There are no known predators of adult Falco biarmicus, and the species thrives in any area where it is left alone by humans. However, F. biarmicus eggs are vulnerable to scavengers that feed on them, as well as humans who rob nests for the pet/falconry industry.
Falco biarmicus is a medium-sized falcon, ranging from 35 to 50 cm long with a wingspan of 90 to 110 cm. The females are heavier, weighing from 700 to 900 g, whereas the males typically weigh from 500 to 600 g. The backs of adult lanner falcons are slate gray, juveniles are brown; both adults and juveniles have off-white or reddish-brown undersides streaked with gray. Northern subspecies have undersides spotted with black; southern subspecies lack spotted undersides. The head is reddish-brown or white with a black 'moustach' stripe. Females typically have darker coloration than males.
Range mass: 500 to 900 g.
Range length: 35 to 50 cm.
Range wingspan: 90 to 110 cm.
Sexual Dimorphism: female larger; sexes colored or patterned differently
Other Physical Features: endothermic ; bilateral symmetry
We do not have information on the lifespan of Falco biarmicus at this time.
Falco biarmicus is found in habitats varying from flat, dry areas near sea level to wet, forested mountains as high as 5000 m. Lanner falcons require large open or lightly wooded hunting areas, as well as rocky formations such as cliffs for nesting. However, F. biarmicus is also known to nest in trees and abandoned structures, as well as near the ground in desert areas.
Range elevation: 50 to 5000 m.
Habitat Regions: temperate ; terrestrial
Terrestrial Biomes: desert or dune ; savanna or grassland ; forest ; scrub forest ; mountains
Falco biarmicus is found as far north as the central/eastern Mediterranean region, extending south throughout most of Africa. Primarily a sedentary species, F. biarmicus does not migrate, though extensive wandering is frequently observed in Africa, especially in juveniles and non-breeding adults. In fact, ringed individuals have been recovered as far as 1528 km from their breeding territory. Lanner falcons are also known to move according to weather patterns, they move into desert areas after rain and out of forested areas during mist/heavy rain.
Biogeographic Regions: palearctic (Native ); ethiopian (Native )
Lanner falcons are carnivores. They feed on a variety of terrestrial and flying prey. Their main food sources are smaller birds, especially quails and columbids. Falco biarmicus also feeds on lizards, rodents, and bats, as well as spiders and scorpions in desert areas. If competition for these food resources is high, or locusts and other flying insects are swarming, F. biarmicus will also gorge itself on insects.
Animal Foods: birds; mammals; reptiles; insects; terrestrial non-insect arthropods
Primary Diet: carnivore (Eats terrestrial vertebrates)
Like all raptors, Falco biarmicus relies mostly on its keen sense of sight to hunt prey both in the air and on the ground. It has a variety of calls for different situations and communicates with other individuals acoustically, especially in territorial disputes and courtship rituals. It is known for its loud, repeated "kak-kak" call.
Communication Channels: acoustic
Perception Channels: visual ; tactile ; acoustic ; chemical
It is estimated that fewer than 1400 breeding pairs of Falco biarmicus exist in the world. Although not listed on the IUCN Red list, it is classified in Appendix II by CITES and is considered endangered at the European level. The population of F. biarmicus has decreased severely in the last fifty years due to destruction and loss of habitat, as well as human persecution (hunting, theft of eggs, and disturbance of nesting sites).
CITES: appendix ii
IUCN Red List of Threatened Species: least concern
Falco biarmicus does not usually affect humans in any way. However, when it inhabits agricultural areas, F. biarmicus frequently hunts domesticated fowl and poultry, typically chickens and ducks. This, unfortunately, prompts farmers to persecute F. biarmicus in order to protect their livestock.
Falco biarmicus is popular in the sport of falconry, and young birds and eggs are often taken from their natural habitat by humans. Over the past decade the numbers of breeding pairs of F. biarmicus have severely declined due to the harvesting of eggs by humans.
Positive Impacts: pet trade
Limited information is available about the role of this species in its ecosystem . Falco biarmicus> shares a niche with many other raptors, and competition between lanner falcons and peregrin falcons is high. Both feed primarily on small birds, however, F. biarmicus is able to adjust its diet accordingly if competition for this resource is too high. Lanner falcons are likely hosts to mites that commonly infest other birds; apart from this relationship, the main role of lanner falcons in an ecosystem is that of predators at the top of the food web.
