Der Griechische Neunstachlige Stichling (Pungitius hellenicus) ist eine in Griechenland endemische Stichlingsart. Er zeichnet sich gegenüber anderen Stichlingen durch bemerkenswerte Reduktionserscheinungen aus.
Diese Stichlingsart kommt ausschließlich in einem eng begrenzten Gebiet im Einzugsbereich des Sperchios vor, wobei inzwischen nur noch an drei Orten Populationen existieren: In den Karstgewässern nahe Agia Paraskevi und damit verbundenen Entwässerungskanälen, in Quellbächen sowie Be- und Entwässerungskanälen in der Umgebung von Moschochori und in einigen Quellbächen in der Nähe des Dorfes Kombotades. Während der Griechische Neunstachlige Stichling bei Moschohori noch relativ häufig gefunden werden kann, sind die anderen beiden Vorkommen deutlich kleiner. Die Bestände am locus typicus, dem Fundort der Typusexemplare, einem Bach in Kombotades, wurden ausgelöscht, als dieser zerstört wurde.
Der typische Lebensraum sind klare Bäche und kleine Flüsse mit sandigem und steinigem Grund und relativ niedrigen Wassertemperaturen von nicht mehr als 20 °C im Sommer. Die Art benötigt außerdem den Schutz dichter Vegetation.
Obwohl der Trivialname anderes vermuten lässt, trägt der Griechische Neunstachlige Stichling in der Regel nur zwei bis sechs Rückenstacheln. Die Bauchflossen fehlen völlig, ebenso wie die bei anderen Stichlingen üblichen Ventralstacheln. Die sonst für Vertreter der Gattung Pungitius typische, zusammenhängende Bauchplatte ist stark reduziert. An den Seiten fehlt jegliche Beschilderung.
Der maximal 50 Millimeter lange Körper ist schlank, der Kopf wirkt im Verhältnis dazu relativ groß. Die blass olivgrüne Färbung wird an den Seiten durch dunkle Streifen oder Flecken unterbrochen.
Anzahl der Flossenstrahlen:
Der Griechische Neunstachlige Stichling lebt zurückgezogen und verbringt den größten Teil des Tages versteckt in dichter Unterwasservegetation. Offene Freiwasserbereiche sucht er nur selten auf. Sein Nahrungsspektrum umfasst Wirbellose, deren Eier und Larvenstadien sowie Fischbrut.
Nach der Laichperiode im Mai und Juni, betreibt das Männchen, wie bei allen Stichlingsarten, eine intensive Brutpflege bis der Nachwuchs eine Größe von etwa fünf bis sechs Millimetern erreicht hat. Mit einer Länge von zehn bis elf Millimetern gleichen die Jungtiere den adulten Exemplaren in Färbung und Gestalt. Mit einer maximalen Lebenserwartung von 18 Monaten erlebt ein Griechischer Neunstachliger Stichling nur eine Fortpflanzungssaison.
Die Art ist durch ihre geringe Größe und eine hohe Reproduktionsrate an kleine und instabile Gewässersysteme angepasst. Sie ist jedoch auch auf ein sehr kleines Verbreitungsgebiet beschränkt und dort fortdauernden Belastungen durch Wasserentnahmen, Umweltverschmutzung und Habitatzerstörungen, wie die Auffüllung von Kleingewässern zur Gewinnung von Nutzflächen, ausgesetzt. In der Folge ist die Population stark fragmentiert und anfällig für zufallsbedingte Störungen wie beispielsweise Gendefekte oder Trockenheiten. Die IUCN führt den Griechischen Neunstachligen Stichling daher als "stark gefährdet". Der griechische Staat stellte ihn mit dem Gesetz 67/1981 unter Schutz.
Der Griechische Neunstachlige Stichling (Pungitius hellenicus) ist eine in Griechenland endemische Stichlingsart. Er zeichnet sich gegenüber anderen Stichlingen durch bemerkenswerte Reduktionserscheinungen aus.
Pungitius hellenicus, the Greek ninespine stickleback or ellinopygósteos, is a species of fish in the family Gasterosteidae. It is endemic to Greece. Its natural habitats are rivers and freshwater spring. It is threatened by habitat loss and considered critically endangered in the International Red List of IUCN, Bern Convention (Appendix III).
