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Hercules' Club

Zanthoxylum clava-herculis L.

Comprehensive Description

provided by North American Flora
Zanthoxylum clava-herculis L,. Sp. PI. 270. 1753
Zanthoxylum cavolinianum Lam. Encyc 2 : 39. 17S6.
Zanthox'ylonfraxinifolium Walt. Fl. Car. 243. 1788. Not Z. J rax ini : folium Marsh. 1785.
Fagara fraxini folia Lam. Tab. Enevc 1 : 334. 1791.
Zanthoxylum tricarpum Michx. FLBor. Am. 2 : 235. 1803.
Zanthoxylum aromaliaim Willd. Sp. PI. 4 : 755. 1806. Kampmania fra i inifolia Raf. Med. Rep. II. 5: 352. 1808.
/, talon glandulosum Raf. Fl. Ludov. 108. 1817. Zanthoxylum t atesbianum Raf. Med. Bot. 2 : 114. 1830. Zanlho I vlum glandulosum Raf. Med. Bot. 2 : 116. 1830. Zaniho vylum macrophyllum Nutt. N. Am. Sylva 3 : 10. 1849. Pseudopetalon tricarpum Raf.; S. Wats. Bibl. Ind. 155, as synonym. 1878. Fagara caroliniana Engler, in E. & P. Nat. Pfl.3<: 117. 1896. Fagara Clava-Herculis .Small, Fl. SE. U. S. 675. 1903.
A prickly shrub, or tree 8-17 m. high, the bark light-gray, bearing short, stout, conic or pyramidal, usually blunt, hard-corky prickles ; branches, petioles, and rachis pubescent when young, with age commonly glabrate, usually armed with sharp, often flat, straight or sometimes curved brownish prickles ; leaves odd-pinnate, 1-3 dm. long, persistent until late in the winter or until spring; leaflets 3-19, opposite or subopposite, oblique, ovate, ovate-lanceolate, or oblong-lanceolate, 2.5-8.5 cm. long, 1-3.5 cm. broad, acute to longacuminate at the apex, rounded or cuneate at the base, often somewhat falcate, crenateserrate, green and lustrous above, paler and glabrous or somewhat pubescent beneath when young, the pellucid glands numerous or scattered ; petioles and rachis terete, stout ; panicles corymb-like, 0.5-1.5 dm. long; staminate flowers: calyx 2.5-3 mm. broad; sepals 5, oblong, ovate-oblong, or ovate-lanceolate, about 1 mm. long, glabrous ; petals 5, green, oval, ovate, ovate-lanceolate, or sometimes obovate, 3-4 mm. long, 1.5-2 mm. broad ; stamens 5, longer than the petals ; filaments stout ; anthers ovate or oval ; pistillate flowers : calyx 2-3 mm. broad ; sepals ovate to ovate-lanceolate or ovate-oblong, about 1 mm. long, deciduous in fruit and sometimes while in flower ; petals narrowly elliptic to oblong or ovate, 2.5-3 mm. long, 1 mm. broad, green ; pistils 2 or 3, rarely 1, sessile ; styles short ; pedicels long, slender ; follicles obliquely-obovoid or globose-obovoid, brown, apiculate, 5-7 mm. long, the surface wrinkled or pitted ; seeds round-ellipsoid to obovoid, 4-5.5 mm. long, rather smooth or coarsely wrinkled, black, shining, at maturity hanging from the follicle.
Type locality : Sea coasts of Carolina.
Distribution : Southern Virginia to Florida, southern Arkansas, and Texas.
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bibliographic citation
John Kunkel Small, Lenda Tracy Hanks, Nathaniel Lord Britton. 1907. GERANIALES, GERANIACEAE, OXALIDACEAE, LINACEAE, ERYTHROXYLACEAE. North American flora. vol 25(1). New York Botanical Garden, New York, NY
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North American Flora

Zanthoxylum clava-herculis

provided by wikipedia EN

Zanthoxylum clava-herculis, the Hercules' club, Hercules-club, pepperwood, or southern prickly ash, is a spiny tree or shrub native to the southeastern United States. It grows to 10–17 m tall and has distinctive spined thick, corky lumps 2–3 cm long on the bark. The leaves are glabrous and leathery,[2] pinnately compound, 20–30 cm long with 7-19 leaflets, each leaflet 4–5 cm long. The flowers are dioecious, in panicles up to 20 cm long, each flower small, 6–8 mm diameter, with 3-5 white petals. The fruit is a two-valved capsule 6 mm diameter with a rough surface, and containing several small black seeds. The tree has also been called Z. macrophyllum. The genus name is sometimes spelled Xanthoxylum.