Relatively little information is available about the mating system of Falco biarmicus. Falco biarmicus is monogamous; both males and females engage in elaborate flying and loud crying as part of the courtship display.
Mating System: monogamous
The breeding season for Falcon biarmicus varies significantly throughout its range. In southern Europe and northern Africa, the laying period is February through May. In the Sahara, western and northeastern Africa, the laying period is from January through March. In east, central and south Africa, the laying period is from June through November. Nesting habitat also varies; typical sites include abandoned raptor or heron nests, in trees, cliff faces, on the ground (desert areas) and buildings. A brood typically includes 3 to 4 eggs with an incubation period of around 32 days, fledging occurs in 35 to 47 days.
Breeding season: Breeding season varies significantly throughout the range.
Range eggs per season: 3 to 4.
Average time to hatching: 32 days.
Range fledging age: 35 to 47 days.
Average time to independence: 3 months.
Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; fertilization ; oviparous
There is little information about parental investment in Falco biarmicus. Eggs are incubated for around 32 days, and chicks fledge in 35 to 47 days. The male hunts alone early on, but the female assists in hunting later in the nesting season and during the fledgling period. Both the male and female take turns incubating the eggs.
Parental Investment: no parental involvement; altricial ; pre-hatching/birth (Protecting: Male, Female); pre-weaning/fledging (Provisioning: Male, Female, Protecting: Male, Female); pre-independence (Provisioning: Male, Female, Protecting: Male, Female)
Resident breeder, winter visitor and regular passage visitor?
The lanner falcon (Falco biarmicus) is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A large falcon, it preys on birds and bats.[2] Most likely either the lanner or peregrine falcon was the sacred species of falcon to the ancient Egyptians,[3] and some ancient Egyptian deities, like Ra and Horus, where often represented as a man with the head of a this falcon.
The lanner falcon was described by the Dutch zoologist Coenraad Jacob Temminck in 1825 under the current binomial name Falco biarmicus.[4] The type locality is Caffraria and the Cape of Good Hope.[5] Falco is Late Latin for a "falcon", from falx, falcis "sickle". The Swedish naturalist Carl Linnaeus had used the specific epithet biarmicus for the bearded reedling and Temminck clearly believed that the word meant "bearded" but it is likely that Linnaeus was using the Latinized form for Bjarmaland, a district in northern Russia.[6] The English word "lanner" is believed to come from the Old French lanier meaning "cowardly". The first recorded use of the word in English is from around 1400.[7]
This is presumably the oldest living hierofalcon species. Support for this assumption comes mainly from biogeography agreeing better with the confusing pattern of DNA sequence data in this case than in others. Nonetheless, there is rampant hybridization (like the perilanner) and incomplete lineage sorting which confounds the data to a massive extent; molecular studies with small sample sizes can simply not be expected to yield reliable conclusions in the entire hierofalcon group.
In any case, the radiation of the entire living diversity of hierofalcons seems to have taken place in the Eemian interglacial at the start of the Late Pleistocene, a mere 130,000–115,000 years ago; the lanner falcons would thus represent the lineage that became isolated in sub-Saharan Africa at some time during the Riss glaciation (200,000 to 130,000 years ago) already.[8][9][10][11]
There are five recognised subspecies:[12]
It is a large falcon, at 43–50 cm (17–20 in) length with a wingspan of 95–105 cm (37–41 in). Eurasian lanner falcons (Falco biarmicus feldeggi, also called Feldegg's falcon) have slate grey or brown-grey upperparts; most African subspecies are a paler blue grey above. The breast is streaked in northern birds, resembling greyish saker falcons, but the lanner has a reddish back to the head. Sexes are similar, but the browner young birds resemble saker falcons even more. However, sakers have a lighter top of the head and less clear head-side patterns. The lanner's call is a harsh "wray-e".
Lanner falcons are predominantly located in open habitats and can range from the forest edge to the desert. However, they are most commonly found in open savannah and sour grasslands. In South Africa, they commonly inhabit the east of the country in grasslands and move into the Fynbos, Nama Karoo and Southern Kalahari during the non-breeding season.[13]
Despite this movement, they are not truly migratory birds, and are usually limited to local movements. These movements may be in response to seasonal rains and altitudes, where higher elevations are inhabited during breeding season, and lower elevations are inhabited out of breeding season.[13]
They are bred in captivity for falconry; hybrids with the peregrine ("perilanners") are also often seen. Merret (1666) claimed that the "lanar" lived in Sherwood Forest and the Forest of Dean in England; such populations would seem to have derived from escaped hunting birds of the nobility.[14] Edward I of England (reigned 1272-1307), who had a passion for falconry, owned at least one lanner.
