Dontostemon tibeticus (Maxim.) Al-Shehbaz

Herbs biennial, (1.5-)4-14(-20) cm tall, pilose with simple trichomes to 1.5 mm, with somewhat fleshy root. Stems often few from base, simple above. Basal and lowermost cauline leaves eglandular; petiole (0.3-)1-2 cm; leaf blade lanceolate, oblong, or oblanceolate in outline, somewhat fleshy, (0.7-)1.2-2.7(-3.5) cm × (3-)5-1(-14) mm, base attenuate or cuneate, margin pectinate-pinnatifid, apex acute; lateral lobes (4-)7-11, to 5 × 2.5 mm, sometimes imbricate, abaxially toothed or entire, adaxially entire, acute; uppermost cauline leaves sessile. Fruiting pedicels divaricate, (2-)3-7(-9) mm, eglandular, stout. Sepals ovate, 3-4 × 1.5-2 mm, glabrous or sparsely pubescent apically, lateral pair slightly saccate. Petals white, with pink or purplish claws, obcordate, 5-8 × 2.5-3.5(-4) mm, apex emarginate; claw 3-4 mm. Filaments of median stamens 3-4 mm, free, flattened and sometimes minutely toothed on 1 side; filaments of lateral stamens 2-2.5 mm, slender; anthers oblong, 0.7-1 mm, apex obtuse. Ovules 12-20 per ovary. Fruit (0.8-)1-1.5(-1.7) cm × (0.8-)1-1.3 mm, straight, torulose, terete; valves with a prominent midvein and multicellular, glandlike warts; style 1-3 mm; stigma prominently 2-lobed, lobes subdecurrent. Seeds brown, oblong-ovate, 1.4-1.6 × 0.9-1 mm, wingless; cotyledons accumbent. Fl. Jun-Jul, fr. Jul-Aug.

Gansu, Qinghai, Xizang.

* Loose scree, disturbed alpine meadows, moist gravelly slopes, permafrost gravel and sandstone, steep rocky slopes; 3200-5200 m.

Nasturtium tibeticum Maximowicz, Fl. Tangut. 54. 1889.