Hence "Hercules club"

Along with the related Zanthoxylum americanum, it is sometimes called "toothache tree"[3][4] or "tingle tongue" because chewing on the leaves, bark, or twigs causes a tingling numbness of the mouth, tongue, teeth and gums. It was used for such medicinal purposes by both Native Americans and early settlers to treat toothache because of this.[5]

The tree has a rounded crown and requires plentiful water and sunlight. Its leaves are browsed by deer and its fruit is eaten by birds. The fruit passes through birds, which helps the seeds to germinate.[5] The new trees tend to sprout below the favorite resting places of the birds, along fence rows and the edge of woods.[6] It is known to be host to a number of insect species, including the Giant Swallowtail (Papilio cresphontes) and the leaf beetle Derospidea brevicollis.

Potentially confused species

The name Hercules' club is also applied to Aralia spinosa, also native to eastern North America. Unlike Zanthoxylum, Aralia has large twice-compound leaves and very large leaf scars, so the trees are easily distinguished.

Chemistry

The benzophenanthridine alkaloid chelerythrine is the major active natural product found in Z. clava-herculis, exhibiting anti-bacterial activity against Staphylococcus aureus.[7]

References

  1. ^ IUCN SSC Global Tree Specialist Group.; Botanic Gardens Conservation International; et al. (BGCI) (2020). "Zanthoxylum clava-herculis". IUCN Red List of Threatened Species. 2020: e.T152906282A152906284. doi:10.2305/IUCN.UK.2020-1.RLTS.T152906282A152906284.en. Retrieved 20 November 2021.
  2. ^ Duncan, Wilbur H. and Marion B. Duncan (1988). Trees of the Southeastern United States. Athens, Georgia: The University of Georgia Press. pp. 77. ISBN 0-8203-1469-2.
  3. ^ "toothache tree". The American Heritage Dictionary of the English Language: Fourth Edition. 2000.
  4. ^ "toothache tree". The Columbia Encyclopedia, Sixth Edition. 2007.
  5. ^ a b Cox, Paul W.; Leslie, Patty (1999). Texas Trees. Corona Publishing Company. pp. 298–300. ISBN 0-931722-67-5.
  6. ^ Simpson, Benny J. (1988). A Field Guide to Texas Trees. Texas Monthly Press. p. 351. ISBN 0-87719-113-1.
  7. ^ Gibbons S., Leimkugel J., Oluwatuyi M. and Heinrich M. (2003). Activity of Zanthoxylum clava‐herculis extracts against multi‐drug resistant methicillin‐resistant Staphylococcus aureus (mdr‐MRSA). Phytotherapy Research, 17(3), 274-275.

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Zanthoxylum clava-herculis: Brief Summary

provided by wikipedia EN

Zanthoxylum clava-herculis, the Hercules' club, Hercules-club, pepperwood, or southern prickly ash, is a spiny tree or shrub native to the southeastern United States. It grows to 10–17 m tall and has distinctive spined thick, corky lumps 2–3 cm long on the bark. The leaves are glabrous and leathery, pinnately compound, 20–30 cm long with 7-19 leaflets, each leaflet 4–5 cm long. The flowers are dioecious, in panicles up to 20 cm long, each flower small, 6–8 mm diameter, with 3-5 white petals. The fruit is a two-valved capsule 6 mm diameter with a rough surface, and containing several small black seeds. The tree has also been called Z. macrophyllum. The genus name is sometimes spelled Xanthoxylum.

Hence "Hercules club"

Along with the related Zanthoxylum americanum, it is sometimes called "toothache tree" or "tingle tongue" because chewing on the leaves, bark, or twigs causes a tingling numbness of the mouth, tongue, teeth and gums. It was used for such medicinal purposes by both Native Americans and early settlers to treat toothache because of this.

The tree has a rounded crown and requires plentiful water and sunlight. Its leaves are browsed by deer and its fruit is eaten by birds. The fruit passes through birds, which helps the seeds to germinate. The new trees tend to sprout below the favorite resting places of the birds, along fence rows and the edge of woods. It is known to be host to a number of insect species, including the Giant Swallowtail (Papilio cresphontes) and the leaf beetle Derospidea brevicollis.

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