Oedenopiforma madecassa (Giordani Soika)

Oedenopiforma madecassa (Giordani Soika)

Paralimna madecassa Giordani Soika, 1956a:123.—Cogan, 1968:321 [revision]; 1980:663 [Afrotropical catalog].

Oedenopiforma madecassa.—Mathis and Zatwarnicki, 1995:117 [catalog, generic recombination].

Paralimna brunneifacies Giordani Soika, 1956a:123.—Cogan, 1968:321 [synonymy].

Paralimna ligabuei Canzoneri, 1987:84.—Mathis and Zatwarnicki, 1995:121 [world catalog]. [New synonym.]

DIAGNOSIS.—This species is distinguished from congeners by the following combination of characters: face and antenna of male appearing velvety, densely microtomentose, dark brown; face of female silvery gray; facial series of setae 2, these well separated; antenna of female dark brown; mesofrons of male dark brown anteriorly, becoming golden brown posteriorly; anterior fronto-orbits concolorous with anterior portion of mesofrons, not appearing velvety; scutum of male either mostly concolorous with scutellum except for golden brown anteromedial area or with disc of scutellum gray, contrasting with dark brown scutum; and forefemur of male usually bearing row of long, fine, pale, ventrally oriented setulae along anteroventral surface.

DESCRIPTION.—Small to moderately small shore flies, body length 1.65–2.40 mm.

Head: Paravertical seta well developed, length about – inner vertical seta. Arista with 6–8 dorsal rays. Face and antenna of male appearing velvety, densely microtomentose, dark brown; face of female silvery gray; antenna of female dark brown; facial series of setae 2, these well separated. Gena-to-eye ratio 0.26–0.36.

Thorax: Scutum of male mostly concolorous with scutellum except for golden brown anteromedial area. Stem vein bearing 1–3 (usually 2) setulae dorsally along posterior margin, length of each about equal to width of vein at level of insertion; costal-vein ratio 0.41–0.42; M-vein ratio 0.80–0.85. Forefemur bearing conspicuous, peg-like setulae anteroventrally; forefemur of male with row of long, fine, pale, ventrally oriented setulae along anteroventral surface.

Abdomen: Male terminalia (Figures 66–68): epandrium about as wide as high in posterior view (Figure 66), dorsum more acutely rounded and more thinly developed than lateral arms, bearing several long setulae toward posterior margin; cercus broadly ovoid, only slightly narrower dorsally, bearing numerous long setulae, evenly scattered; presurstylus in posterior view (Figure 66) as a nearly parallel-sided, horizontal bar bearing tiny setulae along ventral margin, apex only slightly narrower than base; postsurstylus in lateral view (Figure 68) with truncate, nearly flat basal margin, base nearly twice apical width, bearing 2 or 3 subapical setulae at level of preapical notch and bearing 5 or 6 setulae on basal of inner curvature; aedeagus with apex irregularly truncate in lateral view (Figure 68); aedeagal apodeme with wide and moderately deep keel, extended processes about equal in length in lateral view (Figure 68); hypandrium in ventral view (Figure 67) wider than long, bluntly rounded V-shaped, apex of posteriorly extended arms with wide, medial process, posterior margin very broadly excavate.

TYPE MATERIAL.—The lectotype male of Paralimna madecassa Giordani Soika, here designated to stabilize and make more universal the use of this name, is labeled “HOLOTYPUS [pink label with black submarginal line]/COLL. MUS. CONGO Madegascar: [Tamatave:] Maroansetra[,] XII-1949 [Dec 1949] J. Vadon/HOLOTYPUS Paralimna madecassa Soika [red label; handwritten except for “HOLOTYPUS″].” The lectotype male is double mounted (minuten), is in good condition (right fronto-orbital seta missing, posterior portion of left wing destroyed), and is deposited in the MRAC. Although Giordani Soika (1956a:123) wrote that the two specimens comprising the type series are females, one of the syntypes that we are here designating as the lectotype is clearly a male.

The lectotype female of Paralimna brunneifacies Giordani Soika, here designated to stabilize and make more universal the use of this name, is labeled “HOLOTYPUS [pink label with black submarginal line]/COLL. MUS. CONGO Madegascar: [Tamatave:] Maroansetra[,] XII-1949 [Dec 1949] J. Vadon/HOLOTYPUS Paralimna brunneifacies Soika [red label; handwritten except for “HOLOTYPUS”].” The lectotype female is double mounted (minuten), is in fair condition (right portion of mesonotum removed, right mid- and hindtarsi missing), and is deposited in the MRAC.

The holotype male of Paralimna ligabuei Canzoneri is labeled “[Sudan] Kartoum Barc (nilo) 6-xii-80 Leg. Rallo [handwritten]/HOLOTYPUS Paralimna ligabuei nov. det. Canzoneri S. [red label; species name handwritten]/MUSEO CIVICO DI STORIA NATURALE DI VENEZIA Materiale tipico descritto Inv. n. 03732 [red label; number handwritten].” The holotype is double mounted (minuten in a long, rectangular block of polyporus), is in fair condition (several setae missing; abdomen removed and dissected, and the parts are in an attached microvial), and is deposited in the MCV.

OTHER SPECIMENS EXAMINED.—Afrotropical. MADAGASCAR. Antananarivo: Ambatondrazaka, Centre Station Agric., Alaotra (800 m), 24 Dec 1957, B.R. Stuckenberg (1; BMNH). Toliara: Ampanihy (250 m), 16–18 Feb 1958, B.R. Stuckenberg (1; BMNH); Morondava (black light near sand beach), 28–31 Mar 1990, W.E. Steiner, C. Kremen, V. Razafimahatratra (3; USNM); Ranohira (860 m), 26 Jan–4 Feb 1958, B.R. Stuckenberg (3; BMNH); Saint Augustin (6 m), 11–13 Feb 1953, B.R. Stuckenberg (2 BMNH, USNM); Fenerive (lagoon shore), Dec 1955, B.R. Stuckenberg (1; USNM).

SOUTH AFRICA. Transvaal: Kruger Park (Olifants River near Balule; riparian woodland), 9 Dec 1972, B.R. and P. Stuckenberg (1; NMSA).

DISTRIBUTION.—Afrotropical: Madagascar, South Africa (Transvaal), Sudan.