In the wild, lanner falcon numbers are somewhat declining in Europe, though the species remains relatively common in parts of Africa. In the Degua Tembien mountains of Ethiopia, it was observed to contribute to controlling pest rodents.[15]
Females usually moult from September to January, after the nesting period is over. Whereas, males moult from November to May, once the chicks can hunt for themselves.[16]
Lanner falcons most commonly prey on other birds, including doves, pigeons and domestic chickens or fowls.[17] These falcons have also been seen to prey on small mammals, reptiles and invertebrates. In Africa and Israel, lanner falcons were observed as hunting bats.[2]
It usually hunts by horizontal pursuit, rather than the peregrine falcon's stoop from a height, and takes mainly bird prey in flight. However, these raptors make use of a multitude of hunting styles, including stooping after soaring, attacking from a perch, attacking from a fast, low coursing flight and aerial attacks from a perch.[16] Lanner falcons also exhibit co-operative hunting with high success rate specifically when hunting jackdaws, where the use of visual contact is used to coordinate the chase.[18] Males most commonly make the initial attack, yet the prey is usually caught by the female.
Before the breeding season, both males and females show mutual soaring and flapping flight which are closely synchronized from April to June. Near nesting sites, pairs have also been seen to power-dive together.[16]
Once eggs are fledged, the female predominantly stays with and feeds the juveniles. However, the male takes over this role when the female needs to hunt. When the juveniles are about 25 days old, the female is commonly seen perched near the nest, but still spends a large amount of time in the nest with the juveniles. When the juveniles are about 39 days old, the female does not spend large amounts of time in the nest anymore and both the female and male are actively hunting.[16]
Lanner falcons usually nest on sheer cliff faces, and lay three or four eggs. They do not have strict nesting regulations and as a result are more common and do not show a patchy distribution. As a result they also make use of old crow nests and are sometimes seen nesting on pylons, trees and less steep cliff faces.[19]
Lanner falcon have a laying period in July which suggests a strategy to fledge young before the heat and heavy rain of the summer as well as before the influx of migratory birds during December to February. This timing may improve foraging conditions for juveniles which can increase the success of breeding. Breeding success for these falcons is largely affected by environmental conditions.[20] The incubation period is thought to be 32 days and the nesting period to be 44 days. However, juveniles have been present near nests up to ten weeks after fledgling.[16] The mean fledgling rate is 2.24 young per successful pair which is typical of large falcons, however, the mean number of individuals fledged overall per territorial pair is 1.3 which is low and may be due to the number of unproductive years towards the end of each territory's occupancy.[16]
Lanner falcons are of Least Concern on the IUCN Red List, yet they can still be negatively affected through anthropogenic process, directly and indirectly.
Direct anthropogenic threats to these raptors include nest robbery, vehicle collisions, electrocution from power lines and persecution. As chicks can barely fly when leaving the nest they are vulnerable and easy to catch, they are threatened by being captured for falconries and through being caught and killed by angry farmers as a result of these birds sometimes preying on domestic chickens.[16]
Indirect anthropogenic threats include loss of suitable habitats as a result of human expansion and the intensive use of pesticides for agriculture.[13]
Lanner falcon at Kgalagadi Transfrontier Park, South Africa
Painting by John Gerrard Keulemans (1884)
Lanner falcon at Plettenberg Bay, South Africa
Adult Falco biarmicus biarmicus, Etosha National Park, Namibia.
Juvenile, probably F. b. feldeggi. Note blue facial skin and overall similarity to saker falcon.
Falco biarmicus feldeggii - MHNT
Falco biarmicus erlangeri - MHNT
The lanner falcon (Falco biarmicus) is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A large falcon, it preys on birds and bats. Most likely either the lanner or peregrine falcon was the sacred species of falcon to the ancient Egyptians, and some ancient Egyptian deities, like Ra and Horus, where often represented as a man with the head of a this falcon.