A combination of five traits are characteristic of the ellinopygósteos species: caudal peduncle keel absent, its ectocoracoid reduced, dorsal spines fewer than seven, pelvic girdle absent or vestigial, and large lateral scutes absent. The body is moderately compressed. The head is conical and the interorbital area flattened. Bones are weakly ossified and sculpturing poorly developed on the cranial bones. Mouth slightly supraterminal, oblique, and continuous groove separating upper lip from maxillary. Numerous small sharp teeth confined to upper and lower jaws, absent on roof of mouth. Gill membranes extending forward, broadly joined to each other, and free from isthmus. Gill rakers 7–10. Opercular opening extending above pectoral fin base. Pectoral fin soft rays 10 (rarely 11). Dorsal fin spines 2–6, isolated (with small triangular fin membrane), directed posteriorly and not inclined from middorsal line, and depressible in shallow groove. First dorsal spine shortest and last one longest. Dorsal fin soft rays 8–11. Anal fin with one spine and 6-10 soft rays. Caudal fin with 12 soft rays, rounded, and deeper than wide. Vertebrae 29-30 (usually 30) with 12-13 precaudal vertebrae. Lateral line inconspicuous with 28-38 (usually 32) small round scutes. The longest reported specimens are 50 mm. Background color pale olive, sides of the body pigmented with dark bars or blotches.[2]
This species is confined to three localities in the Spercheios valley in central Greece: the Aghia Paraskevi Spring, 5 km east of Lamia, and an associated system of drainage channels extending over the areas Diplosoudi and Bourdara, a large system of connected drainage and irrigation channels and natural wells extending over the areas Lykochoria and Kaikia near the village of Moschochori, and a small number of natural wells near the village of Kompotades. The species is abundant in some channels of the Lykochoria area, but is moderately abundant or rare in the rest of its range.[2]
The typical habitat consists of relatively cool spring or slow-running waters (maximum temperature rarely exceeding 20 °C in summer) with rich vegetation. Specimens inhabit small water bodies and, with its camouflage coloration, exhibit cryptic behavior, remaining among aquatic vegetation during most of the day, and rarely seen in the open areas. In most areas, this species occurs in sympatry with Pelasgus marathonicus, Alburnoides bipunctatus, Gambusia holbrooki, and Gasterosteus aculeatus. Seasonal presence of Squalius and Barbus sp. has been recorded in some wells and localities that communicate with the Sperchios river. Competition for food (both intra- and inter-specific) does not appear to be a critical survival factor. Stomach content analysis of sympatric species has failed to identify predation on P. hellenicus, but some cases of cannibalism are known. Food consists mainly of amphipods, isopods, benthic copepods, bivalves, gastropods, oligochaetes, insects and their larvae, fish larvae, invertebrate eggs, and demersal prey or prey associated with vegetation. Maximum recorded age is 18 months.[2]
Spawning occurs mainly in May and June, with, only one breeding period in their lifetime. During the reproductive period the male acquires breeding coloration, constructs nests from plant material where the female deposits the eggs, and provides care to the eggs until the time they hatch. The morphological and morphometric development of larvae and juveniles is known from both field observations and breeding in aquaria. Larvae hatch at about 5.5 mm SL, exhibit the general morphological characteristics of the family Gasterosteidae, have a well-developed pigmentation pattern and exhibit cryptic behavior. Adult morphology is acquired at about 10–11 mm SL.[2]
The type locality (a spring in Kombotades village) is destroyed but small populations remain in nearby wells. The status of the species in Aghia Paraskevi Spring is satisfactory. Water removal and summer drought may reduce the area of the spring basins available for this population, but the presence of this species in associated channels reduces the danger of extinction. The population in the system of channels and wells of the Moschohori area is relatively safe. Although individual wells are filled in with earth to increase the cultivation area and the channels are occasionally disturbed for maintenance purposes (excavations, plant cleaning, etc.), the system as a whole is relatively stable. The species seems to be well adapted to small and unstable water systems by means of its life-history strategy (small body size, annual life-cycle, high reproductive effort) which permit an opportunistic response to windows of environmental variability, high probability of survival up to the age of reproduction and high colonization efficiency. Aghia Paraskevi Spring and the species have been granted protection by law No. 67/1981 of the Greek state and official decision of the Phthiotis Prefect.[2]
Much basic ecological information is needed for this species, including habitat requirements, life history and reproduction behavior, and surveys of existing populations. No holotype was designated for this species, but eight syntypes are in the Muséum National d'Histoire Naturelle, Paris (MNHN 1975-0867).[2]
Pungitius hellenicus, the Greek ninespine stickleback or ellinopygósteos, is a species of fish in the family Gasterosteidae. It is endemic to Greece. Its natural habitats are rivers and freshwater spring. It is threatened by habitat loss and considered critically endangered in the International Red List of IUCN, Bern Convention (Appendix III).
Pungitius hellenicus Pungitius generoko animalia da. Arrainen barruko Gasterosteidae familian sailkatzen da.
Pungitius hellenicus Pungitius generoko animalia da. Arrainen barruko Gasterosteidae familian sailkatzen da.
Pungitius hellenicus is een straalvinnige vissensoort uit de familie van de stekelbaarzen (Gasterosteidae).[1] De wetenschappelijke naam van de soort is voor het eerst geldig gepubliceerd in 1971 door Stephanidis.
Bronnen, noten en/of referenties
Pungitius hellenicus é uma espécie de peixe da família Gasterosteidae.[1]
Os seus habitats naturais são: rios e nascentes de água doce.[1]
Está ameaçada por perda de habitat.[1]
Pungitius hellenicus é uma espécie de peixe da família Gasterosteidae.
É endémica de Grécia.
Os seus habitats naturais são: rios e nascentes de água doce.
Está ameaçada por perda de habitat.
Pungitius hellenicus — вид риб родини колючкових (Gasterosteidae). Прісноводна бентопелагічна риба, що сягає 4.5 см довжиною. Є ендеміком Греції, де мешкає виключно у нижньому басейні річки Сперхіос.
Pungitius hellenicus — вид риб родини колючкових (Gasterosteidae). Прісноводна бентопелагічна риба, що сягає 4.5 см довжиною. Є ендеміком Греції, де мешкає виключно у нижньому басейні річки Сперхіос.
Pungitius hellenicus là một loài cá thuộc họ Gasterosteidae.[1][2][3] Nó là loài đặc hữu của Hy Lạp. Các môi trường sống tự nhiên của chúng là sông và suối. Nó bị đe dọa do mất môi trường sống và được IUCN xếp vào loài bị de dọa.[4]
|accessdate= (trợ giúp)
Bài viết Lớp Cá vây tia này vẫn còn sơ khai. Bạn có thể giúp Wikipedia bằng cách mở rộng nội dung để bài được hoàn chỉnh hơn. Pungitius hellenicus là một loài cá thuộc họ Gasterosteidae. Nó là loài đặc hữu của Hy Lạp. Các môi trường sống tự nhiên của chúng là sông và suối. Nó bị đe dọa do mất môi trường sống và được IUCN xếp vào loài bị de dọa.
