 mm, Lemma with 1 awn, Lemma awn less than 1 cm long, Lemma awn 1-2 cm long, Lemma awn 2-4 cm long or longer, Lemma awned from tip, Lemma awn twisted, spirally coiled at base, like a corkscrew, Lemma awn once geniculate, bent once, Lemma awn twice geniculate, bent twice, Lemma margins thin, lying flat, Lemma straight, Callus or base of lemma evidently hairy, Callus hairs shorter than lemma, Lemma surface pilose, setose or bristly, Stamens 3, Styles 2-fid, deeply 2-branched, Stigmas 2, Fruit - caryopsis, Caryopsis ellipsoid, longitudinally grooved, hilum long-linear.
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Dichanthium annulatum

provided by wikipedia EN

Dichanthium annulatum is a species of grass in the family Poaceae. It is commonly used as a forage for livestock.

Spikelet with long awn

Common names include marvel grass, Diaz bluestem, Kleberg bluestem, Hindi grass, ringed dichanthium, sheda grass, medio bluestem (var. papillosum),[1] jargu grass, Delhi grass, vuda bluegrass, two-flowered golden-beard,[2] Santa Barbara grass.

It is native to tropical Asia, the Middle East, and parts of Africa. In India, it is very important in agriculture.[3] It has been introduced to many other parts of the world for cultivation, and it has become naturalized in some places, such as Australia.[1]

Description

This species is quite variable.[3][4] In general, it is a perennial grass often with stolons. The stems spread outwards, then grow erect at the ends. A stem may be a meter long and reach about 60 centimeters in erect height.[5] Each node on the stem is encircled with a ring of hairs.[2] The rough-haired leaf blades are up to 30 centimeters long.[5] The inflorescence is an array of purplish or green branches each up to 7 centimeters long. There are generally 2 to 9 branches, but sometimes up to 15.[2] The spikelet may be well over 2 centimeters long, including its long awn. The awn is twisted and has two bends in it.[5] The root system goes no deeper than one meter.[2][6]

This grass often reproduces by apomixis, producing seeds without fertilization. It does reproduce sexually at times. Plants can be diploid, tetraploid, or hexaploid.[2]

In cultivation

This is a popular pasture grass in many areas. It can be used in fields for grazing livestock, and cut for hay and silage. It is tolerant of varied soil conditions, including soils high in clay and sand, poorly drained soils, and soils that are somewhat alkaline and saline. It forms a turf that can stand up to grazing pressure. It can recover from fire and drought, but it is less tolerant of frost and shade. It does not require fertilizer but it does respond well to a small amount of supplemental nitrogen.[2] Horses and cattle find it very palatable.[3]

While it can aggressively outcompete many other plants, it thrives with some types of companion species, such as the grasses Bothriochloa insculpta, Dichanthium aristatum, and D. caricosum, and the legumes Medicago sativa, Stylosanthes hamata, and S. seabrana.[2]

The grass can also be used to revegetate degraded grasslands.[2] It is a very effective binding plant for erosion control.[7]

This is an especially favored pasture grass species in India. There it is familiar, widely planted, and successful.[4][6][7]

Cultivars include 'Marvel 8'.[4]

Ecology

This grass is host to a number of fungus species, such as Balansia sclerotica, Cerebella andropogonis, Chaetostroma atrum, Cochliobolus cymbopogonis, Curvularia andropogonis, C. lunata, C. robusta, Ellisiella caudata, Jamesdicksonia obesa, Phyllachora ischaemi, Physoderma dichanthicola, Pithomyces graminicola, Puccinia cesatii, P. duthiae, P. propinqua, Sclerospora dichanthicola, Sphacelotheca annulata, S. andropogonis-annulati, Tolyposporella obesa, Uredo susica, Uromyces andropogonis-annulati, U. clignyi, and Ustilago duthiei. Most are not very destructive to the grass, but it is susceptible to ergot (Claviceps spp.).[2]

It is also a host for the parasitic plant Striga lutea.[2]

This grass can escape cultivation and take hold in the wild. It is able to grow in harsh and disturbed habitat types, such as roadsides. It can become a weed.[4] It is an invasive species in Fiji, Hawaii, and New Caledonia and has displaced native grasses in large areas of south Texas.[8] It is cited as a factor in the decline of the slender rushpea (Hoffmannseggia tenella) a federally listed endangered plant of the United States.[9]

References

  1. ^ a b "Dichanthium annulatum". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA). Retrieved 21 January 2018.
  2. ^ a b c d e f g h i j Cook, B. G., et al. Dichanthium annulatum. Tropical Forages. CSIRO, DPI&F (Qld), CIAT and ILRI, Brisbane, Australia.
  3. ^ a b c Quattrocchi, U. CRC World Dictionary of Grasses: Common Names, Scientific Names, Eponyms, Synonyms, and Etymology, Volume 1. CRC Press. 2006. pg. 633.
  4. ^ a b c d Dichanthium annulatum (Forsk.) Stapf. The Center for New Crops & Plant Products. Purdue University.
  5. ^ a b c Dichanthium annulatum. Grass Manual. Flora of North America.
  6. ^ a b Dichanthium annulatum (Forsk.) Stapf. Grassland Species Profiles. FAO.
  7. ^ a b Heuzé V., Tran G., Archimède H., 2015. Marvel grass (Dichanthium annulatum). Feedipedia, a programme by INRA, CIRAD, AFZ and FAO. https://www.feedipedia.org/node/463 Last updated on May 11, 2015, 14:30
  8. ^ Dichanthium annulatum. Pacific Island Ecosystems at Risk (PIER).
  9. ^ USFWS. Slender Rushpea Five-year Review. July 2008.
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Dichanthium annulatum: Brief Summary

provided by wikipedia EN

Dichanthium annulatum is a species of grass in the family Poaceae. It is commonly used as a forage for livestock.

Spikelet with long awn

Common names include marvel grass, Diaz bluestem, Kleberg bluestem, Hindi grass, ringed dichanthium, sheda grass, medio bluestem (var. papillosum), jargu grass, Delhi grass, vuda bluegrass, two-flowered golden-beard, Santa Barbara grass.

It is native to tropical Asia, the Middle East, and parts of Africa. In India, it is very important in agriculture. It has been introduced to many other parts of the world for cultivation, and it has become naturalized in some places, such as Australia.

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